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Articles

New stegosaurs from the Middle Jurassic Lower Member of the Shaximiao Formation of Chongqing, China

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Article: e1995737 | Received 06 Oct 2020, Accepted 30 Aug 2021, Published online: 03 Mar 2022

Figures & data

FIGURE 1. Maps showing the location where the Bashanosaurus primitivus was recovered. A, sketch map of China showing the location of Chongqing Municipality. B, sketch map of Yunyang County. C, sketch map of the fossil locality. D, stratigraphic chart with Lower Member of the Shaximiao Formation in Yunyang County, Chongqing Municipality, China.

FIGURE 1. Maps showing the location where the Bashanosaurus primitivus was recovered. A, sketch map of China showing the location of Chongqing Municipality. B, sketch map of Yunyang County. C, sketch map of the fossil locality. D, stratigraphic chart with Lower Member of the Shaximiao Formation in Yunyang County, Chongqing Municipality, China.

FIGURE 2. Lithological section of the Yunyang dinosaur locality in the Lower Member of the Shaximiao Formation in this study (left). AD: concordia diagrams of detrital zircon U-Pb ages for the samples analyzed in this study; EF: histogram of zircons for the samples analyzed in this study.

FIGURE 2. Lithological section of the Yunyang dinosaur locality in the Lower Member of the Shaximiao Formation in this study (left). A–D: concordia diagrams of detrital zircon U-Pb ages for the samples analyzed in this study; E–F: histogram of zircons for the samples analyzed in this study.

FIGURE 3. A, photograph (top) and interpretative sketch (bottom) of Bashanosaurus primitivus (CLGPR V00006-1) on the west side of the Dinosaur Fossil Wall. B, photograph (top) and interpretative sketch (bottom) of Bashanosaurus primitivus (CLGPR V00006-2) in the middle of the Dinosaur Fossil Wall. Abbreviations: cav, caudal vertebra; co, coracoid; dv, dorsal vertebra; f, femur; fl, fibula; mt, metatarsal; pl, plate; r, rib; sc, scapula; sp, spine; t, tibia.

FIGURE 3. A, photograph (top) and interpretative sketch (bottom) of Bashanosaurus primitivus (CLGPR V00006-1) on the west side of the Dinosaur Fossil Wall. B, photograph (top) and interpretative sketch (bottom) of Bashanosaurus primitivus (CLGPR V00006-2) in the middle of the Dinosaur Fossil Wall. Abbreviations: cav, caudal vertebra; co, coracoid; dv, dorsal vertebra; f, femur; fl, fibula; mt, metatarsal; pl, plate; r, rib; sc, scapula; sp, spine; t, tibia.

FIGURE 4. A–F, dorsal vertebrae referred to Bashanosaurus primitivus (CLGPR V00006-3) from the Middle Jurassic of China. A, anterior; B, left lateral; C, dorsal; D, posterior; E, right lateral; F, ventral; G, posterior views of dorsal vertebrae referred to Bashanosaurus primitivus (CLGPR V00006-1) from the Middle Jurassic of China. H, posterior view of dorsal vertebrae referred to Bashanosaurus primitivus (CLGPR V00006-2) from the Middle Jurassic of China. I, anterior view of dorsal vertebrae referred to Bashanosaurus primitivus (CLGPR V00006-2) from the Middle Jurassic of China. J–O, anterior caudal vertebrae referred to Bashanosaurus primitivus (CLGPR V00006-1) from the Middle Jurassic of China. J, anterior; K, left lateral; L, dorsal; M, posterior; N, right lateral; O, ventral. P–R, middle caudal vertebrae referred to Bashanosaurus primitivus (CLGPR V00006-1) from the Middle Jurassic of China. P, right lateral; Q, posterior; R, ventral. Abbreviations: acpl, anterior centroparapophyseal lamina; dia, diapophysis; ipzs, intraprezygapophyseal shelf; nc, neural canal; ns, neural spine; para, parapophysis; podl, postzygodiapophyseal lamina; pozyg, postzygapophysis; ppdl, paradiapophyseal lamina; przyg, prezygapophysis.

