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ARTICLES

Angonisaurus and Shansiodon, dicynodonts (Therapsida, Anomodontia) from subzone C of the Cynognathus Assemblage Zone (Middle Triassic) of South Africa

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Pages 655-676 | Received 05 Jun 2012, Accepted 17 Aug 2012, Published online: 07 May 2013

LITERATURE CITED

  • Abdala , F. and Ribeiro , A. M. 2010 . Distribution and diversity patterns of Triassic cynodonts (Therapsida, Cynodontia) in Gondwana . Palaeogeography, Palaeoclimatology, Palaeoecology , 286 : 202 – 217 .
  • Abdala , F. , Hancox , P. J. and Neveling , J. 2005 . Cynodonts from the uppermost Burgersdorp Formation (Karoo Supergroup), South Africa, and their bearing on the biostratigraphic subdivision and correlation of the Triassic Cynognathus Assemblage Zone . Journal of Vertebrate Paleontology , 25 : 192 – 199 .
  • Abdala , F. J. , Neveling and Welman , J. 2006 . A new trirachodontid cynodont from the lower levels of the Burgersdorp Formation (Lower Triassic) of the Beaufort Group, South Africa and the cladistic relationships of Gondwanan gomphodonts . Zoological Journal of the Linnean Society , 147 : 383 – 413 .
  • Anderson , J. M. and Cruickshank , A. R. I. 1978 . The biostratigraphy of the Permian and the Triassic. Part 5. A review of the classification and distribution of Permo-Triassic tetrapods . Palaeontologia africana , 21 : 15 – 44 .
  • Angielczyk , K. D. and Sullivan , C. 2008 . Diictodon feliceps (Owen, 1876), a dicynodont (Therapsida, Anomodontia) species with a Pangaean distribution . Journal of Vertebrate Paleontology , 28 : 788 – 802 .
  • Angielczyk , K. D. , Steyer , J.-S. , Sidor , C. A. , Smith , R. M. H. , Whatley , R. L. and Tolan , S. “ Permian and Triassic dicynodont (Therapsida: Anomodontia) faunas of the Luangwa Basin, Zambia: taxonomic update and implications for dicynodont biogeography and biostratigraphy ” . In The Early Evolutionary History of the Synapsida , Edited by: Kammerer , C. F. , Angielczyk , K. D. and Fröbisch , J. Dordrecht , , The Netherlands : Springer . In press
  • Battail , B. 1993 . “ On the biostratigraphy of Triassic therapsid-bearing formations ” . In The Nonmarine Triassic , Edited by: Lucas , S. G. and Morales , M. 31 – 35 . New Mexico Museum of Natural History and Science Bulletin 3 .
  • Broom , R. 1905 . On the use of the term Anomodontia . Albany Museum Records , 1 : 266 – 269 .
  • Butler , R. J. , Barrett , P. M. , Abel , R. L. and Gower , D. J. 2009 . A possible ctenosauriscid archosaur from the Middle Triassic Manda beds of Tanzania . Journal of Vertebrate Paleontology , 29 : 1022 – 1031 .
  • Butler , R. J. , Brusatte , S. L. , Reich , M. , Nesbitt , S. J. , Schoch , R. R. and Hornung , J. J. 2011 . The sail-backed reptile Ctenosauriscus from the latest Early Triassic of Germany and the timing and biogeography of the early archosaur radiation . PloS ONE , 6 : e25693
  • Catuneanu , O. , Wopfner , H. , Eriksson , P. G. , Cairncross , B. , Rubidge , B. S. , Smith , R. M. H. and Hancox , P. J. 2005 . The Karoo basins of south-central Africa . Journal of African Earth Sciences , 43 : 211 – 253 .
  • Cheng , Z. 1980 . “ Vertebrate animals ” . In Institute of Geology, Chinese Academy of Sciences (ed.), Mesozoic Stratigraphy and Paleontology of the Shansi-Gansu-Ningxia Basin, Volume 2 , 115 – 146 . Beijing , , China : Geological Publishing House .
  • Cheng , Z. 1981 . “ Permo-Triassic continental deposits and vertebrate faunas of China ” . In Gondwana Five , Edited by: Cresswell , M. M. and Vella , P. 65 – 70 . Rotterdam , , The Netherlands : A. A. Balkema .
  • Cluver , M. A. 1974 . The skull and mandible of a new cistecephalid dicynodont . Annals of the South African Museum , 64 : 137 – 155 .
