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This work was supported by the Natural History Museum, London, and I am grateful to the curators in Paris (PC) and Geneva (G) for the loan of specimens. I also thank Professor Dr Ryszard Ochyra (KRAM), Dr Michelle Price (G) and Jo Wilbraham (BM), for their constructive advice with regard to the interpretation of the nomenclatural complexities surrounding the taxa in this article.
Taxonomic Changes and Additions: Nil.
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