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This work was supported in part by the National Natural Science Foundation of China (grant NO 31372448), Special Fund for Agro-scientific Research in the Public Interest (201303042), Science and Technology Planning Project of Guangdong Province (20140224), Guangdong Provincial Key Laboratory of Prevention and Control for Severe Clinical Animal Diseases (2013A061401013), the Promote Scientific and Technological Development Program, China (2013B040200032) and a grant from the US National Institute of Allergy and Infectious Diseases (R01AI108993 to Dr Gregory C Gray).
Supplementary information of this article can be found on the Emerging Microbes and Infections's website: http://www.nature.com/emi.