  • Cooper , M. R. 1980 . “ The origins and classification of Triassic Dicynodonts ” . Edited by: Keyser , A. W. and Cruickshank , A. R. I. Vol. 83 , 107 – 110 . Discussion. Transactions of the Geological Society of South Africa
  • Cooper , M. R. 1982 . A mid-Permian to earliest Jurassic tetrapod biostratigraphy and its significance . Arnoldia , 9 ( 7 ) : 77 – 104 .
  • Cox , C. B. 1965 . New Triassic dicynodonts from South America, their origins and relationships . Philosophical Transactions of the Royal Society of London, Series B , 248 : 457 – 514 .
  • Cox , C. B. 1969 . Two new dicynodonts from the Triassic Ntawere Formation, Zambia . Bulletin of the British Museum (Natural History), Geology , 17 : 257 – 294 .
  • Cox , C. B. 1972 . “ A new digging dicynodont from the Upper Permian of Tanzania ” . In Studies in Vertebrate Evolution , Edited by: Joysey , K. A. and Kemp , T. S. 173 – 189 . Edinburgh : Oliver and Boyd .
  • Cox , C. B. 1991 . The Pangaea dicynodont Rechnisaurus and the comparative biostratigraphy of Triassic dicynodont faunas . Palaeontology , 34 : 767 – 784 .
  • Cox , C. B. and Li , J. 1983 . A new genus of Triassic dicynodont from east Africa and its classification . Palaeontology , 26 : 389 – 406 .
  • Cruickshank , A. R. I. On a specimen of the anomodont reptile Kannemeyeria latifrons (Broom) from the Manda Formation of Tanganyika, Tanzania . Proceedings of the Linnean Society of London , Vol. 172 , pp. 149 – 157 .
  • Cruickshank , A. R. I. 1967 . A new dicynodont genus from the Manda Formation of Tanzania (Tanganyika) . Journal of Zoology , 153 : 163 – 208 .
  • Cruickshank , A. R. I. 1986 . Biostratigraphy and classification of a new Triassic dicynodont from East Africa . Modern Geology , 10 : 121 – 131 .
  • Damiani , R. , Vasconcelos , C. , Renaut , A. , Hancox , J. and Yates , A. 2007 . Dolichuranus primaevus (Therapsida: Anomodontia) from the Middle Triassic of Namibia and its phylogenetic relationships . Palaeontology , 50 : 1531 – 1546 .
  • Domnanovich , N. S. and Marsicano , C. A. 2012 . The Triassic dicynodont Vinceria (Therapsida, Anomodontia) from Argentina and a discussion on basal Kannemeyeriiformes . Geobios , 45 : 173 – 186 .
  • Farabee , M. J. , Taylor , E. L. and Taylor , T. N. 1990 . Correlation of Permian and Triassic palynomorph assemblages from the central Transantarctic Mountains, Antarctica . Review of Palaeobotany and Palynology , 65 : 257 – 265 .
  • Fröbisch , J. 2008 . Global taxonomic diversity of anomodonts (Tetrapoda, Therapsida) and the terrestrial rock record across the Permian-Triassic boundary . PloS ONE , 35 : e3733
  • Fröbisch , J. 2009 . Composition and similarity of global anomodont-bearing tetrapod faunas . Earth Science Reviews , 95 : 119 – 157 .
  • Fröbisch , J. “ Synapsid diversity and the rock record in the Permian-Triassic Beaufort Group (Karoo Supergroup), South Africa ” . In The Early Evolutionary History of the Synapsida , Edited by: Kammerer , C. F. , Angielczyk , K. D. and Fröbsich , J. Dordrecht , , The Netherlands : Springer . In press
  • Fröbisch , J. , Angielczyk , K. D. and Sidor , C. A. 2010 . The Triassic dicynodont Kombuisia (Synapsida, Anomodontia) from Antarctica, a refuge from the terrestrial Permo-Triassic mass extinction . Naturwissenschaften , 97 : 187 – 196 .
  • Gao , K.-Q. , Fox , R. C. , Zhou , C.-F. and Li , D.-Q. 2010 . A new nonmammalian eucynodont (Synapsida: Therapsida) from the Triassic of northern Gansu Province, China, and its biostratigraphic and biogeographic implications . American Museum Novitates , 3685 : 1 – 25 .
  • Gay , S. A. and Cruickshank , A. R. I. 1999 . Biostratigraphy of the Permian tetrapod faunas from the Ruhuhu Valley, Tanzania . Journal of African Earth Sciences , 29 : 195 – 210 .
