Abstract
The Harpellales genus Smittium is based on a type species, S. arvernense, which was described by Poisson in 1937 without designation of a type specimen. Smittium arvernense has not been reported since its originalPUBLICation. Because the other 79 species of Smittium cannot be compared to the type species, a lectotype is proposed as well as an epitype for that lectotype that is also the holotype of S. mucronatum. Because Smittium is believed to be polyphyletic these type designations will provide stable application of names and, as well resolved phylogenetic analyses of member species emerge based on morphological and DNA sequence characters, they will provide a foundation for a more robust and revised classification.
This research was supported by National Science Foundation REVSYS collaborative grants DEB-0918169 (to RWL) and DEB-0918182 (to MMW). Discovery and culture of S. mucronatum by the authors and their various former and current collaborators during surveys from several countries were accomplished with the aid of prior NSF support (GB-7072X, DEB-9220518, DEB-0108110, PEET Award DEB-9521811, and DEB-0344722). We are greatly indebted to Drs Scott Redhead and Paul M. Kirk and anonymous reviewers for valuable suggestions.