FIGURE 4. A–F, dorsal vertebrae referred to Bashanosaurus primitivus (CLGPR V00006-3) from the Middle Jurassic of China. A, anterior; B, left lateral; C, dorsal; D, posterior; E, right lateral; F, ventral; G, posterior views of dorsal vertebrae referred to Bashanosaurus primitivus (CLGPR V00006-1) from the Middle Jurassic of China. H, posterior view of dorsal vertebrae referred to Bashanosaurus primitivus (CLGPR V00006-2) from the Middle Jurassic of China. I, anterior view of dorsal vertebrae referred to Bashanosaurus primitivus (CLGPR V00006-2) from the Middle Jurassic of China. J–O, anterior caudal vertebrae referred to Bashanosaurus primitivus (CLGPR V00006-1) from the Middle Jurassic of China. J, anterior; K, left lateral; L, dorsal; M, posterior; N, right lateral; O, ventral. P–R, middle caudal vertebrae referred to Bashanosaurus primitivus (CLGPR V00006-1) from the Middle Jurassic of China. P, right lateral; Q, posterior; R, ventral. Abbreviations: acpl, anterior centroparapophyseal lamina; dia, diapophysis; ipzs, intraprezygapophyseal shelf; nc, neural canal; ns, neural spine; para, parapophysis; podl, postzygodiapophyseal lamina; pozyg, postzygapophysis; ppdl, paradiapophyseal lamina; przyg, prezygapophysis.

FIGURE 5. A, lateral view of right scapula and coracoid referred to Bashanosaurus primitivus (CLGPR V00006-1) from the Middle Jurassic of China. B, C, E, dermal armor referred to Bashanosaurus primitivus (CLGPR V00006-1) from the Middle Jurassic of China. D, dermal armor referred to Bashanosaurus primitivus (CLGPR V00006-2) from the Middle Jurassic of China. BD, lateral view of plates; E, lateral view of spine. Abbreviations: ap, acromial process; bl, scapula blade; cor, coracoid; glf, glenoid fossa; pp, scapula proximal plate.

FIGURE 5. A, lateral view of right scapula and coracoid referred to Bashanosaurus primitivus (CLGPR V00006-1) from the Middle Jurassic of China. B, C, E, dermal armor referred to Bashanosaurus primitivus (CLGPR V00006-1) from the Middle Jurassic of China. D, dermal armor referred to Bashanosaurus primitivus (CLGPR V00006-2) from the Middle Jurassic of China. B–D, lateral view of plates; E, lateral view of spine. Abbreviations: ap, acromial process; bl, scapula blade; cor, coracoid; glf, glenoid fossa; pp, scapula proximal plate.

FIGURE 6. AD, hind limb referred to Bashanosaurus primitivus (CLGPR V00006-1) from the Middle Jurassic of China. A, left femur in posterior view; B, left tibia in medial view; C, left fibula in lateral view; D, left metatarsal III in lateral view. EJ, right tibia referred to Bashanosaurus primitivus (CLGPR V00006-2) from the Middle Jurassic of China. E, anterior; F, lateral; G, posterior; H, media; I, dorsal; J, ventral. K, lateral view of fibula referred to Bashanosaurus primitivus (CLGPR V00006-2) from the Middle Jurassic of China. Abbreviations: cc, cnemial crest; ft, fourth trochanter; gr, groove; gt, greater trochanter; hd, head; lm, lateral malleolus; lp, lateral process; mm, medial malleolus; ri, ridge.

