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Articles

New megaraptorid (Dinosauria: Theropoda) remains from the Lower Cretaceous Eumeralla Formation of Cape Otway, Victoria, Australia

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Article: e1666273 | Received 08 Oct 2018, Accepted 10 Jul 2019, Published online: 10 Oct 2019

LITERATURE CITED

  • Agnolin, F. L., M. D. Ezcurra, D. F. Pais, and S. W. Salisbury. 2010. A reappraisal of the Cretaceous non-avian dinosaur faunas from Australia and New Zealand: evidence for their Gondwanan affinities. Journal of Systematic Palaeontology 8:257–300. doi: 10.1080/14772011003594870
  • Allain, R. 2005. The postcranial anatomy of the megalosaur Dubreuillosaurus valesdunensis (Dinosauria Theropoda) from the Middle Jurassic of Normandy, France. Journal of Vertebrate Paleontology 25:850–858. doi: 10.1671/0272-4634(2005)025[0850:TPAOTM]2.0.CO;2
  • Apesteguía, S., N. D. Smith, R. Juárez Valieri, and P. J. Makovicky. 2016. An unusual new theropod with a didactyl manus from the Upper Cretaceous of Patagonia, Argentina. PLoS One 11:e0157793. doi: 10.1371/journal.pone.0157793
  • Aranciaga Rolando, A. M., F. E. Novas, and F. L. Agnolín. 2019. A reanalysis of Murusraptor barrosaensis Coria & Currie (2016) affords new evidence about the phylogenetical relationships of Megaraptora. Cretaceous Research 99:104–127. doi: 10.1016/j.cretres.2019.02.021
  • Azuma, Y., and P. J. Currie. 2000. A new carnosaur (Dinosauria: Theropoda) from the Lower Cretaceous of Japan. Canadian Journal of Earth Sciences 37:1735–1753. doi: 10.1139/e00-064
  • Bakker, R. T. 1986. The Dinosaur Heresies. William Morrow and Company, Inc., New York, 481 pp.
  • Barrett, P. M., B. P. Kear, and R. B. J. Benson. 2010. Opalized archosaur remains from the Bulldog Shale (Aptian: Lower Cretaceous) of South Australia. Alcheringa 34:293–301. doi: 10.1080/03115511003664440
  • Barrett, P. M., R. B. J. Benson, T. H. Rich, and P. Vickers-Rich. 2011. First spinosaurid dinosaur from Australia and the cosmopolitanism of Cretaceous dinosaur faunas. Biology Letters 7:933–936. doi: 10.1098/rsbl.2011.0466
  • Bell, P. R., A. Cau, F. Fanti, and E. Smith. 2016. A large-clawed theropod (Dinosauria: Tetanurae) from the Lower Cretaceous of Australia and the Gondwanan origin of megaraptorid theropods. Gondwana Research 36:473–487. doi: 10.1016/j.gr.2015.08.004
  • Bell, P. R., F. Fanti, L. J. Hart, L. A. Milan, S. J. Craven, T. Brougham, and E. Smith. 2019. Revised geology, age, and vertebrate diversity of the dinosaur-bearing Griman Creek Formation (Cenomanian), Lightning Ridge, New South Wales, Australia. Palaeogeography, Palaeoclimatology, Palaeoecology 514:655–671. doi: 10.1016/j.palaeo.2018.11.020
  • Benson, R. B. J., and X. Xu. 2008. The anatomy and systematic position of the theropod dinosaur Chilantaisaurus tashuikouensis Hu, 1964 from the Early Cretaceous of Alanshan, People’s Republic of China. Geological Magazine 145:778–789. doi: 10.1017/S0016756808005475
  • Benson, R. B. J., M. T. Carrano, and S. L. Brusatte. 2010. A new clade of archaic large-bodied predatory dinosaurs (Theropoda: Allosauroidea) that survived to the latest Mesozoic. Naturwissenschaften 97:71–78. doi: 10.1007/s00114-009-0614-x
  • Benson, R. B. J., T. H. Rich, P. Vickers-Rich, and M. Hall. 2012. Theropod fauna from southern Australia indicates high polar diversity and climate-driven dinosaur provinciality. PLoS ONE 7:e37122. doi: 10.1371/journal.pone.0037122
  • Brochu, C. A. 2003. Osteology of Tyrannosaurus rex: Insights from a Nearly Complete Skeleton and High-Resolution Computed Tomographic Analysis. Society of Vertebrate Paleontology Memoir 7. Journal of Vertebrate Paleontology 22(4, Supplement). doi: 10.1080/02724634.2003.10010947
  • Brougham, T., E. T. Smith, and P. R. Bell. 2019. New theropod (Tetanurae: Avetheropoda) material from the ‘mid’-Cretaceous Griman Greek [sic (Creek)] Formation at Lightning Ridge, New South Wales, Australia. Royal Society Open Science 6:180826. doi: 10.1098/rsos.180826
  • Brusatte, S. L., R. B. J. Benson, and S. Hutt. 2008. The osteology of Neovenator salerii (Dinosauria: Theropoda) from the Wealden Group (Barremian) of the Isle of Wight. Palaeontographical Society Monographs 162:1–75.
