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Journal of Vertebrate Paleontology Volume 37, 2017 - Issue 4
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The second titanosaurian (Dinosauria: Sauropoda) from the middle Cretaceous Galula Formation, southwestern Tanzania, with remarks on African titanosaurian diversity

Eric GorscakDepartment of Biological Sciences, Ohio University, 107 Irvine Hall, Athens, Ohio45701, U.S.A.;Ohio Center for Ecology and Evolutionary Studies, Ohio University, 107 Irvine Hall, Athens, Ohio45701, U.S.A.Correspondence[email protected]
ORCID Iconhttp://orcid.org/0000-0002-4019-8301
ORCID Icon,
Patrick M. O'ConnorOhio Center for Ecology and Evolutionary Studies, Ohio University, 107 Irvine Hall, Athens, Ohio45701, U.S.A.;Department of Biomedical Sciences, Ohio University Heritage College of Osteopathic Medicine, 228 Irvine Hall, Athens, Ohio45701, U.S.A., [email protected]; [email protected]ORCID Iconhttp://orcid.org/0000-0002-6762-3806
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Eric M. RobertsDepartment of Geosciences, College of Science and Engineering, James Cook University, Townsville, Queensland, 4811Australia, [email protected]
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Nancy J. StevensOhio Center for Ecology and Evolutionary Studies, Ohio University, 107 Irvine Hall, Athens, Ohio45701, U.S.A.;Department of Biomedical Sciences, Ohio University Heritage College of Osteopathic Medicine, 228 Irvine Hall, Athens, Ohio45701, U.S.A., [email protected]; [email protected]ORCID Iconhttp://orcid.org/0000-0002-2402-6526
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Article: e1343250 | Received 19 Dec 2016, Accepted 15 May 2017, Published online: 24 Aug 2017

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J. Logan King, Julia B. McHugh, Mathew J. Wedel & Brian Curtice. (2023) A previously unreported form of dorsal rib pneumaticity in Apatosaurus (Dinosauria: Sauropoda) and implications for pneumatic variation among diplodocid dorsal ribs. Journal of Vertebrate Paleontology 43:5.
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Eric Gorscak, Matthew C. Lamanna, Daniela Schwarz, Verónica Díez Díaz, Belal S. Salem, Hesham M. Sallam & Marc Filip Wiechmann. (2022) A new titanosaurian (Dinosauria: Sauropoda) from the Upper Cretaceous (Campanian) Quseir Formation of the Kharga Oasis, Egypt. Journal of Vertebrate Paleontology 42:6.
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Felix J. Augustin, Andreas T. Matzke, Michael W. Maisch & Hans-Ulrich Pfretzschner. (2021) Dinosaur taphonomy of the Jurassic Shishugou Formation (Northern Junggar Basin, NW China) – insights from bioerosional trace fossils on bone. Ichnos 28:2, pages 87-96.
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Articles from other publishers (27)

Theresa J. Orr, Eric M. Roberts, Michael I. Bird, Cassy Mtelela, Patrick M. O'Connor & Nancy J. Stevens. (2023) Paleosol-derived paleoclimate and paleoenvironment reconstruction of the Rukwa Rift Basin, Tanzania: implications for faunal dispersal in the Miocene–Pliocene. Journal of Sedimentary Research 93:5, pages 309-326.
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Stephen F. Poropat, Philip D. Mannion, Samantha L. Rigby, Ruairidh J. Duncan, Adele H. Pentland, Joseph J. Bevitt, Trish Sloan & David A. Elliott. (2023) A nearly complete skull of the sauropod dinosaur Diamantinasaurus matildae from the Upper Cretaceous Winton Formation of Australia and implications for the early evolution of titanosaurs . Royal Society Open Science 10:4.
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Agustín Pérez Moreno, Alejandro Otero, José L. Carballido, Leonardo Salgado & Jorge O. Calvo. (2023) The appendicular skeleton of Rinconsaurus caudamirus (Sauropoda: Titanosauria) from the Upper Cretaceous of Patagonia, Argentina. Cretaceous Research 142, pages 105389.