FIGURE 6. A–D, hind limb referred to Bashanosaurus primitivus (CLGPR V00006-1) from the Middle Jurassic of China. A, left femur in posterior view; B, left tibia in medial view; C, left fibula in lateral view; D, left metatarsal III in lateral view. E–J, right tibia referred to Bashanosaurus primitivus (CLGPR V00006-2) from the Middle Jurassic of China. E, anterior; F, lateral; G, posterior; H, media; I, dorsal; J, ventral. K, lateral view of fibula referred to Bashanosaurus primitivus (CLGPR V00006-2) from the Middle Jurassic of China. Abbreviations: cc, cnemial crest; ft, fourth trochanter; gr, groove; gt, greater trochanter; hd, head; lm, lateral malleolus; lp, lateral process; mm, medial malleolus; ri, ridge.

FIGURE 7. A–F, braincase referred to Stegosauria gen. et sp. indet. (CLGPRV0005-4) from the Middle Jurassic of China. A, anterior; B, left lateral; C, right lateral; D, posterior; E, dorsal; F, ventral. G–L, dorsal vertebrae referred to Stegosauria gen. et sp. indet. (CLGPRV0005-4) from the Middle Jurassic of China. G, anterior; H, left lateral; I, dorsal; J, posterior; K, right lateral; L, ventral. M–R, dermal armor referred to Stegosauria gen. et sp. indet. (CLGPRV0005-4) from the Middle Jurassic of China. M, anterior; N, posterior; O, left lateral; P, right lateral; Q, dorsal; R, ventral. Abbreviations: boc, basioccipital; bpt, basipterygoid process; bsp, basiphenoid; cc, carotid canal; c.n.I, III, V, VII, XII exit for cranial nerves; dia, diapophysis; f, frontal; fime, fissura metotica; fm, foramen magnum; fo, foramen ovale; ipzs, intraprezygapophyseal shelf; ist, integumentary structure; nc, neural canal; ns, neural spine; oc, occipital condyle; os, orbitosphenoid; p, parietal; para, parapophysis; poc, paroccipital process; pozyg, postzygapophysis; przyg, prezygapophysis; ps, parasphenoid; soc, supraoccipital; sf, supratemporal fossae; sq, squamosal.

FIGURE 7. A–F, braincase referred to Stegosauria gen. et sp. indet. (CLGPRV0005-4) from the Middle Jurassic of China. A, anterior; B, left lateral; C, right lateral; D, posterior; E, dorsal; F, ventral. G–L, dorsal vertebrae referred to Stegosauria gen. et sp. indet. (CLGPRV0005-4) from the Middle Jurassic of China. G, anterior; H, left lateral; I, dorsal; J, posterior; K, right lateral; L, ventral. M–R, dermal armor referred to Stegosauria gen. et sp. indet. (CLGPRV0005-4) from the Middle Jurassic of China. M, anterior; N, posterior; O, left lateral; P, right lateral; Q, dorsal; R, ventral. Abbreviations: boc, basioccipital; bpt, basipterygoid process; bsp, basiphenoid; cc, carotid canal; c.n.I, III, V, VII, XII exit for cranial nerves; dia, diapophysis; f, frontal; fime, fissura metotica; fm, foramen magnum; fo, foramen ovale; ipzs, intraprezygapophyseal shelf; ist, integumentary structure; nc, neural canal; ns, neural spine; oc, occipital condyle; os, orbitosphenoid; p, parietal; para, parapophysis; poc, paroccipital process; pozyg, postzygapophysis; przyg, prezygapophysis; ps, parasphenoid; soc, supraoccipital; sf, supratemporal fossae; sq, squamosal.

FIGURE 8. A single most parsimonious tree recovered by the phylogenetic analysis. The numbers below the line are Bremer supports, the numbers above the line are GC frequencies. Ankylosauria was represented by three taxa, Gastonia, Euoplocephalus, and Sauropelta in the analysis, but these are shown here as a single terminal taxon for clarity.

FIGURE 8. A single most parsimonious tree recovered by the phylogenetic analysis. The numbers below the line are Bremer supports, the numbers above the line are GC frequencies. Ankylosauria was represented by three taxa, Gastonia, Euoplocephalus, and Sauropelta in the analysis, but these are shown here as a single terminal taxon for clarity.
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