  • Brusatte, S. L., T. D. Carr, and M. A. Norell. 2012. The osteology of Alioramus, a gracile and long-snouted tyrannosaurid (Dinosauria: Theropoda) from the Late Cretaceous of Mongolia. Bulletin of the American Museum of Natural History 366:1–197. doi: 10.1206/770.1
  • Brusatte, S. L., G. T. Lloyd, S. C. Wang, and M. A. Norell. 2014. Gradual assembly of avian body plan culminated in rapid rates of evolution across the dinosaur–bird transition. Current Biology 24:2386–2392. doi: 10.1016/j.cub.2014.08.034
  • Bryan, S. E., A. E. Constantine, C. J. Stephens, A. Ewart, R. W. Schon, and J. Parianos. 1997. Early Cretaceous volcano-sedimentary successions along the eastern Australian continental margin: implications for the break-up of eastern Gondwana. Earth and Planetary Science Letters 153:85–102. doi: 10.1016/S0012-821X(97)00124-6
  • Burch, S. H., and M. T. Carrano. 2012. An articulated pectoral girdle and forelimb of the abelisaurid theropod Majungasaurus crenatissimus from the Late Cretaceous of Madagascar. Journal of Vertebrate Paleontology 32:1–16. doi: 10.1080/02724634.2012.622027
  • Calvo, J. O., J. D. Porfiri, C. Veralli, F. Novas, and F. Poblete. 2004. Phylogenetic status of Megaraptor namunhuaiquii Novas based on a new specimen from Neuquén, Patagonia, Argentina. Ameghiniana 41:565–575.
  • Carrano, M. T., and J. Choiniere. 2016. New information on the forearm and manus of Ceratosaurus nasicornis Marsh, 1884 (Dinosauria, Theropoda), with implications for theropod forelimb evolution. Journal of Vertebrate Paleontology. doi: 10.1080/02724634.2015.1054497.
  • Carrano, M. T., and S. D. Sampson. 2008. The phylogeny of Ceratosauria (Dinosauria: Theropoda). Journal of Systematic Palaeontology 6:183–236. doi: 10.1017/S1477201907002246
  • Carrano, M. T., R. B. J. Benson, and S. D. Sampson. 2012. The phylogeny of Tetanurae (Dinosauria: Theropoda). Journal of Systematic Palaeontology 10:211–300. doi: 10.1080/14772019.2011.630927
  • Carrano, M. T., M. A. Loewen, and J. J. W. Sertich. 2011. New materials of Masiakasaurus knopfleri Sampson, Carrano, and Forster, 2001, and implications for the morphology of the Noasauridae (Theropoda: Ceratosauria). Smithsonian Contributions to Paleobiology 95:1–53. doi: 10.5479/si.00810266.95.1
  • Casal, G. A., R. D. Martínez, M. Luna, and L. M. Ibiricu. 2016. Ordenamiento y caracterización faunística del Cretácico Superior del Grupo Chubut, Cuenca del Golfo San Jorge, Argentina. Revista Brasileira de Paleontologia 19:53–70. doi: 10.4072/rbp.2016.1.05
  • Charig, A. J., and A. C. Milner. 1997. Baryonyx walkeri, a fish-eating dinosaur from the Wealden of Surrey. Bulletin of the Natural History Museum, London (Geology) 53:11–70.
  • Choiniere, J. N. 2010. Anatomy and systematics of coelurosaurian theropods from the Late Jurassic of Xinjiang, China, with comments on forelimb evolution in Theropoda. Ph.D. dissertation, George Washington University, Washington, D.C., 994 pp.
