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Articles

A novel archosauromorph from Antarctica and an updated review of a high-latitude vertebrate assemblage in the wake of the end-Permian mass extinction

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Article: e1536664 | Received 19 Sep 2017, Accepted 15 Aug 2018, Published online: 31 Jan 2019

LITERATURE CITED

  • Abdala, F., J. Neveling, and J. Welman. 2006. A new trirachodontid cynodont from the lower levels of the Burgersdorp Formation (Lower Triassic) of the Beaufort Group, South Africa and the cladistic relationships of Gondwanan gomphodonts. Zoological Journal of the Linnean Society 147:383–413.
  • Angielczyk, K. D. 2007. New specimens of the Tanzanian dicynodont “Cryptocynodon” parringtoni von Huene, 1942 (Therapsida, Anomodontia), with an expanded analysis of Permian dicynodont phylogeny. Journal of Vertebrate Paleontology 27:116–131.
  • Barrett, P. J. 1969. Stratigraphy and Petrology of the Mainly Fluviatile Permian and Triassic Beacon Rocks, Beardmore Glacier Area, Antarctica. Institute of Polar Studies Report 34. Ohio State University, Columbus, Ohio, 132 pp.
  • Barrett, P. J., R. J. Baillie, and E. H. Colbert. 1968. Triassic amphibian from Antarctica. Science 161:460–462.
  • Barrett, P. J., D. H. Elliot, and J. F. Lindsay. 1986. The Beacon Supergroup (Devonian-Triassic) and Ferrar Group (Jurassic) in the Beardmore Glacier area, Antarctica; pp. 339–428 in M. D. Turner and J. E. Splettstoesser (eds.), Geology of the Central Transantarctic Mountains. Antarctic Research Series 161. American Geophysical Union, Washington, D.C.
  • Bartholomai, A. 1979. New lizard-like reptiles from the Early Triassic of Queensland. Alcheringa 3:225–234.
  • Beightol, C. V., A. K. Huttenlocker, B. R. Peecook, C. A. Sidor, and R. M. H. Smith. 2013. A new basal stereospondyl (Temnospondyli) from the Lower Triassic Fremouw Formation of Antarctica. Journal of Vertebrate Paleontology 33(Programs and Abstracts):84A.
  • Benton, M. J. 2016. The Triassic. Current Biology 26:R1205–R1225.
  • Botha, J., and R. M. H. Smith. 2006. Rapid vertebrate recuperation in the Karoo Basin of South Africa following the end-Permian extinction. Journal of African Earth Sciences 45:502–514.
  • Botha, J., and R. M. H. Smith. 2007. Lystrosaurus species composition across the Permo-Triassic boundary in the Karoo Basin of South Africa. Lethaia 40:125–137.
  • Broom, R. 1906. On the Permian and Triassic faunas of South Africa. Geological Magazine 5:29–30.
  • Butler, R. J., S. L. Brusatte, M. Reich, S. J. Nesbitt, R. R. Schoch, and J. J. Hornung. 2011. The sail-backed reptile Ctenosauriscus from the latest Early Triassic of Germany and the timing and biogeography of the early archosaur radiation. PLoS ONE 6:e25693.
  • Camp, C. L., and M. R. Banks. 1978. A proterosuchian reptile from the Early Triassic of Tasmania. Alcheringa 2:143–158.
  • Colbert, E. H. 1970. The fossil tetrapods of Coalsack Bluff. Antarctic Journal of the United States 5:57–61.
  • Colbert, E. H. 1974. Lystrosaurus from Antarctica. American Museum Novitates 2535:1–44.
  • Colbert, E. H. 1975. Further determinations of Antarctic Triassic tetrapods. Antarctic Journal of the United States 10:250–252.
  • Colbert, E. H. 1978. Proto-lizards from the Triassic of Antarctica. Antarctic Journal 13:20–21.
  • Colbert, E. H. 1987. The Triassic reptile Prolacerta in Antarctica. American Museum Novitates 2882:1–19.
  • Colbert, E. H., and J. W. Kitching. 1975. The Triassic reptile Procolophon in Antarctica. American Museum Novitates 2566:1–23.
  • Colbert, E. H., and J. W. Kitching. 1977. Triassic cynodont reptiles from Antarctica. American Museum Novitates 2611:1–30.
  • Colbert, E. H., and J. W. Kitching. 1981. Scaloposaurian reptiles from the Triassic of Antarctica. American Museum Novitates 2709:1–22.
