ABSTRACT
The relationship between pupil shape and accommodation was examined in two Red Sea fishes, Hemibalistes chrysoptera and Fistutaria petimba. Both species have markedly asymmetrical pupils and both species exhibit large accommodative lens movement ability in the pupil plane. Accommodative variation in the location of the lens within the pupil, as well as the absence of a binocular field of vision in F. petimba, indicate that pupil asymmetry can best be explained in terms of the need for space for accommodative lens movement.