  • Close, R. A., P. Vickers-Rich, P. Trusler, L. M. Chiappe, J. O’Connor, T. H. Rich, L. Kool, and P. Komarower. 2009. Earliest Gondwanan bird from the Cretaceous of southeastern Australia. Journal of Vertebrate Paleontology 29:616–619. doi: 10.1671/039.029.0214
  • Colbert, E. H. 1989. The Triassic dinosaur Coelophysis. Museum of Northern Arizona Bulletin 57:1–160.
  • Colbert, E. H., and D. Merrilees. 1967. Cretaceous dinosaur footprints from Western Australia. Journal of the Royal Society of Western Australia 50:21–25.
  • Coombs, W. P., Jr., and R. E. Molnar. 1981. Sauropoda (Reptilia, Saurischia) from the Cretaceous of Queensland. Memoirs of the Queensland Museum 20:351–373.
  • Coria, R. A., and P. J. Currie. 2016. A new megaraptoran dinosaur (Dinosauria, Theropoda, Megaraptoridae) from the Late Cretaceous of Patagonia. PLoS ONE 11:e0157973. doi: 10.1371/journal.pone.0157973
  • Cuesta, E., F. Ortega, and J. L. Sanz. 2019. Axial osteology of Concavenator corcovatus (Theropoda; Carcharodontosauria) from the Lower Cretaceous of Spain. Cretaceous Research 95:106–120. doi: 10.1016/j.cretres.2018.10.026
  • Currie, P. J., and Y. Azuma. 2006. New specimens, including a growth series, of Fukuiraptor (Dinosauria, Theropoda) from the Lower Cretaceous Kitadani Quarry of Japan. Journal of the Paleontological Society of Korea 22:173–193.
  • Currie, P. J., and K. Carpenter. 2000. A new specimen of Acrocanthosaurus atokensis (Theropoda, Dinosauria) from the Lower Cretaceous Antlers Formation (Aptian) of Oklahoma, USA. Geodiversitas 22:207–246.
  • Currie, P. J., and X.-J. Zhao. 1993. A new carnosaur (Dinosauria, Theropoda) from the Jurassic of Xinjiang, People’s Republic of China. Canadian Journal of Earth Sciences 30:2037–2081. doi: 10.1139/e93-179
  • Debenham, N., N. J. C. Farrell, S. P. Holford, R. C. King, and D. Healy. 2019. Spatial distribution of micrometre-scale porosity and permeability across the damage zone of a reverse-reactivated normal fault in a tight sandstone: insights from the Otway Basin, SE Australia. Basin Research 31:640–658. doi: 10.1111/bre.12345
  • Dong, Z., S. Zhou, and Z. Zhang. 1983. Dinosaurs from the Jurassic of Sichuan. Palaeontologia Sinica (Series C) 162:1–145.
  • Duddy, I. R. 1983. The geology, petrology and geochemistry of the Otway Formation volcanogenic sediments. Ph.D. dissertation, University of Melbourne, Melbourne, Australia, 426 pp.
  • Duddy, I. R. 2003. Mesozoic; pp. 239–286 in W. D. Birch (ed.), Geology of Victoria, third edition. Geological Society of Australia (Victoria Division), Melbourne, Australia.
  • Edwards, A. B., and G. Baker. 1943. Jurassic arkose in southern Victoria. Proceedings of the Royal Society of Victoria 55:195–228.
  • Edwards, J., J. G. Leonard, G. R. Pettifer, and P. A. McDonald. 1996. Colac 1:250 000 Map. Geological Survey of Victoria Report 98:1–168.
  • Ezcurra, M. D., and F. E. Novas. 2016. Theropod dinosaurs from Argentina; pp. 139–156 in F. L. Agnolin, G. L. Lio, F. Brissón Egli, N. R. Chimento, and F. E. Novas (eds.), Historia evolutiva y paleobiogeográfica de los vertebrados de América del Sur. Museo Argentino de Ciencias Naturales ‘Bernardino Rivadavia,’ Buenos Aires, Argentina.
  • Felton, E. A. 1997a. A non-marine Lower Cretaceous rift-related epiclastic volcanic unit in southern Australia: the Eumeralla Formation in the Otway Basin. Part I: lithostratigraphy and depositional environments. AGSO Journal of Australian Geology & Geophysics 16:717–730.
