New material and phylogenetic position of Arenysaurus ardevoli, a lambeosaurine dinosaur from the late Maastrichtian of Arén (northern Spain)

LITERATURE CITED

• Blain , H. A. , Canudo , J. I. , Cuenca-Bescós , G. and López-Martínez , N. 2010 . Amphibians and squamates from the latest Maastrichtian (Upper Cretaceous) of Blasi 2 (Huesca, Spain) . Cretaceous Research , 31 : 433 – 446 .
   Web of Science ®Google Scholar
• Brett-Surman , M. K. 1979 . Phylogeny and paleobiogeography of hadrosaurian dinosaurs . Nature , 277 : 560 – 562 .
   Web of Science ®Google Scholar
• Brett-Surman , M. K. and Wagner , J. R. 2006 . “ Discussion of character analysis of the appendicular anatomy in Campanian and Maastrichtian North American hadrosaurids—variation and ontogeny ” . In Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs , Edited by: Carpenter , K. 125 – 169 . Bloomington and Indianapolis , Indiana : Indiana University Press .
   Google Scholar
• Brown , B. 1912 . A crested dinosaur from the Edmonton Cretaceous . Bulletin of the American Museum of Natural History , 31 : 131 – 136 .
   Google Scholar
• Brown , B. 1914 . Corythosaurus casuarius, a new crested dinosaur from the Belly River Cretaceous, with provisional classification of the family Trachodontidae . Bulletin of the American Museum of Natural History , 33 : 559 – 565 .
   Google Scholar
• Canudo , J. I. , Cruzado-Caballero , P. and Moreno-Azanza , M. 2005 . Possible theropod predation evidence in hadrosaurid dinosaurs from the upper Maastrichtian (Upper Cretaceous) of Arén (Huesca, Spain). Kaupia . Darmstädter Beiträge zur Naturgeschichte , 14 : 9 – 13 .
   Google Scholar
• Casanovas-Cladellas , M. L. , Santafé-Llopis , J. V. and Isidro-Llorens , A. 1993 . Pararhabdodon isonense n. gen. n. sp. (Dinosauria). Estudio morfológico, radio-tomográfico y consideraciones biomecánicas . Paleontogia i Evolució , 26–27 : 121 – 131 .
   Google Scholar
• Casanovas-Cladellas , M. L. , Pereda-Suberbiola , X. , Santafé , J. V. and Weishampel , D. B. 1999 . First lambeosaurine hadrosaurid from Europe: palaeobiogeographical implications . Geological Magazine , 136 : 205 – 211 .
   Web of Science ®Google Scholar
• Cope , E. D. 1870 . Synopsis of the extinct Batrachia, Reptilia, and Aves of North America . Transactions of the American Philosophical Society , 14 : 1 – 252 .
   Google Scholar
• Cruzado-Caballero , P. 2012 . “ Restos directos de dinosaurios hadrosáuridos (Ornithopoda, Hadrosauridae) del Maastrichtiense superior (Cretácico Superior) de Arén (Huesca) ” . 411 Zaragoza , , Spain : Ph. D. thesis, University of Zaragoza .
   Google Scholar
• Cruzado-Caballero , P. and Canudo , J. I. 2005 . Sobre la diversidad de los hadrosaurios en el Maastricthiense superior (Cretácico superior) de Arén (Huesca); pp. 115–116 in E. Bernáldez, E. Mayoral and A. Guerreiro dos Santos (eds.), XXI Jornadas de la Sociedad Española de Paleontología. Gestión e Investigación de la Paleontología en el Siglo XXI, Program and Abstracts, October 4–8, 2005, Sevilla .
   Google Scholar
• Cruzado-Caballero , P. , Canudo , J. I. and Ruiz-Omeñaca , J. I. 2005 . Nuevas evidencias de la presencia de hadrosaurios lambeosaurinos (Dinosauria) en el Maastrichtiense superior de la Península Ibérica (Arén, Huesca) . Geogaceta , 38 : 47 – 50 .
   Google Scholar
• Cruzado-Caballero , P. , Canudo , J. I. and Ruiz-Omeñaca , J. I. 2009 . “ Los fémures de Blasi (Arén, Huesca, Spain): una contribución a los hadrosauroideos europeos del Maastrichtiense superior ” . In Actas de las IV Jornadas Internacionales sobre Paleontología de Dinosaurios y su Entorno , Edited by: Hurtado , P. Huerta and Fernández-Baldor , F. Torcida . 197 – 205 . Burgos , , Spain : Colectivo Arqueológico y Paleontológico de Salas, Salas de los Infantes .
