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ABSTRACT
Additional information about one of the oldest known gecko-like lizards, Hoburogekko suchanovi from the Early Cretaceous of Mongolia, is provided in light of recent advances in the understanding of gekkotan osteology. The phylogenetic affinities of this taxon were assessed by adding it into an existing morphological data set of 224 taxa coded for 364 characters. The effects of character ordering and scoring on tree topology and the composition of Gekkonomorpha were also explored. The results of the analysis indicate that Gekkonomorpha is highly unstable and additional revision of stem gekkotans (e.g., Parviraptor, Gobekko) is required to refine the basal relationships of this clade. Like modern gekkotans, Hoburogekko possess a tubular frontal bone. In contrast to extant gekkotans, the Meckelian canal is only partially fused and may represent an incipient form of the gekkotan condition. The posterior shelf of the maxilla is wider than in modern gekkotans, perhaps related to a plesiomorphically large jugal. The short, rounded snout may indicate a lifestyle as found in some modern gekkotans found in equivalent xeric environments to the inferred paleoenviroment of the Höövor (Mongolia) locality during the Early Cretaceous.
ACKNOWLEDGMENTS
We thank S. V. Saveliev for the help in photographing of fossil specimens and S. E. Evans and M. E. H. Jones for their useful comments on an earlier version of the manuscript. C. J. Bell provided osteological specimens for comparative purposes. We thank the Willi Hennig Society for sponsoring the program TNT and making it freely available. J. S. Arias assisted in writing computer macros for the phylogenetic analyses. This research was supported by grant DEB 0844523 from the National Science Foundation and by the Gerald M. Lemole, M.D., Endowed Chair in Integrative Biology Fund at Villanova University.
Handling editor: Johannes Müller