  • Felton, E. A. 1997b. A non-marine Lower Cretaceous rift-related epiclastic volcanic unit in southern Australia: the Eumeralla Formation in the Otway Basin. Part II: fluvial systems. AGSO Journal of Australian Geology and Geophysics 16:731–757.
  • Flannery, T. F., and T. H. Rich. 1981. Dinosaur digging in Victoria. Australian Natural History 20:195–198.
  • Gao, Y. 1993. A new species of Szechuanosaurus from the Middle Jurassic of Dashanpu, Zigong, Sichuan. Vertebrata PalAsiatica 31:308–314.
  • Gauthier, J. 1986. Saurischian monophyly and the origin of birds; pp. 1–55 in K. Padian (ed.), The Origin of Birds and the Evolution of Flight. California Academy of Sciences, San Francisco, California.
  • Gilmore, C. W. 1920. Osteology of the carnivorous Dinosauria in the United States National Museum, with special reference to the genera Antrodemus (Allosaurus) and Ceratosaurus. Bulletin of the United States National Museum 110:1–154.
  • Gleadow, A. J. W., and I. R. Duddy. 1980. Early Cretaceous volcanism and the early breakup history of south-eastern Australia: evidence from fission-track dating of volcanoclastic sediments; pp. 295–300 in M. M. Cresswell and P. Vella (eds.), Gondwana Five: Proceedings of the Fifth International Geological Symposium, Wellington, February 11–16, New Zealand. Balkema, Rotterdam, The Netherlands.
  • Guinard, G. 2016. Limusaurus inextricabilis (Theropoda: Ceratosauria) gives a hand to evolutionary teratology: a complementary view on avian manual digits identities. Zoological Journal of the Linnean Society 176:674–685. doi: 10.1111/zoj.12329
  • Harris, J. D. 1998. A reanalysis of Acrocanthosaurus atokensis, its phylogenetic status, and paleobiogeographic implications, based on a new specimen from Texas. New Mexico Museum of Natural History and Science Bulletin 13:1–75.
  • Hedrick, B. P., L. E. Zanno, D. G. Wolfe, and P. Dodson. 2015. The slothful claw: osteology and taphonomy of Nothronychus mckinleyi and N. graffami (Dinosauria: Theropoda) and anatomical considerations for derived therizinosaurids. PLoS ONE 10:e0129449. doi: 10.1371/journal.pone.0129449
  • Hendrickx, C., O. Mateus, and R. Araújo. 2015. A proposed terminology of theropod teeth (Dinosauria, Saurischia). Journal of Vertebrate Paleontology. doi: 10.1080/02724634.2015.982797.
  • Herne, M. C., A. M. Tait, V. Weisbecker, M. Hall, J. P. Nair, M. Cleeland, and S. W. Salisbury. 2018. A new small-bodied ornithopod (Dinosauria, Ornithischia) from a deep, high-energy Early Cretaceous river of the Australian–Antarctic rift system. PeerJ 5:e4113. doi: 10.7717/peerj.4113
  • Hill, K. A., D. M. Finlayson, K. C. Hill, and G. T. Cooper. 1995. Mesozoic tectonics of the Otway Basin region: the legacy of Gondwana and the active Pacific margin—a review and ongoing research. APEA Journal 35:467–493.
  • Hocknull, S. A., M. A. White, T. R. Tischler, A. G. Cook, N. D. Calleja, T. Sloan, and D. A. Elliott. 2009. New mid-Cretaceous (latest Albian) dinosaurs from Winton, Queensland, Australia. PLoS ONE 4:e6190. doi: 10.1371/journal.pone.0006190
  • Hu, S.-y. 1964. Carnosaurian remains from Alashan, Inner Mongolia. Vertebrata PalAsiatica 8:42–63.
  • Ibrahim, N., P. C. Sereno, C. Dal Sasso, S. Maganuco, M. Fabbri, D. M. Martill, S. Zouhri, N. Myhrvold, and D. A. Iurino. 2014. Semiaquatic adaptations in a giant predatory dinosaur. Science 345:1613–1616. doi: 10.1126/science.1258750
  • Kellner, A. W. A. 2002. A review of avian Mesozoic feathers; pp. 389–404 in L. M. Chiappe and L. M. Witmer (eds.), Mesozoic Birds: Above the Heads of the Dinosaurs. University of California Press, Berkeley, California.