  • Collinson, J. W. 1991. The palaeo-Pacific margin as seen from East Antarctica; pp. 199–204 in M. R. A. Thomson, J. A. Crame, and J. W. Thomson (eds.), Geological Evolution of Antarctica. Cambridge University Press, Cambridge, U.K.
  • Collinson, J. W., W. R. Hammer, R. A. Askin, and D. H. Elliot. 2006. Permian-Triassic boundary in the central Transantarctic Mountains, Antarctica. Geological Society of America Bulletin 118:747–763.
  • Collinson, J. W., J. L. Isbell, D. H. Elliot, M. F. Miller, and J. M. G. Miller. 1994. Permian-Triassic Transantarctic Basin; pp. 173–222 in J. J. Veevers and C. M. Powell (eds.), Permian-Triassic Pangean Basins and Foldbelts along the Panthalassan Margin of Gondwanaland. Geological Society of America Memoir 184. Boulder, Colorado.
  • Cosgriff, J. W. 1983. Large thecodont reptiles from the Fremouw Formation. Antarctic Journal of the United States 18:52–55.
  • Cosgriff, J. W., and W. R. Hammer. 1984. New material of labyrinthodont amphibians from the Lower Triassic Fremouw Formation of Antarctica. Journal of Vertebrate Paleontology 4:47–56.
  • Cosgriff, J. W., W. R. Hammer, and W. J. Ryan. 1982. The Pangaean reptile Lystrosaurus maccaigi, in the Lower Triassic of Antarctica. Journal of Paleontology 56:371–385.
  • Cosgriff, J. W., W. R. Hammer, J. M. Zawiskie, and N. R. Kemp. 1978. New Triassic vertebrates from the Fremouw Formation of the Queen Maud Mountains. Antarctic Journal of the United States 13:23–24.
  • Dilkes, D. W. 1998. The Early Triassic rhynchosaur Mesosuchus browni and the interrelationships of basal archosauromorph reptiles. Philosophical Transactions of the Royal Society of London B: Biological Sciences 353:501–541.
  • Elliot, D. H., E. H. Colbert, J. Jensen, W. Breed, and J. S. Powell. 1970. Triassic tetrapods from Antarctica: evidence for continental drift. Science 169:1197–1201.
  • Evans, S. E., and M. S. King. 1993. A new specimen of Protorosaurus (Reptilia: Diapsida) from Marl Slate (Late Permian) of Britain. Proceedings of the Yorkshire Geological Society 49:229–234.
  • Ezcurra, M. D. 2014. The osteology of the basal archosauromorph Tasmaniosaurus triassicus from the Lower Triassic of Tasmania, Australia. PLoS ONE 9:e86864.
  • Ezcurra, M. D. 2016. The phylogenetic relationships of basal archosauromorphs, with an emphasis on the systematics of proterosuchian archosauriforms. PeerJ 4:e1778.
  • Ezcurra, M. D., and R. J. Butler. 2015. Taxonomy of the proterosuchid archosauriforms (Diapsida: Archosauromorpha) from the earliest Triassic of South Africa, and implications for the early archosauriform radiation. Palaeontology 58:141–170.
  • Ezcurra, M. D., R. J. Butler, and D. J. Gower. 2013. ‘Proterosuchia’: the origin and early history of Archosauriformes; pp. 9–33 in S. J. Nesbitt, J. B. Desojo, and R. B. Irmis (eds.), Anatomy, Phylogeny, and Palaeobiology of Early Archosaurs and Their Kin. Geographical Society, London, Special Publications 379.
  • Ezcurra, M. D., T. M. Scheyer, and R. J. Butler. 2014. The origin and early evolution of Sauria: reassessing the Permian saurian fossil record and the timing of the crocodile-lizard divergence. PLoS ONE 9:e89165.
  • Foth, C., M. D. Ezcurra, R. B. Sookias, S. L. Brusatte, and R. J. Butler. 2016. Unappreciated diversification of stem archosaurs during the Middle Triassic predated the dominance of dinosaurs. BMC Evolutionary Biology 16:188. doi: 10.1186/s12862-016-0761-6.
  • Fröbisch, J. 2007. The cranial anatomy of Kombuisia frerensis Hotton (Synapsida, Dicynodontia) and a new phylogeny of anomodont therapsids. Zoological Journal of the Linnean Society 150:117–144.