   Google Scholar
• Cruzado-Caballero , P. , Pereda-Suberbiola , X. and Ruiz-Omeñaca , J. I. 2010a . Blasisaurus canudoi gen. et sp. nov., a new lambeosaurine dinosaur (Hadrosauridae) from the latest Cretaceous of Arén (Huesca, Spain) . Canadian Journal of Earth Sciences , 47 : 1507 – 1517 .
   Web of Science ®Google Scholar
• Cruzado-Caballero , P. , Ruiz-Omeñaca , J. I. and Canudo , J. I. 2010b . Evidencias de la coexistencia de dinosaurios hadrosaurinos y lambeosaurinos en el Maastrichtiano superior de la Península Ibérica (Arén, Huesca, España) . Ameghiniana , 47 : 153 – 164 .
   Web of Science ®Google Scholar
• Cruzado-Caballero , P. , Canudo , J. I. , Moreno-Azanza , M. and Ruiz-Omeñaca , J. I. 2011 . “ The complex fauna of European Maastrichtian hadrosaurids: contributions of the lambeosaurines from the Iberian Peninsula ” . In International Hadrosaur Symposium at the Royal Tyrrell Museum of Palaeontology, September 22–23, 2011. Program and Abstracts , Edited by: Braman , D. R. , Eberth , D. A. , Evans , D. C. and Taylor , W. 33 – 37 . Drumheller : Royal Tyrrell Museum of Palaeontology .
   Google Scholar
• Cruzado-Caballero , P. , Ruiz-Omeñaca , J. I. , Gaete , R. , Riera , V. , Oms , O. and Canudo , J. I. 2013 . A new hadrosaurid dentary from the latest Maastrichtian of the Pyrenees (north Spain) and the high diversity of the duck-billed dinosaurs of the Ibero-Armorican Realm at the very end of the Cretaceous. Historical Biology
   Google Scholar
• Dalla Vecchia , F. M. 2009a . “ European hadrosauroids ” . In Actas de las IV Jornadas Internacionales sobre Paleontología de Dinosaurios y su Entorno , Edited by: Hurtado , P. Huerta and Fernández-Baldor , F. Torcida . 45 – 74 . Burgos , , Spain : Colectivo Arqueológico y Paleontológico de Salas, Salas de los Infantes .
   Google Scholar
• Dalla Vecchia , F. M. 2009b . Tethyshadros insularis, a new hadrosauroid dinosaur (Ornithischia) from the Upper Cretaceous of Italy . Journal of Vertebrate Paleontology , 29 : 1100 – 1116 .
   Web of Science ®Google Scholar
• Dilkes , D. W. 2000 . Appendicular myology of the hadrosaurian dinosaur Maiasaura peeblesorum from the Late Cretaceous (Campanian) of Montana . Transactions of the Royal Society of Edinburgh: Earth Sciences , 90 : 87 – 125 .
   Google Scholar
• Evans , D. C. and Reisz , R. R. 2007 . Anatomy and relationships of Lambeosaurus magnicristatus, a crested hadrosaurid dinosaur (Ornithischia) from the Dinosaur Park Formation, Alberta . Journal of Vertebrate Paleontology , 27 : 373 – 393 .
   Web of Science ®Google Scholar
• Evans , D. C. , Reisz , R. R. and Dupuis , K. 2007 . A juvenile Parasaurolophus (Ornithischia: Hadrosauridae) braincase from Dinosaur Provincial Park, Alberta, with comments on crest ontogeny in the genus . Journal of Vertebrate Paleontology , 27 : 642 – 650 .
   Web of Science ®Google Scholar
• Fiorillo , A. R. 2008 . “ Dinosaurs of Alaska: implications for the Cretaceous origin of Beringia ” . In The Terrane Puzzle: New Perspectives on Paleontology and Stratigraphy from the North American Cordillera Edited by: Blodgett , R. B. and Stanley , G. D. Jr. 313 – 326 . Special Paper of the Geological Society of America 442
   Google Scholar
• Gates , T. A. , Sampson , S. D. , Delgado de Jesús , C. R. , Zanno , L. E. , Eberth , D. , Hernández-Rivera , R. , Aguillón Martínez , M. C. and Kirkland , J. I. 2007 . Velafrons coahuilensis, a new lambeosaurine hadrosaurid (Dinosauria: Ornithopoda) from the late Campanian Cerro del pueblo Formation, Coahuila, México . Journal of Vertebrate Paleontology , 27 : 917 – 930 .