  • Korasidis, V. A., B. E. Wagstaff, S. J. Gallagher, I. R. Duddy, A.-M. P. Tosolini, D. J. Cantrill, and M. S. Norvick. 2016. Early angiosperm diversification in the Albian of southeast Australia: implications for flowering plant radiation across eastern Gondwana. Review of Palaeobotany and Palynology 232:61–80. doi: 10.1016/j.revpalbo.2016.04.005
  • Krassay, A. A., D. L. Cathro, and D. J. Ryan. 2004. A regional tectonostratigraphic framework for the Otway Basin; pp. 97–116 in P. J. Boult, D. R. Johns, and S. C. Lang (eds.), PESA’s Eastern Australasian Basins Symposium II: Conference Proceedings. Petroleum Exploration Society of Australia (PESA), 19–22 September, Adelaide, Australia.
  • Krumenacker, L. J., D. J. Simon, G. Scofield, and D. J. Varricchio. 2017. Theropod dinosaurs from the Albian–Cenomanian Wayan Formation of eastern Idaho. Historical Biology 29:170–186. doi: 10.1080/08912963.2015.1137913
  • Kurochkin, E. N., and R. E. Molnar. 1997. New material of enantiornithine birds from the Early Cretaceous of Australia. Alcheringa 21:291–297. doi: 10.1080/03115519708619169
  • Lamanna, M. C. 2004. Late Cretaceous dinosaurs and crocodyliforms from Egypt and Argentina. Ph.D. dissertation, University of Pennsylvania, Philadelphia, Pennsylvania, 292 pp.
  • Lambe, L. M. 1917. The Cretaceous theropodous dinosaur Gorgosaurus. Memoirs of the Geological Survey of Canada 100:1–84.
  • Long, J. A. 1992. First dinosaur bones from Western Australia. The Beagle, Records of the Northern Territory Museum of Arts and Sciences 9:21–27.
  • Long, J. A. 1995. A theropod dinosaur bone from the Late Cretaceous Molecap Greensand, Western Australia. Records of the Western Australian Museum 17:143–146.
  • Long, J. A., and A. R. I. Cruickshank. 1996. First record of an Early Cretaceous theropod dinosaur bone from Western Australia. Records of the Western Australian Museum 18:219–221.
  • Long, J. A., and R. E. Molnar. 1998. A new Jurassic theropod dinosaur from Western Australia. Records of the Western Australian Museum 19:121–129.
  • Madsen, J. H., Jr. 1976. Allosaurus fragilis: a revised osteology. Utah Geological Survey Bulletin 109:1–163.
  • Madsen, J. H., Jr., and S. P. Welles. 2000. Ceratosaurus (Dinosauria: Theropoda), a revised osteology. Utah Geological Survey, Miscellaneous Publication OO-2:1–80.
  • Marsh, O. C. 1881. Principal characters of American Jurassic dinosaurs. Part V. American Journal of Science 21 (series 3):417–423. doi: 10.2475/ajs.s3-21.125.417
  • Martin, A. J., P. Vickers-Rich, T. H. Rich, and M. Hall. 2014. Oldest known avian footprints from Australia: Eumeralla Formation (Albian), Dinosaur Cove, Victoria. Palaeontology 57:7–19. doi: 10.1111/pala.12082
  • Mitchell, M. M. 1997. Identification of multiple detrital sources for Otway Supergroup sedimentary rocks: implications for basin models and chronostratigraphic correlations. Australian Journal of Earth Sciences 44:743–750. doi: 10.1080/08120099708728351
  • Molnar, R. E. 1986. An enantiornithine bird from the Lower Cretaceous of Queensland, Australia. Nature 322:736–738. doi: 10.1038/322736a0
  • Molnar, R. E. 1991. Fossil reptiles in Australia; pp. 605–702 in P. Vickers-Rich, J. M. Monaghan, R. F. Baird, and T. H. Rich (eds.), Vertebrate Palaeontology of Australasia. Pioneer Design Studio, Melbourne, Australia.
