Abstract
Auxospore formation and vegetative cell division are described for Mastogloia smithii. This species is unique among raphid diatoms for its cis-type behavioural anisogamy. This results in both auxospores being formed within one gametangial frustule. Auxospore formation in M. smithii has its closest parallels with certain araphid pennate taxa and seems to represent a link between the reproductive methods of araphid and raphid pennate taxa.