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Abstract
In this study we sequenced and analyzed the complete mitochondrial genome (mitogenome) of the lined shore crab Pachygrapsus crassipes Randall, 1840 (Crustacea: Grapsidae). The full-length P. crassipes mitogenome is 15,652 bp in size, which encodes the same 37 genes as all metazoan mitogenomes. Both AT contents of the entire molecule as well as putative control region display lowest values among all mitogenomes of the brachyuran crabs determined to date. The mitochondrial gene order follows a classic crab-type arrangement that underwent a unique tRNA translocation from the pancrustacean ancestral pattern. Our results will provide important data for phylogenetic as well as biogeographic studies.