  • Molnar, R. E. 1999. Avian tibiotarsi from the Early Cretaceous of Lightning Ridge, New South Wales; pp. 197–209 in Y. Tomida, T. H. Rich, and P. Vickers-Rich (eds.), Proceedings of the Second Gondwanan Dinosaur Symposium, National Science Museum, July 12–13, Tokyo, Japan.
  • Molnar, R. E., and N. S. Pledge. 1980. A new theropod dinosaur from South Australia. Alcheringa 4:281–287. doi: 10.1080/03115518008558972
  • Molnar, R. E., and S. W. Salisbury. 2005. Observations on Cretaceous sauropods from Australia; pp. 454–465 in V. Tidwell and K. Carpenter (eds.), Thunder-lizards: The Sauropodomorph Dinosaurs. Indiana University Press, Bloomington, Indiana.
  • Molnar, R. E., I. Obata, M. Tanimoto, and M. Matsukawa. 2009. A tooth of Fukuiraptor aff. F. kitadaniensis from the Lower Cretaceous Sebayashi Formation, Sanchu Cretaceous, Japan. Bulletin of Tokyo Gakugei University, Division of Natural Sciences 61:105–117.
  • Motta, M. J., A. M. Aranciaga Rolando, S. Rozadilla, F. E. Agnolín, N. R. Chimento, F. Brissón Egli, and F. E. Novas. 2016. New theropod fauna from the Upper Cretaceous (Huincul Formation) of northwestern Patagonia, Argentina. New Mexico Museum of Natural History and Science Bulletin 71:231–253.
  • Noll, C. A., and M. Hall. 2003. Fluvial architecture and the tectonic control on deposition of onshore Eumeralla Formation, Otway Ranges, Victoria: implications for exploration in the Early Cretaceous Otway Basin. The APPEA Journal 43:99–116. doi: 10.1071/AJ02005
  • Norvick, M. S., and M. A. Smith. 2001. Mapping the plate tectonic reconstruction of southern and southeastern Australia and implications for petroleum systems. The APPEA Journal 41:15–35. doi: 10.1071/AJ00001
  • Novas, F. E. 1998. Megaraptor namunhuaiquii, gen. et sp. nov., a large-clawed, Late Cretaceous theropod from Patagonia. Journal of Vertebrate Paleontology 18:4–9. doi: 10.1080/02724634.1998.10011030
  • Novas, F. E., A. M. Aranciaga Rolando, and F. L. Agnolín. 2016. Phylogenetic relationships of the Cretaceous Gondwanan theropods Megaraptor and Australovenator: the evidence afforded by their manual anatomy. Memoirs of Museum Victoria 74:49–61. doi: 10.24199/j.mmv.2016.74.05
  • Novas, F. E., M. D. Ezcurra, and A. Lecuona. 2008. Orkoraptor burkei nov. gen. et sp., a large theropod from the Maastrichtian Pari Aike Formation, southern Patagonia, Argentina. Cretaceous Research 29:468–480. doi: 10.1016/j.cretres.2008.01.001
  • Novas, F. E., F. L. Agnolín, M. D. Ezcurra, J. Porfiri, and J. I. Canale. 2013. Evolution of the carnivorous dinosaurs during the Cretaceous: the evidence from Patagonia. Cretaceous Research 45:174–215. doi: 10.1016/j.cretres.2013.04.001
  • Novas, F. E., L. Salgado, M. Suárez, F. L. Agnolín, M. D. Ezcurra, N. R. Chimento, R. De la Cruz, M. P. Isasi, A. O. Vargas, and D. Rubilar-Rogers. 2015. An enigmatic plant-eating theropod from the Late Jurassic period of Chile. Nature 522:331–334. doi: 10.1038/nature14307
  • O’Connor, P. M. 2007. The postcranial axial skeleton of Majungasaurus crenatissimus (Theropoda: Abelisauridae) from the Late Cretaceous of Madagascar; pp. 127–162 in S. D. Sampson and D. W. Krause (eds.), Majungasaurus crenatissimus (Theropoda : Abelisauridae) from the Late Cretaceous of Madagascar. Society of Vertebrate Paleontology Memoir 8. Journal of Vertebrate Paleontology 27(2, Supplement).
  • Ortega, F., F. Escaso, and J. L. Sanz. 2010. A bizarre, humped Carcharodontosauria (Theropoda) from the Lower Cretaceous of Spain. Nature 467:203–206. doi: 10.1038/nature09181
  • Perincek, D., and C. D. Cockshell. 1995. The Otway Basin: Early Cretaceous rifting to Neogene inversion. APEA Journal 35:451–466.
