Acknowledgments
Yong-Fu Qiu, Yi-Yu Yan, and Bin Lu contributed equally to this work. This study was supported by the National Natural Science Foundation of China (Nos. 31600740 and 81903427), the Natural Science Foundation of Jiangsu Province (BK20160443), the Six Talent Peaks Project in Jiangsu Province (SWYY-094), the Jiangsu Provincial Key Laboratory for Bioresources of Saline Soils (Nos. JKLBS2016013 and JKLBS2017010) and the college students practice innovation training program of Yancheng Teachers University (Provincial key projects).