  • Fröbisch, J., K. D. Angielczyk, and C. A. Sidor. 2010. The Triassic dicynodont Kombuisia (Synapsida, Anomodontia) from Antarctica, a refuge from the terrestrial Permian-Triassic mass extinction. Naturwissenschaften 97:187–196.
  • Gao, K.-Q., R. C. Fox, C.-F. Zhou, and D.-Q. Li. 2010. A new nonmammalian eucynodont (Synapsida: Therapsida) from the Triassic of northern Gansu Province, China, and its biostratigraphic and biogeographic implications. American Museum Novitates 3685:1–5.
  • Gastaldo, R. A., S. L. Kamo, J. Neveling, J. W. Geissman, M. Bamford, and C. V. Looy. 2015. Is the vertebrate-defined Permian-Triassic boundary in the Karoo Basin, South Africa, the terrestrial expression of the end-Permian marine event? Geology 43:939–942.
  • Gauthier, J. A. 1984. A cladistic analysis of the higher categories of the Diapsida. Ph.D. dissertation, University of California, Berkeley, California, 564 pp.
  • Gauthier, J. A., A. G. Kluge, and T. Rowe. 1988. Amniote phylogeny and the importance of fossils. Cladistics 4:105–209.
  • Goloboff, P. A., J. S. Farris, and K. C. Nixon. 2008. TNT, a free program for phylogenetic analysis. Cladistics 24:774–786.
  • Golubev, V. K. 2000. The faunal assemblages of Permian terrestrial vertebrates from Eastern Europe. Paleontological Journal 34(2, Supplement):S211–S224.
  • Gow, C. 1992. An enigmatic new reptile from the Lower Triassic Fremouw Formation of Antarctica. Palaeontologica africana 29:21–23.
  • Gow, C. 1999. The Triassic reptile Palacrodon browni Broom, synonymy and a new specimen. Palaeontologica africana 35:21–23.
  • Gower, D. J. 2003. Osteology of the early archosaurian reptile Erythrosuchus africanus, Broom. Annals of the South African Museum 110:1–84.
  • Gower, D. J., and A. G. Sennikov. 1996. Morphology and phylogenetic informativeness of early archosaur braincases. Palaeontology 39:883–906.
  • Gower, D. J., and A. G. Sennikov. 1997. Sarmatosuchus and the early history of the Archosauria. Journal of Vertebrate Paleontology 17:60–73.
  • Gower, D. J., P. J. Hancox, J. Botha-Brink, A. G. Sennikov, and R. J. Butler. 2014. A new species of Garjainia Ochev, 1958 (Diapsida: Archosauriformes: Erythrosuchidae) from the Early Triassic of South Africa. PLoS ONE 9:e111154.
  • Hammer, W. R. 1995. New therapsids from the upper Fremouw Formation (Triassic) of Antarctica. Journal of Vertebrate Paleontology 15:105–112.
  • Hammer, W. R., and J. W. Cosgriff. 1981. Myosaurus gracilis, an anomodont reptile from the Lower Triassic of Antarctica and South Africa. Journal of Paleontology 55:410–424.
  • Hammer, W. R., and W. J. Hickerson. 1994. A crested theropod dinosaur from Antarctica. Science 264:828–830.
  • Hammer, W. R., and N. D. Smith. 2008. A tritylodont postcanine from the Hanson Formation of Antarctica. Journal of Vertebrate Paleontology 28:269–273.
  • Hammer, W. R., J. W. Collinson, and W. J. Ryan Jr. 1990. A new Triassic vertebrate fauna from Antarctica and its depositional setting. Antarctic Science 2:163–167.
  • Hammer, W. R., W. J. Ryan, and S. L. DeFauw. 1987. Comments of the vertebrate fauna from the Fremouw Formation (Triassic), Beardmore Glacier region Antarctica. Antarctic Journal of the United States 22:32–33.
  • Hammer, W. R., J. W. Collinson, R. A. Askin, and W. J. Hickerson. 2004. The first Upper Triassic vertebrate locality in Antarctica. Gondwana Research 7:199–204.
  • Hotton, N. 1974. A new dicynodont (Reptilia, Therapsida) from Cynognathus Zone deposits of South Africa. Annals of the South African Museum 64:157–165.
  • Huene, F. von. 1940. Eine reptilfauna aus der ältesten Trias Nordrusslands. Neues Jahrbuch für Mineralogie, Geologie, und Paläontologie, Beilage-Band 84:1–23.