  • Perincek, D., B. Simons, and G. R. Pettifer. 1994. The tectonic framework, and associated play types of the Western Otway Basin, Victoria, Australia. APEA Journal 34:460–478.
  • Pol, D., and O. W. M. Rauhut. 2012. A Middle Jurassic abelisaurid from Patagonia and the early diversification of theropod dinosaurs. Proceedings of the Royal Society B: Biological Sciences 279:3170–3175. doi: 10.1098/rspb.2012.0660
  • Porfiri, J. D., D. Dos Santos, and J. O. Calvo. 2007. New information on Megaraptor namunhuaiquii (Theropoda: Tetanurae), Patagonia: considerations on palaeoecological aspects. Arquivos do Museu Nacional, Rio de Janeiro 65:545–550.
  • Porfiri, J. D., R. D. Juárez Valieri, D. D. D. Santos, and M. C. Lamanna. 2018. A new megaraptoran theropod dinosaur from the Upper Cretaceous Bajo de la Carpa Formation of northwestern Patagonia. Cretaceous Research 89:302–319. doi: 10.1016/j.cretres.2018.03.014
  • Porfiri, J. D., F. E. Novas, J. O. Calvo, F. L. Agnolin, M. D. Ezcurra, and I. A. Cerda. 2014. Juvenile specimen of Megaraptor (Dinosauria, Theropoda) sheds light about tyrannosauroid radiation. Cretaceous Research 51:35–55. doi: 10.1016/j.cretres.2014.04.007
  • Poropat, S. F., P. D. Mannion, P. Upchurch, S. A. Hocknull, B. P. Kear, and D. A. Elliott. 2015a. Reassessment of the non-titanosaurian somphospondylan Wintonotitan wattsi (Dinosauria: Sauropoda: Titanosauriformes) from the mid-Cretaceous Winton Formation, Queensland, Australia. Papers in Palaeontology 1:59–106. doi: 10.1002/spp2.1004
  • Poropat, S. F., S. K. Martin, A.-M. P. Tosolini, B. E. Wagstaff, L. B. Bean, B. P. Kear, P. Vickers-Rich, and T. H. Rich. 2018. Early Cretaceous polar biotas of Victoria, southeastern Australia—an overview of research to date. Alcheringa 42:157–229. doi: 10.1080/03115518.2018.1453085
  • Poropat, S. F., P. Upchurch, P. D. Mannion, S. A. Hocknull, B. P. Kear, T. Sloan, G. H. K. Sinapius, and D. A. Elliott. 2015b. Revision of the sauropod dinosaur Diamantinasaurus matildae Hocknull et al. 2009 from the middle Cretaceous of Australia: implications for Gondwanan titanosauriform dispersal. Gondwana Research 27:995–1033. doi: 10.1016/j.gr.2014.03.014
  • Poropat, S. F., J. P. Nair, C. E. Syme, P. D. Mannion, P. Upchurch, S. A. Hocknull, A. G. Cook, T. R. Tischler, and T. Holland. 2017. Reappraisal of Austrosaurus mckillopi Longman, 1933 from the Allaru Mudstone of Queensland, Australia’s first named Cretaceous sauropod dinosaur. Alcheringa 41:543–580. doi: 10.1080/03115518.2017.1334826
  • Poropat, S. F., P. D. Mannion, P. Upchurch, S. A. Hocknull, B. P. Kear, M. Kundrát, T. R. Tischler, T. Sloan, G. H. K. Sinapius, J. A. Elliott, and D. A. Elliott. 2016. New Australian sauropods shed light on Cretaceous dinosaur palaeobiogeography. Scientific Reports 6:34467. doi: 10.1038/srep34467
  • Rauhut, O. W. M., and M. T. Carrano. 2016. The theropod dinosaur Elaphrosaurus bambergi Janensch, 1920, from the Late Jurassic of Tendaguru, Tanzania. Zoological Journal of the Linnean Society 178:546–610. doi: 10.1111/zoj.12425
  • Rich, T. H., and P. V. Rich. 1989. Polar dinosaurs and biotas of the Early Cretaceous of southeastern Australia. National Geographic Research 5:15–53.
