ACKNOWLEDGMENTS
We are grateful to M. Traverse for collecting these fossils and to the Université Montpellier II for making these specimens available for study. We thank the OCP, LACM, National Museum of Natural History (Washington), Natural History Museum (London), and MHNL for access to comparative material. Some of the photographs were taken by D. Serrette and P. Loubry (Muséum National d’Histoire Naturelle, Paris). We thank the NMNH for providing photographs of USNM 425111. This work has been supported by the Marie Curie Intra European Fellowship to E.B. G. Mayr, C. Mourer-Chauviré, and T. Worthy provided useful comments on the manuscript.
Handling editor: Trevor Worthy