  • Huene, F. von. 1946. Die grossen Stämme der Tetrapoden in den geologischen Zeiten. Biologisches Zentralblatt 65:268–275.
  • Huttenlocker, A. K., and C. A. Sidor. 2012. Taxonomic revision of therocephalians (Therapsida: Theriodontia) from the Lower Triassic of Antarctica. American Museum Novitates 3738:1–19.
  • Isbell, J. L., 1991. Evidence for a low-gradient alluvial fan from the palaeo-Pacific margin in the Upper Permian Buckley Formation, Beardmore Glacier area, Antarctica; pp. 215–217 in M. R. A. Thomson, J. A. Crame, and J. W. Thomson (eds.), Geological Evolution of Antarctica. Cambridge University Press, Cambridge, U.K.
  • Jiang, D.-Y., R. Motani, A. Tintori, O. Rieppel, G.-B. Chen, J.-D. Huang, R. Zhang, Z.-Y. Sun, and C. Ji. 2014. The early Triassic eosauropterygian Majiashanosaurus discocoracoidis gen. et sp. nov. (Reptilia, Sauropterygia), from Chaohu, Anhui Province, People’s Republic of China. Journal of Vertebrate Paleontology 34:1044–1052.
  • Kitching, J. W., J. W. Collinson, D. H. Elliot, and E. H. Colbert. 1972. Lystrosaurus Zone (Triassic) fauna from Antarctica. Science 175:524–527.
  • Krull, E. S., and G. J. Retallack. 2000. ∂13C depth profiles from paleosols across the Permian-Triassic boundary: evidence for methane release. Geological Society of America Bulletin 112:1459–1472.
  • Laurenti, J. N. 1768. Classis Reptilium. Specimen medicum, exhibens synopsis Reptilium emendatum, cum experimentis circa venena et antidote Reptilium Austriacorum. Joan. Thom., Nob. de Trattnern, Vienna.
  • Lucas, S. G. 2001. Chinese Fossil Vertebrates. Columbia University Press, New York, 375 pp.
  • Maddison, W. P., and D. R. Maddison. 2008. Mesquite: a modular system for evolutionary analysis, version 3.02. Available at mesquiteproject.org. Accessed July 1, 2017.
  • Meyer, H. von. 1832. Palaeologica zur Geschichte der Erde und ihrer Geschöpfe. Verlag von Siegmund Schmerber, Frankfurt am Main, 560 pp.
  • Modesto, S. P., and J. S. Anderson. 2004. The phylogenetic definition of Reptilia. Systematic Biology 53:815–821.
  • Modesto, S. P., and J. Botha-Brink. 2008. Evidence of a second, large archosauriform reptile in the Lower Triassic Katberg Formation of South Africa. Journal of Vertebrate Paleontology 28:914–917.
  • Modesto, S. P., and H.-D. Sues. 2004. The skull of the Early Triassic archosauromorph reptile Prolacerta broomi and its phylogenetic significance. Zoological Journal of the Linnean Society 140:335–351.
  • Motani, R., D.-Y. Jiang, G.-B. Chen, A. Tintori, O. Rieppel, C. Ji, and J.-D. Huang. 2015a. A basal ichthyosauriform with a short snout from the Lower Triassic of China. Nature 517:485–488.
  • Motani, R., D.-Y. Jiang, A. Tintori, O. Rieppel, G.-B. Chen, and H. You. 2015b. Status of Chaohusaurus chaoxianensis (Chen, 1985). Journal of Vertebrate Paleontology. doi: 10.1080/02724634.2014.892011.
  • Nesbitt, S. J. 2011. The early evolution of archosaurs: relationships and the origin of major clades. Bulletin of the American Museum of Natural History 352:1–292.
  • Nesbitt, S. J., J. Liu, and C. Li. 2011. A sail-backed suchian from the Heshanggou Formation (Early Triassic: Olenekian) of China. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 101:271–284.
  • Nesbitt, S. J., J. J. Flynn, A. C. Pritchard, J. M. Parrish, L. Ranivoharimanana, and A. R. Wyss. 2015. Postcranial anatomy and relationships of Azendhosaurus madagaskarensis. Bulletin of the American Museum of Natural History 398:1–126.
  • Neveling, J. 2004. Stratigraphic and Sedimentological Investigation of the Contact between the Lystrosaurus and Cynognathus Assemblage Zones (Beaufort Group: Karoo Supergroup). Bulletin 137. Council for Geoscience, Pretoria, South Africa, 164 pp.
