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Articles

Homogenization techniques for population dynamics in strongly heterogeneous landscapes

ORCID Icon &
Pages 171-193 | Received 03 Aug 2017, Accepted 17 Nov 2017, Published online: 11 Dec 2017

Figures & data

Figure 1. Comparison of the travelling wave solution ρ with the leading order approximation ρ0 at time t=10 in the logistic growth example. The inset graph shows a ‘zoomed-in’ plot of the leading order approximation ρ0 (red dashed) and the numerical non-homogenized solution ρ (black). The main graph shows the numerical non-homogenized solution (black curve) and the upper and lower bound obtained from the homogenized solution. The top red curve is the upper bound g while the bottom red curve is the lower bound g/k. The parameters are D1=D2=1, l1=l2=0.1, λi=μi=1, and α=0.75 (k=3).

Figure 1. Comparison of the travelling wave solution ρ with the leading order approximation ρ0 at time t=10 in the logistic growth example. The inset graph shows a ‘zoomed-in’ plot of the leading order approximation ρ0 (red dashed) and the numerical non-homogenized solution ρ (black). The main graph shows the numerical non-homogenized solution (black curve) and the upper and lower bound obtained from the homogenized solution. The top red curve is the upper bound g while the bottom red curve is the lower bound g/k. The parameters are D1=D2=1, l1=l2=0.1, λi=μi=1, and α=0.75 (k=3).

Figure 2. The effect of varying l2 on the accuracy of the leading order approximation of the travelling wave profile. In plots (a–d) l2=1.9 and the numerical non-homogenized solution ρ (black) and leading order approximation ρ0 (red) are plotted as a function of the scaled spatial variable ξ (left column) and the unscaled spatial variable x (right column). The first row corresponds to the case where α=0.25 andthere is greater preference for patch 2. The second row corresponds to the case where α=0.75 and there is greater preference for patch 1. Finally, plot (e) shows the maximum absolute error (maxξ|ρρ0|) as l2 is varied. Plot (f) shows the averaged absolute error across rescaled space ξ. In all cases we ignore population dynamics with f1=f2=0 and D1=D2=1 and l1=0.1. We use a Gaussian initial condition and compare solutions at t=3.

Figure 2. The effect of varying l2 on the accuracy of the leading order approximation of the travelling wave profile. In plots (a–d) l2=1.9 and the numerical non-homogenized solution ρ (black) and leading order approximation ρ0 (red) are plotted as a function of the scaled spatial variable ξ (left column) and the unscaled spatial variable x (right column). The first row corresponds to the case where α=0.25 andthere is greater preference for patch 2. The second row corresponds to the case where α=0.75 and there is greater preference for patch 1. Finally, plot (e) shows the maximum absolute error (maxξ|ρ−ρ0|) as l2 is varied. Plot (f) shows the averaged absolute error across rescaled space ξ. In all cases we ignore population dynamics with f1=f2=0 and D1=D2=1 and l1=0.1. We use a Gaussian initial condition and compare solutions at t=3.

Figure 3. Asymptotic wave speed plotted as a function of patch preference, α. The solid line illustrates the homogenized wave speed prediction (Equation Equation79) in the case λ1=λ2=1, and the dashed line illustrates the case λ1=1, λ2=0.5. The grey stars are the numerically simulated wave speeds obtained from solving the original non-homogenized Equations (Equation2), (Equation3), (Equation7). The solution was solved until t=50, and the last 30 time steps were used to estimate wave speed, taking a threshold of 0.01 to track the location of the wave front at each time point. In all cases the population dynamics are given by logistic growth (fi(ρ)=(λiμiρ)ρ), and parameters are D1=D2=1, μi=1 and l1=l2=1.

Figure 3. Asymptotic wave speed plotted as a function of patch preference, α. The solid line illustrates the homogenized wave speed prediction (Equation Equation79(79) c∗=2l1+l2l1+l2/kλ1l1+λ2l2/kl1/D1+(l2/k)/(D2/k2).(79) ) in the case λ1=λ2=1, and the dashed line illustrates the case λ1=1, λ2=0.5. The grey stars are the numerically simulated wave speeds obtained from solving the original non-homogenized Equations (Equation2(2) ∂τρ=Di∂y2ρ+ε2fi(ρ)for y∈(yi−1,yi),i∈Z.(2) ), (Equation3(3) Di+1∂yρ(yi+,τ)=Di∂yρ(yi−,τ).(3) ), (Equation7(7) ρ(yi+,τ)=kiρ(yi−,τ).(7) ). The solution was solved until t=50, and the last 30 time steps were used to estimate wave speed, taking a threshold of 0.01 to track the location of the wave front at each time point. In all cases the population dynamics are given by logistic growth (fi(ρ)=(λi−μiρ)ρ), and parameters are D1=D2=1, μi=1 and l1=l2=1.