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Julian C. G. Silva Junior, Agustín G. Martinelli, Thiago S. Marinho, João Ismael da Silva & Max C. Langer. (2022) New specimens of Baurutitan britoi and a taxonomic reassessment of the titanosaur dinosaur fauna (Sauropoda) from the Serra da Galga Formation (Late Cretaceous) of Brazil . PeerJ 10, pages e14333.
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Stephen F. Poropat, Timothy G. Frauenfelder, Philip D. Mannion, Samantha L. Rigby, Adele H. Pentland, Trish Sloan & David A. Elliott. (2022) Sauropod dinosaur teeth from the lower Upper Cretaceous Winton Formation of Queensland, Australia and the global record of early titanosauriforms. Royal Society Open Science 9:7.
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Flavio Bellardini, Rodolfo A Coria, Diego A Pino, Guillermo J Windholz, Mattia A BaianoAugustin G Martinelli. (2022) Osteology and phylogenetic relationships of Ligabuesaurus leanzai (Dinosauria: Sauropoda) from the Early Cretaceous of the Neuquén Basin, Patagonia, Argentina . Zoological Journal of the Linnean Society.
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Agustín Pérez MorenoJosé L. CarballidoAlejandro OteroLeonardo SalgadoJorge O. Calvo. (2022) The Axial Skeleton of Rinconsaurus caudamirus (Sauropoda: Titanosauria) from the Late Cretaceous of Patagonia, Argentina. Ameghiniana 59:1.
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Mauro Aranciaga Rolando, Jordi A. García Marsà, Federico L. Agnolín, Matías J. Motta, Sebastián Rozadilla & Fernando E. Novas. (2022) The sauropod record of Salitral Ojo del Agua: An Upper Cretaceous (Allen Formation) fossiliferous locality from northern Patagonia, Argentina. Cretaceous Research 129, pages 105029.
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José L. Carballido, Alejandro Otero, Philip D. Mannion, Leonardo Salgado & Agustín Pérez Moreno. 2022. South American Sauropodomorph Dinosaurs. South American Sauropodomorph Dinosaurs 269 298 .
Jose L. Carballido, Flavio Bellardini & Leonardo Salgado. 2022. South American Sauropodomorph Dinosaurs. South American Sauropodomorph Dinosaurs 237 268 .
F. Bellardini, G.J. Windholz, M.A. Baiano, A.C. Garrido & L.S. Filippi. (2021) New titanosaur remains from the Portezuelo Formation (Turonian–Coniancian) and their implications for the sauropod faunal diversity of the southern Neuquén Basin, Patagonia, Argentina. Journal of South American Earth Sciences 111, pages 103457.
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Theresa J. Orr, Eric M. Roberts, Christopher M. Wurster, Cassy Mtelela, Nancy J. Stevens & Patrick M. O'Connor. (2021) Paleoclimate and paleoenvironment reconstruction of paleosols spanning the Lower to Upper Cretaceous from the Rukwa Rift Basin, Tanzania. Palaeogeography, Palaeoclimatology, Palaeoecology 577, pages 110539.
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Belal S. Salem, Patrick M. O'Connor, Eric Gorscak, Sanaa El-Sayed, Joseph J.W. Sertich, Erik Seiffert & Hesham M. Sallam. (2021) Dinosaur remains from the Upper Cretaceous (Campanian) of the Western Desert, Egypt. Cretaceous Research 123, pages 104783.
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Alexander O. Averianov, Alexander V. Sizov & Pavel P. Skutschas. (2021) Gondwanan affinities of Tengrisaurus, Early Cretaceous titanosaur from Transbaikalia, Russia (Dinosauria, Sauropoda). Cretaceous Research 122, pages 104731.
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Adrián Páramo, Pedro Mocho & Francisco Ortega. (2020) Three-dimensional analysis of the titanosaurian limb skeleton: implications for systematic analysis. Journal of Iberian Geology 46:4, pages 369-402.