   Web of Science ®Google Scholar
• Gilmore , C. W. 1933 . On the dinosaurian fauna of the Iren Dabasu Formation . Bulletin of the American Museum of Natural History , 67 : 23 – 78 .
   Google Scholar
• Godefroit , P. , Bolotsky , Y. and Alifanov , V. 2003 . A remarkable hollow-crested hadrosaur from Russia: an Asian origin for lambeosaurines . Comptes Rendus Palevol , 2 : 143 – 151 .
   Web of Science ®Google Scholar
• Godefroit , P. , Bolotsky , Y. L. and Bolotsky , I. Y. 2012 . Osteology and relationships of Olorotitan arharensis, a hollow-crested hadrosaurid dinosaur from the latest Cretaceous of Far Eastern Russia . Acta Palaeontologica Polonica , 57 : 527 – 560 .
   Web of Science ®Google Scholar
• Godefroit , P. , Bolotsky , Y. L. and Van Itterbeeck , J. 2004 . The lambeosaurine dinosaur Amurosaurus riabinini, from the Maastrichtian of Far Eastern Russia . Acta Palaeontologica Polonica , 49 : 585 – 618 .
   Web of Science ®Google Scholar
• Godefroit , P. , Zan , S. and Jin , L. 2000 . Charonosaurus jiayinensis n. g., n. sp., a lambeosaurine dinosaur from the late Maastrichtian of northeastern China. Comptes Rendus de l’Académie des Sciences . Série IIa, Sciences de la Terre et des planètes , 330 : 875 – 882 .
   Google Scholar
• Godefroit , P. , Zan , S. and Jin , L. 2001 . The Maastrichtian (Late Cretaceous) lambeosaurine dinosaur Charonosaurus jiayinensis from north-eastern China . Bulletin de l’Institut Royal des Sciences Naturelles de Belgique, Sciences de la Terre , 71 : 119 – 168 .
   Google Scholar
• Godefroit , P. , Hai , S. , Yu , T. and Lauters , P. 2008 . New hadrosaurid dinosaurs from the uppermost Cretaceous of northeastern China . Acta Palaeontologica Polonica , 53 : 47 – 74 .
   Web of Science ®Google Scholar
• Godefroit , P. , Dong , Z.-M. , Bultynck , P. , Li , H. and Feng , L. 1998 . Sino-Belgian Cooperation Program “Cretaceous dinosaurs and mammals from Inner Mongolia”; 1. New Bactrosaurus (Dinosauria, Hadrosauroidea) material from Iren Dabasu (Inner Mongolia, P. R. China). Bulletin de l’Institut Royal des Sciences Naturelles de Belgique . Sciences de la Terre , 68 ( Supplement ) : 3 – 70 .
   Google Scholar
• Goloboff , P. , Farris , J. C. and Nixon , K. C. 2008 . TNT, a free program for phylogenetic analysis . Cladistics , 24 : 774 – 786 .
   Web of Science ®Google Scholar
• Head , J. J. 1998 . A new species of basal hadrosaurid (Dinosauria, Ornithischia) from the Cenomanian of Texas . Journal of Vertebrate Paleontology , 18 : 718 – 738 .
   Web of Science ®Google Scholar
• Head , J. J. 2001 . A reanalysis of the phylogenetic position of Eolambia caroljonesa (Dinosauria, Iguanodontia) . Journal of Vertebrate Paleontology , 21 : 392 – 396 .
   Web of Science ®Google Scholar
• Horner , J. R. , Weishampel , D. B. and Forster , C. A. 2004 . “ Hadrosauridae ” . In The Dinosauria, second edition , Edited by: Weishampel , D. B. , Dodson , P. and Osmólska , H. 438 – 463 . Berkeley , California : University of California Press .
   Google Scholar
• Lambe , L. M. 1914 . On Gryposaurus notabilis, a new genus and species of trachodont dinosaur from the Belly River Formation of Alberta, with a description of the skull of Chasmosaurus belli . The Ottawa Naturalist , 27 : 145 – 155 .
   Google Scholar
• Laurent , Y. , Cavin , L. and Bilotte , M. 1999 . Découverte d’un gisement à vertébrés dans le Maastrichtien supérieur des Petites-Pyrénées . Comptes rendus de l’Académie des Sciences. Série IIa, Sciences de la Terre et des planètes , 328 : 781 – 787 .
   Google Scholar
• Leidy , J. 1858 . Hadrosaurus foulkii, a new saurian from the Cretaceous of New Jersey . Proceedings of the Academy of Natural Sciences of Philadelphia , 10 : 215 – 218 .
