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ABSTRACT
Germlings of Uromyces appendicular develop appressoria when they grow on a substrate with a certain topography. Such a topography can be a stomatal guard cell, or a scratch on an artificial substrate with a mean depth of 0.4 um and a width of 0.95 urn. The microtubule and F-actin cytoskeleton was observed to become reoriented parallel to such scratches.