A revision of the moss Syrrhopodon spiculosus Hook. & Grev. (Calymperaceae, Bryopsida) and associated taxa, including two new species described from Borneo

Abstract

Syrrhopodon spiculosus Hook. & Grev. and some associated taxa are revised. Syrrhopodon elimbatus Dixon and Syrrhopodon patens Dixon are confirmed as synonymous with S. spiculosus; Syrrhopodon hispidissimus Dixon and Syrrhopodon scalariformis Dixon are recognized as distinct species; S. spiculosus ssp. bokorenis Tixier is recognized as a synonym of S. mammillosus Müll.Hal. and Syrrhopodon perarmatus Broth. in Broth & Watts is placed in synonymy with Syrrhopodon subciliatus Müll. Hal. Two new species are described from Borneo: Syrrhopodon teniolatus L.T.Ellis, sp. nov., which resembles S. hispidissimus but has a differentiated rib (teniola) within the margin of the distal hyaline leaf base, and Syrrhopodon multipapillosus L.T.Ellis, sp. nov. This resembles a miniature version of Syrrhopodon kilauae Müll.Hal. but has cells in the chlorophyllose lamina exserted dorsally and/or ventrally as tall, papillose projections.

Keywords:

• CALYMPERACEAE
• SYRRHOPODON SPICULOSUS HOOK. AND GREV
• SYRRHOPODON TENIOLATUS L.T.ELLIS
• SP. NOV
• SYRRHOPODON MULTIPAPILLOSUS L.T.ELLIS
• SP. NOV
• BORNEO