   Google Scholar
• López-Martínez , N. , Canudo , J. I. , Ardévol , L. , Pereda-Suberbiola , X. , Orue-Etxebarria , X. , Cuenca-Bescós , G. , Ruiz-Omeñaca , J. I. , Murelaga , X. and Feist , M. 2001 . New dinosaur sites correlated with Upper Maastrichtian pelagic deposits in the Spanish Pyrenees: implications for the dinosaur extinction pattern in Europe . Cretaceous Research , 22 : 41 – 61 .
   Web of Science ®Google Scholar
• Lund , E. K. and Gates , T. A. 2006 . “ A historical and biogeographical examination of hadrosaurian dinosaurs ” . In Late Cretaceous Vertebrates from the Western Interior Edited by: Lucas , S. G. and Sullivan , R. M. 263 – 276 . New Mexico Museum of Natural History and Science Bulletin 35
   Google Scholar
• Maddison , W. P. and Maddison , D. R. 2010 . “ Mesquite: A Modular System for Evolutionary Analysis, version 2.73 ” . Available at mesquiteproject.org. Accessed May 30, 2012
   Google Scholar
• Marsh , O. C. 1881 . Principal characters of American Jurassic dinosaurs. Part V . American Journal of Science, Series 3 , 21 : 417 – 423 .
   Google Scholar
• Maryanska , T. and Osmólska , H. 1981 . Postcranial anatomy of Saurolophus angustirostris with comments on other hadrosaurs . Palaeontologia Polonica , 46 : 119 – 141 .
   Google Scholar
• Nagao , T. 1936 . Nipponosaurus sachalinensis, a new genus and species of trachodont dinosaur from Japanese Saghalien . Journal of the Faculty of Science, Hokkaido Imperial University. Series 4, Geology and Mineralogy , 3 : 185 – 220 .
   Google Scholar
• Nopcsa , F. 1903 . Telmatosaurus, new name for the dinosaur Limnosaurus . Geological Magazine, Series 4 , 10 : 94 – 95 .
   Google Scholar
• Ogg , J. G. , Ogg , G. and Gradstein , F. M. 2008 . “ The Concise Geologic Time Scale ” . vi + 177 New York : Cambridge University Press .
   Google Scholar
• Oms , O. and Canudo , J. I. 2004 . Datación magnetoestratigráfica de los dinosaurios del Cretácico terminal (Maastrichtiense superior) de Arén (Huesca, Unidad Surpirenaica Central) . Geo-Temas 6 , 5 : 51 – 54 .
   Google Scholar
• Ostrom , J. H. 1961 . Cranial morphology of the hadrosaurian dinosaurs of North America . Bulletin of the American Museum of Natural History , 122 : 33 – 186 .
   Google Scholar
• Owen , R. 1842 . Report on British fossil reptiles. Part II . Reports of the British Association for the Advancement of Science , 11 : 60 – 204 .
   Google Scholar
• Parks , W. A. 1922 . Parasaurophus walkeri, a new genus and species of crested trachodont dinosaur . University of Toronto Studies, Geological Series , 13 : 1 – 32 .
   Google Scholar
• Parks , W. A. 1923 . Corythosaurus intermedius, a new species of trachodont dinosaur . University of Toronto Studies, Geological Series , 15 : 1 – 57 .
   Google Scholar
• Pereda-Suberbiola , X. 2009 . Biogeographical affinities of Late Cretaceous continental tetrapods of Europe: a review, affinités paléobiogeographiques des tétrapodes continentaux du Crétacé supérieur d’Europe: une mise au point . Bulletin de la Société Géologique de France , 180 : 57 – 71 .
   Web of Science ®Google Scholar
• Pereda-Suberbiola , X. , Canudo , J. I. , Cruzado-Caballero , P. , Barco , J. L. , López-Martínez , N. , Oms , O. and Ruiz-Omeñaca , J. I. 2009 . The last hadrosaurid dinosaurs of Europe: a new lambeosaurine from the uppermost Cretaceous of Arén (Huesca, Spain) . Comptes Rendus Palevol , 8 : 559 – 572 .
   Web of Science ®Google Scholar
• Prieto-Marquez , A. 2007 . “ Postcranial osteology of the hadrosaurid dinosaur Brachylophosaurus canadensis from the Late Cretaceous of Montana ” . In Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs , Edited by: Carpenter , K. 91 – 116 . Bloomington and Indianapolis , Indiana : Indiana University Press .
   Google Scholar
• Prieto-Marquez , A. 2010 . Global phylogeny of Hadrosauridae (Dinosauria: Ornithopoda) using parsimony and Bayesian methods . Zoological Journal of the Linnean Society , 159 : 435 – 502 .
