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- Nicholas, W. L. and Kershaw, W. E. (1954). Studies on the intake of microfilariae by their insect vectors, their survival, and their effect on the survival of their vectors. III: The intake of the microfilariae of Acanthocheilonema perstans by Culicoides austeni and C. grahamii. Ibid., 48, 201.
- Nicholas, W. L. and Kershaw, W. E. Keay, R. W. J., and Zahra, A. (1953). Studies on the epidemiology of filariasis in West Africa, with special reference to the British Cameroons and the Niger delta. III: The distribution of Culicoides spp. biting man in the rain-forest, the forest fringe and the mountain grasslands of the British Cameroons. Ibid., 47, 95.
- Sharp, N. A. Dyce (1928). Filaria perstans; its development in Culicoides austeni. Trans. R. Soc. Trop. Med. Hyg., 21, 371.
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