Figures & data
Figure 1 Photographs of well preserved enantiornithine skulls: A, Rapaxavis pani, DNHM D2522; B, Longipteryx sp., DNHM D2889; C, Pengornis houi, IVPP V15336; D, Shenqiornis mengi, DNHM D2950; E, Longipteryx chaoyangensis, IVPP V12552; F, Cathayornis yandica, IVPP V9769; G, juvenile Euenantiornithes indet, GMV 2158. All scale bars = 1 cm.
![Figure 1 Photographs of well preserved enantiornithine skulls: A, Rapaxavis pani, DNHM D2522; B, Longipteryx sp., DNHM D2889; C, Pengornis houi, IVPP V15336; D, Shenqiornis mengi, DNHM D2950; E, Longipteryx chaoyangensis, IVPP V12552; F, Cathayornis yandica, IVPP V9769; G, juvenile Euenantiornithes indet, GMV 2158. All scale bars = 1 cm.](/cms/asset/6b192954-82c2-41fe-8433-8aaf0cb7094f/tjsp_a_526639_o_f0001ag.gif)
![Figure 1 Photographs of well preserved enantiornithine skulls: A, Rapaxavis pani, DNHM D2522; B, Longipteryx sp., DNHM D2889; C, Pengornis houi, IVPP V15336; D, Shenqiornis mengi, DNHM D2950; E, Longipteryx chaoyangensis, IVPP V12552; F, Cathayornis yandica, IVPP V9769; G, juvenile Euenantiornithes indet, GMV 2158. All scale bars = 1 cm.](/cms/asset/aeb8870e-1dfe-455e-9e0e-f537ed154c83/tjsp_a_526639_o_f0001bg.gif)
Figure 2 Camera lucida drawings of select enantiornithine skulls: A, Rapaxavis pani, DNHM D2522; B, Longirostravis hani, IVPP V11309; C, Longipteryx chaoyangensis, IVPP V12325; D, Eoenantiornis buhleri, IVPP 11537; E, Longipteryx sp., DNHM D2889; F, Longipteryx chaoyangensis, IVPP V12552; G, Shenqiornis mengi, DNHM D2951; H, Pengornis houi, IVPP 15336; I, Alethoalaornis agitornis, LPM 00009; J, Cathayornis yandica, IVPP V9769; K, Dapingfangornis sentisorhinus, LPM 00039; L, M, Gobipteryx minuta, IGM-100/1011, right and left lateral views.
![Figure 2 Camera lucida drawings of select enantiornithine skulls: A, Rapaxavis pani, DNHM D2522; B, Longirostravis hani, IVPP V11309; C, Longipteryx chaoyangensis, IVPP V12325; D, Eoenantiornis buhleri, IVPP 11537; E, Longipteryx sp., DNHM D2889; F, Longipteryx chaoyangensis, IVPP V12552; G, Shenqiornis mengi, DNHM D2951; H, Pengornis houi, IVPP 15336; I, Alethoalaornis agitornis, LPM 00009; J, Cathayornis yandica, IVPP V9769; K, Dapingfangornis sentisorhinus, LPM 00039; L, M, Gobipteryx minuta, IGM-100/1011, right and left lateral views.](/cms/asset/b70ce040-a6f3-4a9a-8335-709fd3bb3871/tjsp_a_526639_o_f0002ag.jpg)
![Figure 2 Camera lucida drawings of select enantiornithine skulls: A, Rapaxavis pani, DNHM D2522; B, Longirostravis hani, IVPP V11309; C, Longipteryx chaoyangensis, IVPP V12325; D, Eoenantiornis buhleri, IVPP 11537; E, Longipteryx sp., DNHM D2889; F, Longipteryx chaoyangensis, IVPP V12552; G, Shenqiornis mengi, DNHM D2951; H, Pengornis houi, IVPP 15336; I, Alethoalaornis agitornis, LPM 00009; J, Cathayornis yandica, IVPP V9769; K, Dapingfangornis sentisorhinus, LPM 00039; L, M, Gobipteryx minuta, IGM-100/1011, right and left lateral views.](/cms/asset/e6636712-6b7b-4fdf-b469-4b4d3e7e9311/tjsp_a_526639_o_f0002bg.jpg)
Table 1 A list of known enantiornithine specimens that preserve skull material; grey indicates unpublished material.
Figure 3 Interpretive drawings of select juvenile enantiornithine skulls: A, GMV 2159; B, C, LP 4450 slab and counterslab. Anatomical abbreviations: cmf, caudal mandibular fenestra; den, dentary; frn, frontal; jug, jugal; la, lacrimal; max, maxilla; nas, nasal; oc, occipital condyle; pmx, premaxilla; po, postorbital; qd, quadrate; rmf, rostral mandibular fenestra; sq, squamosal; sur, surangular.
