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ABSTRACT
A new dryosaurid ornithopod, Eousdryosaurus nanohallucis, gen. et sp. nov., is described here based on a single specimen from the Late Jurassic Alcobaça Formation of Portugal. Eousdryosaurus nanohallucis is distinguished from all other dryosaurids by eight autapomorphic features and an unique combination of characters, some of which are also shared by other dryosaurids. Eousdryosaurus is linked with Dryosauridae, because the fourth trochanter is proximally placed and widely separated from the scar for the insertion of the M. caudifemoralis longus, which is restricted to the medial surface of the femoral shaft. Phylogenetic analysis nests Eousdryosaurus in an unresolved polytomy at the base of Dryosauridae together with Callovosaurus, Dryosaurus, and Kangnasaurus. The complete pes of Eousdryosaurus, which has a phalangeal formula of 1-3-4-5-0, supports the putative autapomorphic reduction of the dryosaurid pes that also occurs in parallel in more derived ornithopods.
SUPPLEMENTAL DATA—Supplemental materials are available for this article for free at www.tandfonline.com/UJVP
ACKNOWLEDGMENTS
We thank J. J. dos Santos who found the specimen and F. Marcos and G. Ramalheiro for its preparation. We are grateful to L. M. Chiappe (The Dinosaur Institute, Natural History Museum of Los Angeles County, Los Angeles, U.S.A.), R. A. Coria (Museo Carmen Funes, Plaza Huincul, Argentina), J. Sertich (Denver Museum of Nature and Science, Denver, U.S.A.), and M. Ramalho and R. Silva (Museu Geológico, Lisbon, Portugal) for their help and for providing access to collections. P. M. Barrett, P. M. Galton, and an anonymous reviewer made constructive suggestions for improving the manuscript. We thank T. Cater for critical reading of the manuscript. We are grateful to C. Gallardo (Departamento de Filología Clásica, Universidad Autónoma de Madrid, Madrid, Spain) for providing suggestions regarding the use of Latin. Research was partially supported by Vicerrectorado de Investigación of the Universidad Nacional de Educación a Distancia (UNED), Project PTDC/CTE–GEX/67723/2006, ‘Study of the vertebrate faunas of the Upper Jurassic of the Lusitanian Basin and paleobiogeographic implications (VERT-JURA)’ (Fundação para a Ciência e a Tecnologia, Portugal), and the Câmara Municipal de Torres Vedras (Portugal). Support was also provided by a donation from A. C. Rodrigues (Torres Vedras, Portugal) to the SHN.
Handling editor: Paul Barrett.
