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Abstract
Diplocaulus minimus Dutuit, 1988, the diplocaulid lepospondyl from the Late Permian of the Argana Basin (Morocco) is revised based on a redescription of the type material, together with a description of recently found specimens. The revised specimens, previously reported to Diplocaulus, prove to be more closely related to the genus Diploceraspis. These animals exhibit a peculiar asymmetry of the skull. They are the only known Gondwanian lepospondyls and their unique co-occurrence with pareiasaurs strengthens the hypotheses of climate-controlled endemism in the end-Permian Pangea.
Acknowledgements
The author greatly acknowledges the ‘Fondations des Treilles’ (for donating a fund to study the Permian vertebrates from the Argana Basin), Philippe Janvier (UMR 7207-CNRS/MNHN) for allowing the study of the specimens from Dutuit's collections, Nour-Eddine Jalil for welcoming him in his team, his help and kindness, J.-S. Steyer for fruitful discussions, Maeva Orliac for the drawings and corrections to the manuscript and Renaud Vacant (UMR 7207-CNRS/MNHN) for the preparation of the specimens. The author finally acknowledges again N.-E. Jalil and J.-S. Steyer for their exhaustive and constructive reviews that strongly improved the manuscript.