  • Govender , R. and Yates , A. 2009 . Dicynodont postcrania from the Triassic of Namibia and their implication for the systematics of Kannemeyeriiforme dicynodonts . Palaeontologia africana , 44 : 41 – 57 .
  • Groenewald , G. H. 1989 . Stratigrafie en sedimentologie van die Groep Beaufort in die Noordoos Vrystaat . Geological Survey South Africa Bulletin , 96 : 1 – 62 .
  • Hammer , W. R. 1990 . “ Triassic terrestrial vertebrate faunas of Antarctica ” . In Antarctic Paleobiology: Its Role in the Reconstruction of Gondwanaland , Edited by: Taylor , T. N. and Taylor , E. L. 42 – 50 . New York : Springer-Verlag .
  • Hammer , W. R. 1995 . New therapsids from the upper Fremouw Formation (Triassic) of Antarctica . Journal of Vertebrate Paleontology , 15 : 105 – 112 .
  • Hancox , P. J. 1998 . A stratigraphic, sedimentological and palaeoenvironmental synthesis of the Beaufort-Molteno contact in the Karoo Basin , 404 Johannesburg , , South Africa : University of the Witwatersrand . Ph.D. dissertation
  • Hancox , P. J. 2000 . The continental Triassic of South Africa . Zentralblatt für Geologie und Paläontologie , 1998 : 1285 – 1324 . Teil 1
  • Hancox , P. J. and Rubidge , B. S. 1994 . A new dicynodont therapsid from South Africa: implications for the biostratigraphy of the Upper Beaufort (Cynognathus Assemblage Zone) . South African Journal of Science , 90 : 98 – 99 .
  • Hancox , P. J. and Rubidge , B. S. 1996 . The first specimen of the mid-Triassic dicynodont Angonisaurus from the Karoo of South Africa: implications for the dating and biostratigraphy of the Cynognathus Assemblage Zone, Upper Beaufort Group . South African Journal of Science , 92 : 391 – 392 .
  • Hancox , P. J. and Rubidge , B. S. 2001 . Breakthroughs in the biodiversity, biostratigraphy, biogeography and basin analysis of the Beaufort Group . Journal of African Earth Sciences , 33 : 563 – 577 .
  • Hancox , P. J. , Shishkin , M. A. , Rubidge , B. S. and Kitching , J. W. 1995 . A threefold subdivision of the Cynognathus Assemblage Zone (Beaufort Group, South Africa) and its palaeogeographical implications . South African Journal of Science , 91 : 143 – 144 .
  • Hopson , J. A. In press . “ The traversodontid cynodont Mandagomphodon (“Scalenodon”) hirschsoni, Crompton, 1972, from the Middle Triassic of the Ruhuhu Valley, Tanzania ” . In The Early Evolutionary History of the Synapsida , Edited by: Kammerer , C. F. , Angielczyk , K. D. and Fröbisch , J. Dordrecht , , The Netherlands : Springer .
  • Huene , F. von . 1935 . “ Die fossilen Reptilien des südamerikanischen Gondwanalandes and der Zeitenwende ” . In Ergebnisse der Sauriergrabungen in Südbrasilien 1928/1929 , 92 Tübingen , , Germany : Lieferung I. F. F. Heine .
  • Huene , F. von . 1942 . Die Anomodontier des Ruhuhu-Gebietes in der Tübinger Sammlung . Palaeontographica, Abteilung A , 44 : 154 – 184 .
  • Huene , F. von . 1948 . “ Short review of the lower tetrapods ” . In Robert Broom Commemorative Volume , Edited by: Du Toit , A. L. 65 – 105 . Cape Town , , South Africa : Royal Society of South Africa .
  • Kammerer , C. F. , Angielczyk , K. D. and Fröbisch , J. 2011 . A comprehensive taxonomic revision of Dicynodon (Therapsida, Anomodontia), and its implications for dicynodont phylogeny, biogeography, and biostratigraphy . Memoir of the Society of Vertebrate Paleontology , 11 : 1 – 158 .
  • Keyser , A. W. 1973 . A preliminary study of the type area of the Cistecephalus Zone of the Beaufort Series, and a revision of the anomodont family Cistecephalidae . Geological Survey of South Africa Memoir , 62 : 1 – 71 .
  • Keyser , A. W. and Cruickshank , A. R. I. 1979 . The origins and classification of Triassic dicynodonts . Transactions of the Geological Society of South Africa , 82 : 81 – 108 .
  • King , G. M. 1988 . “ Anomodontia ” . In Handbook of Paleoherpetology, Part 17C , Edited by: Wellnhofer , P. 174 Stuttgart , , Germany : Gustav Fischer .
