A new dryosaurid ornithopod (Dinosauria, Ornithischia) from the Late Jurassic of Portugal

LITERATURE CITED

• Barrett, P. M., R. J. Butler, R. J. Twitchett, and S. Hutt. 2011. New material of Valdosaurus canaliculatus (Ornithischia: Ornithopoda) from the Lower Cretaceous of southern England. Special Papers in Palaeontology 86:131–163.
   Google Scholar
• Buffetaut, E., and E. Cacheleux. 1997. The dinosaur Dryosaurus (Ornithopoda, Dryosauridae) in the Kimmeridgian (Late Jurassic) of Normandy: paleobiogeographical implications. Comptes Rendus de l’Académie des Sciences, Series IIA 324:499–503.
   Google Scholar
• Butler, R. J. 2005. The ‘fabrosaurid’ ornithischian dinosaurs of the Upper Elliot Formation (Lower Jurassic) of South Africa and Lesotho. Zoological Journal of the Linnean Society 145:175–218.
   Web of Science ®Google Scholar
• Butler, R. J., P. Upchurch, and D. B. Norman. 2008. The phylogeny of the ornithischian dinosaurs. Journal of Systematic Palaeontology 6:1–40.
   Web of Science ®Google Scholar
• Cooper, M. R. 1985. A revision of the ornithischian dinosaur Kangnasaurus coetzeei Haughton, with a classification of Ornithischia. Annals of the South African Museum 95:281–317.
   Google Scholar
• Coria, R. A., and J. O. Calvo. 2002. A new iguanodontian ornithopod from the Neuquén Basin, Patagonia, Argentina. Journal of Vertebrate Paleontology 22:503–509.
   Web of Science ®Google Scholar
• Coria, R. A., and L. Salgado. 1996. A basal iguanodontian (Ornithischia: Ornithopoda) from the Late Cretaceous of South America. Journal of Vertebrate Paleontology 16:445–457.
   Web of Science ®Google Scholar
• Coria, R. A., J. J. Moly, M. Reguero, S. Santillana, and S. Marenssi. 2013. A new ornithopod (Dinosauria; Ornithischia) from Antarctica. Cretaceous Research 41:186–193.
   Web of Science ®Google Scholar
• Dantas, P. P., C. J. Yagüe, F. Hazevoet, V. Ortega, J. L. dos Santos, M. Sanz, A. M. Cachão, de Carvalho Galopim, and J. J. Santos. 2000. Estudo preliminar de um novo Iguanodontia basal do Jurássico Superior Português; pp. 12–13 in J. B. Diez and A. C. Balbino (eds.), Livro de Resumos do I Congresso Ibérico de Paleontologia/XVI Jornadas de la Sociedad Española de Paleontología, Évora (Portugal), 12–14 Outubro 2000. Universidade de Évora, Évora, Portugal.
   Google Scholar
• DiCroce, T., and K. Carpenter. 2001. New ornithopod from the Cedar Mountain Formation (Lower Cretaceous) of eastern Utah; pp. 183–196 in D. H. Tanke and K. Carpenter (eds.), Mesozoic Vertebrate Life. Indiana University Press, Bloomington, Indiana.
   Google Scholar
• Galton, P. M. 1974. The ornithischian dinosaur Hypsilophodon from the Wealden of the Isle of Wight. Bulletin of the British Museum (Natural History) Geology 25:1–152.
   Google Scholar
• Galton, P. M. 1981. Dryosaurus, a hypsilophodontid dinosaur from the Upper Jurassic of North America and Africa. Postcranial skeleton. Paläontologische Zeitschrift 55:271–312.
   Google Scholar
• Galton, P. M., and P. Taquet. 1982. Valdosaurus, a hypsilophodontid dinosaur from the Lower Cretaceous of Europe and Africa. Geobios 15:147–159.
   Google Scholar
• Gilmore, C. W. 1909. Osteology of the Jurassic reptile Camptosaurus, with a revision of the species of the genus, and descriptions of two new species. Proceedings of the United States National Museum 36:197–332.
   Google Scholar
• Goloboff, P., J. C. Farris, and K. C. Nixon. 2008. TNT, a free program for phylogenetic analysis. Cladistics 24:774–786.
   Web of Science ®Google Scholar
• Han, F.-L., P. M. Barrett, R. J. Butler, and X. Xu. 2012. Postcranial anatomy of Jeholosaurus shangyuanensis (Dinosauria, Ornithischia) from the Lower Cretaceous Yixian Formation of China. Journal of Vertebrate Paleontology 32:1370–1395.
   Web of Science ®Google Scholar
• Horner J. R., D. B. Weishampel, and C. A. Foster. 2004. Hadrosauridae; pp. 438–463 in D. B. Weishampel, P. Dodson, and H. Osmólska (eds.), The Dinosauria, second edition. University of California Press, Berkeley, California.
   Google Scholar
• Horner J. R., A. de Ricqlès, K. Padian, and R. D. Scheetz. 2009. Comparative long bone histology and growth of the ‘hypsilophodontid’ dinosaurs Orodromeus makelai, Dryosaurus altus, and Tenontosaurus tillettii (Ornithischia: Euornithopoda). Journal of Vertebrate Paleontology 29:734–747.