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Christina Shears Ozeki, David M. Martill, Roy E. Smith & Nizar Ibrahim. (2020) Biological modification of bones in the Cretaceous of North Africa. Cretaceous Research 114, pages 104529.
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Jean-Claude Rage*Emmanuel Gheerbrant. 2020. Biological Consequences of Plate Tectonics. Biological Consequences of Plate Tectonics 251 264 .
Kamila L.N. Bandeira, Elaine Batista Machado, Diogenes Campos & Alexander W.A. Kellner. (2019) New titanosaur (Sauropoda, Dinosauria) records from the Morro do Cambambe Unit (Upper Cretaceous), Mato Grosso state, Brazil. Cretaceous Research 103, pages 104155.
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Philip D. Mannion, Paul Upchurch, Xingsheng Jin & Wenjie Zheng. (2019) New information on the Cretaceous sauropod dinosaurs of Zhejiang Province, China: impact on Laurasian titanosauriform phylogeny and biogeography. Royal Society Open Science 6:8, pages 191057.
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David W. Krause, Joseph J.W. SertichPatrick M. O'ConnorKristina Curry RogersRaymond R. Rogers. (2019) The Mesozoic Biogeographic History of Gondwanan Terrestrial Vertebrates: Insights from Madagascar's Fossil Record. Annual Review of Earth and Planetary Sciences 47:1, pages 519-553.
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Prince C. Owusu Agyemang, Eric M. Roberts, Robert Bussert, David Evans & Johannes Müller. (2019) U-Pb detrital zircon constraints on the depositional age and provenance of the dinosaur-bearing Upper Cretaceous Wadi Milk formation of Sudan. Cretaceous Research 97, pages 52-72.
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P. Mocho, A. Pérez-García, M. Martín Jiménez & F. Ortega. (2019) New remains from the Spanish Cenomanian shed light on the Gondwanan origin of European Early Cretaceous titanosaurs. Cretaceous Research 95, pages 164-190.
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Eric Gorscak & Patrick M. O’Connor. (2019) A new African Titanosaurian Sauropod Dinosaur from the middle Cretaceous Galula Formation (Mtuka Member), Rukwa Rift Basin, Southwestern Tanzania. PLOS ONE 14:2, pages e0211412.
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BERNARDO J. GONZÁLEZ RIGA, MATTHEW C. LAMANNA, ALEJANDRO OTERO, LEONARDO D. ORTIZ DAVID, ALEXANDER W.A. KELLNER & LUCIO M. IBIRICU. (2019) An overview of the appendicular skeletal anatomy of South American titanosaurian sauropods, with definition of a newly recognized clade. Anais da Academia Brasileira de Ciências 91:suppl 2.
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Verónica Díez Díaz, Géraldine Garcia, Xabier Pereda-Suberbiola, Benjamin Jentgen-Ceschino, Koen Stein, Pascal Godefroit & Xavier Valentin. (2018) The titanosaurian dinosaur Atsinganosaurus velauciensis (Sauropoda) from the Upper Cretaceous of southern France: New material, phylogenetic affinities, and palaeobiogeographical implications. Cretaceous Research 91, pages 429-456.
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Bernardo J Gonzàlez Riga, Philip D Mannion, Stephen F Poropat, Leonardo D Ortiz David & Juan Pedro Coria. (2018) Osteology of the Late Cretaceous Argentinean sauropod dinosaur Mendozasaurus neguyelap: implications for basal titanosaur relationships. Zoological Journal of the Linnean Society 184:1, pages 136-181.
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Y. Ranjeev Epa, Alycia L. Stigall, Eric M. Roberts, Haley D. O'Brien & Nancy J. Stevens. (2018) Morphological diversification of ampullariid gastropods (Nsungwe Formation, Late Oligocene, Rukwa Rift Basin, Tanzania) is coincident with onset of East African rifting. Papers in Palaeontology 4:3, pages 327-348.
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