![Figure 3 Interpretive drawings of select juvenile enantiornithine skulls: A, GMV 2159; B, C, LP 4450 slab and counterslab. Anatomical abbreviations: cmf, caudal mandibular fenestra; den, dentary; frn, frontal; jug, jugal; la, lacrimal; max, maxilla; nas, nasal; oc, occipital condyle; pmx, premaxilla; po, postorbital; qd, quadrate; rmf, rostral mandibular fenestra; sq, squamosal; sur, surangular.](/cms/asset/4ae99cad-bc67-4bfa-8548-2c0e9d9c520e/tjsp_a_526639_o_f0003g.gif)
Figure 4 Close-up of the skull of Eoenantiornis buhleri IVPP showing the two alternate interpretations of the specimen: A, the S-shaped bone fragment is inferred to be part of the maxilla as in CitationZhou et al. (2005); B, the S-shaped bone as the lacrimal. See for anatomical abbreviations.
![Figure 4 Close-up of the skull of Eoenantiornis buhleri IVPP showing the two alternate interpretations of the specimen: A, the S-shaped bone fragment is inferred to be part of the maxilla as in CitationZhou et al. (2005); B, the S-shaped bone as the lacrimal. See Fig. 3 for anatomical abbreviations.](/cms/asset/96847c6c-9605-455b-8092-ee3308f60f84/tjsp_a_526639_o_f0004g.gif)
Figure 5 Basicrania: A, the Early Cretaceous Shenqiornis mengi, DNHM D2951; B, the Late Cretaceous Neuquenornis volans, MUCPv 142. Abbreviations: boc, basioccipital; bt, basal tubera; cerebral prominence; eoc, exoccipital; fm, foramen magnum; fov, foramen for the external occipital vein; nh, canals for the hypglossus nerve; oc, occipital condyle; ocoe, ostium canalis ophthalmici externi; pop, paraoccipital process; soc, supraoccipital.
![Figure 5 Basicrania: A, the Early Cretaceous Shenqiornis mengi, DNHM D2951; B, the Late Cretaceous Neuquenornis volans, MUCPv 142. Abbreviations: boc, basioccipital; bt, basal tubera; cerebral prominence; eoc, exoccipital; fm, foramen magnum; fov, foramen for the external occipital vein; nh, canals for the hypglossus nerve; oc, occipital condyle; ocoe, ostium canalis ophthalmici externi; pop, paraoccipital process; soc, supraoccipital.](/cms/asset/71cbd8d5-5a8d-4ec0-9f4c-42d63041ce3a/tjsp_a_526639_o_f0005g.gif)
Figure 6 Ternary diagram showing the relative proportions of skull units among Enantiornithes, relative to Neornithes. Abbreviations: B, braincase; O, orbit; R, rostrum.
![Figure 6 Ternary diagram showing the relative proportions of skull units among Enantiornithes, relative to Neornithes. Abbreviations: B, braincase; O, orbit; R, rostrum.](/cms/asset/06deabdf-8362-4b26-bca2-029d6d458c05/tjsp_a_526639_o_f0006g.gif)
Figure 7 Enantiornithine teeth, photographs and interpretative drawings: A, B, Eoenantiornis buhleri, IVPP 11537; C, D, Shenqiornis mengi, DNHM D2951; E, F, Pengornis houi, IVPP 15336; G, H, Longirostravis hani, IVPP V11309; I, J, Longipteryx chaoyangensis, IVPP V12552; K, L, Longipteryx sp., DNHM D2889.
![Figure 7 Enantiornithine teeth, photographs and interpretative drawings: A, B, Eoenantiornis buhleri, IVPP 11537; C, D, Shenqiornis mengi, DNHM D2951; E, F, Pengornis houi, IVPP 15336; G, H, Longirostravis hani, IVPP V11309; I, J, Longipteryx chaoyangensis, IVPP V12552; K, L, Longipteryx sp., DNHM D2889.](/cms/asset/9b8721a7-d89d-4249-ba1c-308ad1c12cd0/tjsp_a_526639_o_f0007g.gif)
Figure 8 Reconstructions of the skulls of enantiornithines: A, Shenqiornis mengi; B, Rapaxavis pani; C, Pengornis houi. All scale bars = 1 cm.
![Figure 8 Reconstructions of the skulls of enantiornithines: A, Shenqiornis mengi; B, Rapaxavis pani; C, Pengornis houi. All scale bars = 1 cm.](/cms/asset/b60b5acd-2f44-4231-9785-193278a67936/tjsp_a_526639_o_f0008g.gif)