  • Kitching , J. W. 1977 . The distribution of the Karroo vertebrate fauna . Bernard Price Institute for Palaeontological Research Memoir , 1 : 1 – 131 .
  • Larkin , N. 1994 . Description of a new Triassic dicynodont from the Manda Formation of Tanzania , 104 London , , U.K. : University College London . M.Sc. thesis
  • Liu , J. and Abdala , F. In press . “ Phylogeny and taxonomy of the Traversodontidae ” . In The Early Evolutionary History of the Synapsida , Edited by: Kammerer , C. F. , Angielczyk , K. D. and Fröbisch , J. Dordrecht , , The Netherlands : Springer .
  • Lucas , S. G. 1993a . The Shansiodon Biochron, nonmarine Middle Triassic of Pangaea . Albertiana , 11 : 40 – 42 .
  • Lucas , S. G. 1993b . “ Vertebrate biochronology of the Triassic of China ” . In The Nonmarine Triassic , Edited by: Lucas , S. G. and Morales , M. 301 – 306 . New Mexico Museum of Natural History and Science Bulletin 3 .
  • Lucas , S. G. 1996 . Vertebrate biochronology of the Mesozoic of China . Memoirs of the Beijing Natural History Museum , 55 : 110 – 148 .
  • Lucas , S. G. 1998 . Global Triassic tetrapod biostratigraphy and biochonology . Palaeogeography Palaeoclimatology, Palaeoecology , 143 : 347 – 384 .
  • Lucas , S. G. 2001 . Chinese Fossil Vertebrates , 375 New York : Columbia University Press .
  • Lucas , S. G. 2010 . “ The Triassic timescale based on nonmarine tetrapod biostratigraphy and biogeography ” . In The Triassic Timescale. Geological Society of London Edited by: Lucas , S. G. 447 – 500 . Special Publication 334
  • Lucas , S. G. and Wild , R. 1995 . A Middle Triassic dicynodont from Germany and the biochronology of Triassic dicynodonts . Stuttgarter Beiträge zur Naturkunde, Serie B (Geologie und Paläontologie) , 220 : 1 – 16 .
  • Lucas , S. G. , Hunt , A. P. , Heckert , A. B. and Spielmann , J. A. 2007 . “ Global Triassic tetrapod biostratigraphy and biochonology: 2007 status ” . In The Global Triassic , Edited by: Lucas , S. G. and Spielmann , J. A. 229 – 240 . New Mexico Museum of Natural History and Science Bulletin 41 .
  • Maisch , M. W. 2001 . Observations on Karoo and Gondwana vertebrates. Part 2: a new skull-reconstruction of Stahleckeria potens von Huene, 1935 (Dicynodontia, Middle Triassic) and a reconsideration of kannemeyeriiform phylogeny . Neues Jahrbuch für Geologie und Paläontologie Abhandlungen , 220 : 127 – 152 .
  • Markwort , S. 1991 . Sedementation und Diagenese der untertriadischen Karoo Schichten des Ruhuhu Beckens, SW Tanzania . Geologisches Institut, Universität Köln, Sonderveröffentlichungen , 80 : 1 – 118 .
  • Nesbitt , S. J. 2011 . The early evolution of archosaurs: relationships and the origin of major clades . Bulletin of the American Museum of Natural History , 352 : 1 – 292 .
  • Nesbitt , S. J. and Angielczyk , K. D. 2002 . New evidence of large dicynodonts in the Upper Moenkopi Formation (Middle Triassic) of northern Arizona . PaleoBios , 22 : 10 – 17 .
  • Neveling , J. 2004 . Stratigraphic and sedimentological investigation of the contact between the Lystrosaurus and the Cynognathus assemblage zones (Beaufort Group: Karoo Supergroup) . Council for Geoscience Bulletin , 137 : 1 – 164 .
  • Nicolas , M. and Rubidge , B. S. 2010 . Changes in Permo-Triassic terrestrial tetrapod ecological representation in the Beaufort Group (Karoo Supergroup) of South Africa . Lethaia , 43 : 45 – 59 .
  • Ochev , V. G. and Shishkin , M. A. 1989 . On the principles of global correlation of the continental Triassic on the tetrapods . Acta Palaeontologica Polonica , 34 : 149 – 173 .
  • Owen , R. 1860 . On some reptilian fossils from South Africa . Quarterly Journal of the Geological Society of London , 16 : 49 – 63 .