   Web of Science ®Google Scholar
• Prieto-Marquez , A. and Salinas , G. C. 2010 . A re-evaluation of Secernosaurus koerneri and Kritosaurus australis (Dinosauria, Hadrosauridae) from the Late Cretaceous of Argentina . Journal of Vertebrate Paleontology , 30 : 813 – 837 .
   Web of Science ®Google Scholar
• Prieto-Marquez , A. and Wagner , J. R. 2009 . Pararhabododon isonensis and Tsintaosaurus spinorhinus: a new clade of lambeosaurine hadrosaurids from Eurasia . Cretaceous Research , 30 : 1238 – 1246 .
   Web of Science ®Google Scholar
• Prieto-Márquez , A. , Gaete , R. , Rivas , G. , Galobart , A. and Boada , M. 2006 . Hadrosauroid dinosaurs from the Late Cretaceous of Spain: Pararhabdodon isonensis revisited and Koutalisaurus kohlerorum, gen. et sp. nov . Journal of Vertebrate Paleontology , 26 : 929 – 943 .
   Web of Science ®Google Scholar
• Puértolas , E. , Canudo , J. I. and Cruzado-Caballero , P. 2011 . A new crocodylian from the Late Maastrichtian of Spain: implications for the initial radiation of crocodyloids . PLoS ONE , 6 : e20011
   PubMed Web of Science ®Google Scholar
• Pujalte , V. and Schmitz , B. 2005 . Revisión de la estratigrafía del Grupo Tremp (“Garumniense”, Cuenca de Tremp-Graus, Pirineos meridionales) . Geogaceta , 38 : 79 – 82 .
   Google Scholar
• Riera , V. , Oms , O. , Gaete , R. and Galobart , A. 2009 . The end-Cretaceous dinosaur succession in Europe: the Tremp Basin record (Spain) . Palaeogeography, Palaeoclimatology, Palaeoecology , 283 : 160 – 171 .
   Web of Science ®Google Scholar
• Rozhdestvensky , A. K. 1966 . [New iguanodonts from Central Asia. Phylogenetic and taxonomic relationships between late Iguanodontidae and early Hadrosauridae]. Palaeontologicheskii Zhurnal 1966:103–116 . [Russian; English translation: Rozhdestvensky, A. K. 1967. New iguanodonts from Central Asia. International Geology Review , 9 : 556 – 566 .
   Google Scholar
• Seeley , H. G. 1887 . On the classification of the fossil animals commonly called Dinosauria . Proceedings of the Royal Society of London , 43 : 165 – 171 .
   Google Scholar
• Sereno , P. C. 1998 . A rationale for phylogenetic definitions, with application to the higher-level taxonomy of Dinosauria . Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen , 210 : 41 – 83 .
   Google Scholar
• Sternberg , C. M. 1953 . A new hadrosaur from the Oldman Formation of Alberta: discussion of nomenclature . National Museum of Canada Natural History Papers , 128 : 1 – 12 .
   Google Scholar
• Sues , H.-D. and Averianov , A. 2009 . A new basal hadrosauroid dinosaur from the Late Cretaceous of Uzbekistan and the early radiation of duck-billed dinosaurs . Proceedings of the Royal Society B , 267 : 2549 – 2555 .
   Web of Science ®Google Scholar
• Suzuki , D. , Weishampel , D. B. and Minoura , N. 2004 . Nipponosaurus sachalienensis (Dinosauria: Ornithopoda): anatomy and systematic position within Hadrosauridae . Journal of Vertebrate Paleontology , 24 : 145 – 164 .
   Web of Science ®Google Scholar
• Walker , J. D. and Geissman , J. W. , eds. 2009 . 2009 Geologic Time Scale , Geological Society of America . doi: 10.1130/2009.CTS004R2C
   Google Scholar
• Weishampel , D. B. , Norman , D. B. and Grigorescu , D. 1993 . Telmatosaurus transsylvanicus from the Late Cretaceous of Romania: the most basal hadrosaurid dinosaur . Palaeontology , 36 : 361 – 385 .
   Web of Science ®Google Scholar
• Williamson , T. E. 2000 . “ Review of Hadrosauridae (Dinosauria, Ornithischia) from the San Juan Basin, New Mexico ” . In Dinosaurs of New Mexico , Edited by: Lucas , S. G. and Heckert , A. B. 191 – 213 . New Mexico Museum of Natural History and Science Bulletin 17 .
   Google Scholar