   Web of Science ®Google Scholar
• Hübner, T. R. 2012. Bone histology in Dysalotosaurus lettowvorbecki (Ornithischia: Iguanodontia)—variation, growth, and implications. PLoS ONE 7:e29958. doi: 10.1371/journal.pone.0029958.
   PubMed Web of Science ®Google Scholar
• Janensch, W. 1955. Der Ornithopode Dysalotosaurus der Tendaguruschichten. Palaeontographica, Supplement 7(3):105–176.
   Google Scholar
• Malafaia, E., F. Ortega, F. Escaso, P. Dantas, N. Pimentel, J. M. Gasulla, F. Barriga, and J. L. Sanz. 2010. Vertebrate fauna at the Allosaurus fossil-site of Andrés (Upper Jurassic), Pombal, Portugal. Journal of Iberian Geology 36:193–204.
   Web of Science ®Google Scholar
• Manupella, G., M. T. Antunes, J. Pais, M. M. Ramalho, and J. Rey. 1999. Notícia Explicativa da Carta Geológica de Portugal, folha 30-A (Lourinhã). Escala 1: 50,000 (2 ed.). Publicações do Instituto Geológico e Mineiro, Lisbon, 83 pp.
   Google Scholar
• Marsh, O. C. 1881. Principal characters of the American Jurassic dinosaurs. Part V. American Journal of Science, Series 3 21:417–423.
   Google Scholar
• Mateus, O. 2007. Notes and review of the ornithischian dinosaurs of Portugal. Journal of Vertebrate Paleontology 27(3, Supplement):114A.
   Web of Science ®Google Scholar
• McDonald, A. T. 2012. Phylogeny of basal iguanodonts (Dinosauria: Ornithischia): an update. PLoS ONE 7:e36745. doi: 10.1371/journal.pone.0036745.
   PubMed Web of Science ®Google Scholar
• Milner, A. R., and D. B. Norman. 1984. The biogeography of advanced ornithopod dinosaurs (Archosauria: Ornithischia)—a cladistic-vicariance model; pp. 145–150 in W.-E. Reif and F. Westphal (eds.), Third Symposium on Mesozoic Terrestrial Ecosystems, Short Papers. Attempto Verlag, Tübingen.
   Google Scholar
• Norman, D. B. 2004. Basal Iguanodontia; pp. 413–437 in D. B. Weishampel, P. Dodson, and H. Osmólska (eds.), The Dinosauria, second edition. University of California Press, Berkeley, California.
   Google Scholar
• Norman, D. B., H.-D. Sues, L. M. Witmer, and R. A. Coria. 2004. Basal Ornithopoda; pp. 393–412 in D. B. Weishampel, P. Dodson, and H. Osmólska (eds.), The Dinosauria, second edition. University of California Press, Berkeley, California.
   Google Scholar
• Ostrom, J. H. 1970. Stratigraphy and pa1eontology of the Cloverly Formation (Lower Cretaceous) of the Bighorn Basin area, Wyoming and Montana. Bulletin of the Peabody Museum of Natural History 35:1–234.
   Google Scholar
• Owen, R. 1842. Report on British fossil reptiles. Part II. Reports of the British Association for the Advancement of Science 11:60–204.
   Google Scholar
• Pincemaille-Quilléyéré, M. 2002. Description d’un squelette partiel de Rhabdodon priscus (Euornithopoda) du Crétacé Superieur de Vitrolles (Bouches du Rhône, France). Oryctos 4:39–70.
   Google Scholar
• Ruiz-Omeñaca, J. L., X. Pereda Suberbiola, and P. M. Galton. 2007. Callovosaurus leedsi, the earliest dryosaurid dinosaur (Ornithischia: Euornithopoda) from the Middle Jurassic of England; pp. 3–16 in K. Carpenter (ed.), Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs. Indiana University Press, Bloomington and Indianapolis, Indiana.
   Google Scholar
• Salgado, L., R. A. Coria, and S. E. Heredia. 1997. New materials of Gasparinisaura cincosaltensis (Ornithischia, Ornithopoda) from the Upper Cretaceous of Argentina. Journal of Vertebrate Paleontology 71:933–940.
   Web of Science ®Google Scholar
• Seeley, H. G. 1887. On the classification of the fossil animals commonly called Dinosauria. Proceedings of the Royal Society of London 43:165–171.
   Google Scholar
• Sereno, P. C. 1986. Phylogeny of the bird-hipped dinosaurs (Order Ornithischia). National Geographic Research 2:234–256.
   Google Scholar
• Shepherd, J. D., P. M. Galton, and J. A. Jensen. 1977. Additional specimens of the hypsilophodontid dinosaur Dryosaurus altus from the Upper Jurassic of western North America. Brigham Young University Geology Studies 24:11–15.
   Google Scholar
• Yagüe, P., P. Dantas, F. Ortega, M. Cachão, F. A. M. Santos, R. Gonçalves, and S. Lopes. 2006. New sauropod material from the Upper Jurassic of Areia Branca (Lourinhã, Portugal). Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen 240:313–342.
   Web of Science ®Google Scholar
• Submitted June 14, 2013; revisions received September 16, 2013; accepted September 21, 2013. Handling editor: Paul Barrett.
   Google Scholar