  • Nicolas, M., and B. S. Rubidge. 2010. Changes in Permo-Triassic terrestrial tetrapod ecological representation in the Beaufort Group (Karoo Supergroup) of South Africa. Lethaia 43:45–59.
  • Ochev, V. G. 1979. New Early Triassic archosaurs from eastern European Russia. Paleontological Journal 1979:104–109.
  • Pritchard, A. C., and S. J. Nesbitt. 2017. A bird-like skull in a Triassic diapsid reptile increases heterogeneity of the morphological and phylogenetic radiation of Diapsida. Royal Society Open Science 4:170499. doi: 10.1098/rsos.170499.
  • Retallack, G. J., and E. S. Krull. 1999. Landscape ecologic shift at the Permian-Triassic boundary in Antarctica. Australian Journal of Earth Sciences 46:785–812.
  • Retallack, G. J. and E. S. Krull. 2006. Carbon isotopic evidence for terminal-Permian methane outbursts and their role in extinctions of animals, plants, coral reefs, and peat swamps; in S. F. Grebs and W. A. DiMichele (eds.) Wetlands Through Time. Geological Society of America Special Paper 399:249–268.
  • Retallack, G. J., E. S. Krull and S. E. Robinson. 1996. Permian and Triassic paleosols and paleoenvironments of the central Transantarctic Mountains, Antarctica. Antarctic Journal of the United States 31:29–32.
  • Retallack, G. J., A. H. Jahren, N. D. Sheldon, R. Chakrabarti, C. A. Metzger, and R. M. H. Smith. 2005. The Permian-Triassic boundary in Antarctica. Antarctic Science 17:241–258.
  • Rubidge, B. S. 2005. Re-uniting lost continents—fossil reptiles from the ancient Karoo and their wanderlust. South African Journal of Geology 108:135–172.
  • Ruta, M., J. Botha-Brink, S. A. Mitchell, and M. J. Benton. 2013. The radiation of cynodonts and the ground plan of mammalian morphological diversity. Proceedings of the Royal Society B: Biological Sciences 280:20131865. doi: 10.1098/rspb.2013.1865.
  • Scheyer, T. M., C. Romano, J. Jenks, and H. Bucher. 2014. Early Triassic marine biotic recovery: the predators’ perspective. PLoS ONE 9:e88987.
  • Schoch, R. R., and A. R. Milner. 2000. Stereospondyli. Handbuch der Paläoherpetologie, Teil 3B. Verlag Dr. Friedrich Pfeil, Munich, Germany, 203 pp.
  • Sennikov, A. G. 1992. Oldest proterosuchids from the Triassic of eastern Europe. Doklady Akademii Nauk 326:896–899.
  • Sennikov, A. G. 1995. Early thecodonts of Eastern Europe. Trudy Paleontologischeskogo Instituta RAN 263:1–141.
  • Sennikov, A. G. 1997. An enigmatic reptile from the upper Permian of the Volga River Basin. Paleontological Journal 31:94–101.
  • Sereno, P. C. 1991. Basal archosaurs: phylogenetic relationships and functional implications. Society of Vertebrate Paleontology Memoir 2. Journal of Vertebrate Paleontology 11(4, Supplement):1–53.
  • Sidor, C. A., R. Damiani, and W. R. Hammer. 2008. A new temnospondyl from Antarctica and a review of Fremouw Formation biostratigraphy. Journal of Vertebrate Paleontology 28:656–663.
  • Sidor, C. A., J. S. Steyer, and W. R. Hammer. 2014a. A new capitosauroid temnospondyl from the Middle Triassic upper Fremouw Formation of Antarctica. Journal of Vertebrate Paleontology 34:539–548.
  • Sidor, C. A., R. M. H. Smith, A. K. Huttenlocker, and B. R. Peecook. 2014b. New Middle Triassic tetrapods from the upper Fremouw Formation of Antarctica and their depositional setting. Journal of Vertebrate Paleontology 34:793–801.
  • Sidor, C. A., D. A. Vilhena, K. D. Angielczyk, A. K. Huttenlocker, S. J. Nesbitt, B. R. Peecook, J. S. Steyer, R. M. H. Smith, and L. A. Tsuji. 2013. Provincialization of terrestrial faunas following the end-Permian mass extinction. Proceedings of the National Academy of Sciences of the United States of America 110:8129–8133.
