Acknowledgements
Thanks to Jonathan Cox (Natural England) for managing the study contract and to Nick Hodgetts (Portree, Scotland, UK), Dave Ottewell (Natural England, UK) and Lara Pike (Natural England, UK) for supplying various useful information. Thanks also to Ingleton Scenery Company for help with fieldwork logistics, and to reviewers of the manuscript for their valuable comments.
Disclosure statement
No potential conflicts of interest were reported by the authors.
Additional information
Funding
Notes on contributors
Des A. Callaghan
Des A. Callaghan is a consultant bryologist operating under Bryophyte Surveys Ltd, working throughout Britain and further afield. His research is focused on threatened species, taxonomy and conservation ecology.