  • Rayfield , E. J. , Barrett , P. M. and Milner , A. R. 2009 . Utility and validity of Middle and Late Triassic ‘land vertebrate faunachrons.’ Journal of Vertebrate Paleontology , 29 : 80 – 87 .
  • Rayfield , E. J. , Barrett , P. M. , McDonnell , R. A. and Willis , K. J. 2005 . A geographical information system (GIS) study of Triassic vertebrate biochronology . Geological Magazine , 142 : 327 – 354 .
  • Renaut , A. J. 2000 . A re-evaluation of the cranial morphology and taxonomy of the Triassic dicynodont genus Kannemeyeria , 191 Johannesburg , , South Africa : University of the Witwatersrand . Ph.D. dissertation
  • Rubidge , B. S. 1995 . Biostratigraphy of the Beaufort Group (Karoo Supergroup) . South African Committee for Stratigraphy Biostratigraphic Series , 1 : 1 – 46 . ed.
  • Rubidge , B. S . 2005 . Re-uniting lost continents—fossil reptiles from the ancient Karoo and their wanderlust . South African Journal of Geology , 108 : 135 – 172 .
  • Sahney , S. and Benton , M. J. Recovery from the most profound mass extinction of all time . Proceedings of the Royal Society B , Vol. 275 , pp. 759 – 765 .
  • Shishkin , M. A. , Rubidge , B. S. and Hancox , P. J. 1995 . “ Vertebrate biozonation of the Upper Beaufort Series of South Africa—a new look on correlation of the Triassic biotic events in Euramerica and southern Gondwana ” . In Sixth Symposium on Mesozoic Terrestrial Ecosystems, Short Papers , Edited by: Sun , A.-L. and Wang , Y. 39 – 41 . Beijing , , China : China Ocean Press .
  • Sidor , C. A. , Damiani , R. and Hammer , W. R. 2008 . A new Triassic temnospondyl from Antarctica and a review of Fremouw Formation biostratigraphy . Journal of Vertebrate Paleontology , 28 : 656 – 663 .
  • Sidor , C. A. , Steyer , J.-S. and Damiani , R. 2007 . Parotosuchus (Temnospondyli: Mastodonsauridae) from the Triassic of Antarctica . Journal of Vertebrate Paleontology , 27 : 232 – 235 .
  • Smith , R. , Rubidge , B. and van der Walt , M. 2011 . “ Therapsid biodiveristy patterns and paleoenvironments of the Karoo Basin, South Africa ” . In Forerunners of Mammals: Radiation, Histology, Biology , Edited by: Chinsamy-Turan , A. 31 – 62 . Bloomington , Indiana : Indiana University Press .
  • Stockley , G. M. 1932 . The geology of the Ruhuhu coalfields, Tanganyika Territory . Quarterly Journal of the Geological Society of London , 88 : 610 – 622 .
  • Sues , H.-D. and Fraser , N. C. 2010 . Triassic Life on Land: The Great Transition , 263 New York : Columbia University Press .
  • Surkov , M. V. and Benton , M. J. 2004 . The basicranium of dicynodonts (Synapsida) and its use in phylogenetic analysis . Palaeontology , 47 : 619 – 638 .
  • Surkov , M. V. and Benton , M. J. 2008 . Head kinematics and feeding adaptations of the Permian and Triassic dicynodonts . Journal of Vertebrate Paleontology , 28 : 1120 – 1129 .
  • Vega-Dias , C. , Maisch , M. W. and Schultz , C. L. 2004 . A new phylogenetic analysis of Triassic dicynodonts (Therapsida) and the systematic position of Jachaleria candelariensis from the Upper Triassic of Brazil . Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen , 231 : 145 – 166 .
  • Welman , J. , Groenewald , G. H. and Kitching , J. W. 1991 . Confirmation of the occurrence of Cynognathus zone deposits (uppermost Beaufort Group) in the north-eastern Orange Free Stata, South Africa . South African Journal of Geology , 94 : 245 – 248 .
  • Watson , D. M. S. 1942 . On the Permian and Triassic tetrapods . Geological Magazine , 79 : 81 – 116 .
  • Wopfner , H. 2002 . Tectonic and climatic events controlling deposition in Tanzanian Karoo basins . Journal of African Earth Sciences , 34 : 167 – 177 .
  • Yeh , H.-K. 1959 . New dicynodont from Sinokannemeyeria-fauna from Shansi . Vertebrata PalAsiatica , 3 : 187 – 204 .
  • Young , C. C. 1959 . Note on the first cynodont from the Sinokannemeyeria faunas in Shansi, China . Vertebrata PalAsiatica , 3 : 124 – 132 .

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