  • Smith, N. D., and D. Pol. 2007. Anatomy of a basal sauropodomorph dinosaur from the Early Jurassic Hanson Formation of Antarctica. Acta Palaeontologica Polonica 52:657–674.
  • Smith, N. D., J. R. Crandall, S. M. Hellert, W. R. Hammer, and P. J. Makovicky. 2011. Anatomy and affinities of large archosauromorphs from the lower Fremouw Formation (Early Triassic) of Antarctica. Journal of Vertebrate Paleontology 31:784–797.
  • Smith, R. M. H. 1995. Changing fluvial environments across the Permo-Triassic boundary in the Karoo Basin, South Africa and possible causes of tetrapod extinctions. Palaeogeography, Palaeoclimatology, Palaeoecology 117:81–104.
  • Smith, R. M. H., and J. Botha-Brink. 2014. Anatomy of a mass extinction: sedimentological and taphonomic evidence for drought-induced die-offs at the Permo-Triassic boundary in the main Karoo Basin, South Africa. Palaeogeography, Palaeoclimatology, Palaeoecology 396:99–118.
  • Smith, R. M. H., and P. D. Ward. 2001. Pattern of vertebrate extinctions across an event bed at the Permian-Triassic boundary in the Karoo Basin of South Africa. Geology 29:1147–1150.
  • Smith, R. M. H., B. Rubidge, and M. van der Walt. 2012. Therapsid biodiversity patterns and paleoenvironments of the Karoo Basin, South Africa; pp. 223–246 in A. Chinsamy-Turan (ed.), Forerunners of Mammals. Indiana University Press, Indianapolis, Indiana.
  • Sookias, R. B. 2016. The relationships of the Euparkeriidae and the rise of Archosauria. Royal Society Open Science 3:150674. doi: 10.1098/rsos.150674.
  • Sookias, R. B., and R. J. Butler. 2013. Euparkeriidae; pp. 35–48 in S. J. Nesbitt, J. B. Desojo, and R. B. Irmis (eds.), Anatomy, Phylogeny, and Palaeobiology of Early Archosaurs and Their Kin. Geographical Society, London, Special Publications 379.
  • Spiekman, S. N. F. In press. A new specimen of Prolacerta broomi from the lower Fremouw Formation (Early Triassic) of Antarctica, its biogeographical implications and a taxonomic revision. Scientific Reports. doi: 10.1038/s41598-D18-36499-6.
  • Tatarinov, L. P. 1960. Discovery of pseudosuchians in the upper Permian of SSSR. Paleontological Journal 1960:74–80.
  • Thulborn, R. A. 1979. A proterosuchian thecodont from the Rewan Formation of Queensland. Memoir of the Queensland Museum 19:331–355.
  • Viglietti, P. A., R. M. H. Smith, K. D. Angielczyk, C. F. Kammerer, J. Fröbisch, and B. S. Rubidge. 2015. The Daptocephalus Assemblage Zone (Lopingian), South Africa: a proposed biostratigraphy based on a new compliation of stratigraphic ranges. Journal of African Earth Sciences. doi: 10.1016/j.jafrearsci.2015.10.011.
  • Ward, P. D., J. Botha, R. Buick, M. O. De Kock, D. H. Erwin, G. H. Garrison, J. L. Kirschvink, R. M. H. Smith. 2005. Abrupt and gradual extinction among Late Permian land vertebrates in the Karoo Basin, South Africa. Science 307:709–714.
  • Warren, A. A., and M. N. Hutchinson. 1990. Lapillopsis, a new genus of temnospondyl amphibians from the Early Triassic of Queensland. Alcheringa 14:149–158.
  • Warren, A. A., and C. Marsicano. 2000. A phylogeny of the Brachypoidea (Temnospondyli, Stereospondyli). Journal of Vertebrate Paleontology 20:462–483.
  • Watson, D. M. S. 1914. The zones of the Beaufort Beds of the Karroo System in South Africa. Geological Magazine, New Series, Decade 6:203–208.
  • Welman, J. 1998. The taxonomy of the South African proterosuchids (Reptilia, Archosauromorpha). Journal of Vertebrate Paleontology 18:340–347.
  • Yates, A. M. 1999. The Lapillopsidae: a new family of small temnospondyls from the Early Triassic of Australia. Journal of Vertebrate Paleontology 19:302–320.
  • Yates, A. M., and D. P. Sengupta. 2002. A lapillopsid temnospondyl from the Early Triassic of India. Alcheringa 26:201–208.

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