  • Rich, T. H., and P. Vickers-Rich. 1994. Neoceratopsians and ornithomimosaurs: dinosaurs of Gondwana origin? National Geographic Research and Exploration 10:129–131.
  • Rich, T. H., P. Vickers-Rich, T. F. Flannery, D. Pickering, L. Kool, A. M. Tait, and E. M. G. Fitzgerald. 2009. A fourth Australian Mesozoic mammal locality; pp. 677–681 in L. B. Albright III (ed.), Papers on Geology, Vertebrate Paleontology, and Biostratigraphy in Honor of Michael O. Woodburne. Museum of Northern Arizona, Flagstaff, Arizona.
  • Ruiz, J., A. Torices, H. Serrano, and V. López. 2011. The hand structure of Carnotaurus sastrei (Theropoda, Abelisauridae): implications for hand diversity and evolution in abelisaurids. Palaeontology 54:1271–1277. doi: 10.1111/j.1475-4983.2011.01091.x
  • Salisbury, S. W., A. Romilio, M. C. Herne, R. T. Tucker, and J. P. Nair. 2017. The Dinosaurian Ichnofauna of the Lower Cretaceous (Valanginian–Barremian) Broome Sandstone of the Walmadany Area (James Price Point), Dampier Peninsula, Western Australia. Society of Vertebrate Paleontology Memoir 16. Journal of Vertebrate Paleontology 36(Supplement 1):1–152. doi: 10.1080/02724634.2016.1269539
  • Samathi, A., P. Chanthasit, and P. M. Sander. 2019. Two new basal coelurosaurian theropod dinosaurs from the Lower Cretaceous Sao Khua Formation of Thailand. Acta Palaeontologica Polonica 64:239–260. doi: 10.4202/app.00540.2018
  • Sereno, P. C., J. A. Wilson, H. C. E. Larsson, D. B. Dutheil, and H.-D. Sues. 1994. Early Cretaceous dinosaurs from the Sahara. Science 266:267–271. doi: 10.1126/science.266.5183.267
  • Sereno, P. C., R. N. Martinez, J. A. Wilson, D. J. Varricchio, O. A. Alcober, and H. C. E. Larsson. 2008. Evidence for avian intrathoracic air sacs in a new predatory dinosaur from Argentina. PLoS ONE 3:e3303. doi: 10.1371/journal.pone.0003303
  • Sereno, P. C., D. B. Dutheil, M. Iarochene, H. C. E. Larsson, G. H. Lyon, P. M. Magwene, C. A. Sidor, D. J. Varricchio, and J. A. Wilson. 1996. Predatory dinosaurs from the Sahara and Late Cretaceous faunal differentiation. Science 272:986–991. doi: 10.1126/science.272.5264.986
  • Sereno, P. C., A. L. Beck, D. B. Dutheil, B. Gado, H. C. E. Larsson, G. H. Lyon, J. D. Marcot, O. W. M. Rauhut, R. W. Sadleir, C. A. Sidor, D. D. Varricchio, G. P. Wilson, and J. A. Wilson. 1998. A long-snouted predatory dinosaur from Africa and the evolution of spinosaurids. Science 282:1298–1302. doi: 10.1126/science.282.5392.1298
  • Smith, N. D., P. J. Makovicky, W. R. Hammer, and P. J. Currie. 2007. Osteology of Cryolophosaurus ellioti (Dinosauria: Theropoda) from the Early Jurassic of Antarctica and implications for early theropod evolution. Zoological Journal of the Linnean Society 151:377–421. doi: 10.1111/j.1096-3642.2007.00325.x
  • Smith, N. D., P. J. Makovicky, F. L. Agnolin, M. D. Ezcurra, D. F. Pais, and S. W. Salisbury. 2008. A Megaraptor-like theropod (Dinosauria: Tetanurae) in Australia: support for faunal exchange across eastern and western Gondwana in the Mid-Cretaceous. Proceedings of the Royal Society B 275:2085–2093. doi: 10.1098/rspb.2008.0504
  • Stromer, E. 1934. Ergebnisse der Forschungsreisen Prof. E. Stromers in den Wüsten Ägyptens. II. Wirbeltier-Reste der Baharîje-Stufe (unterstes Cenoman). 13. Dinosauria. Abhandlungen der Bayerischen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Abteilung 22 (Neue Folge):1–79.
  • Talent, J. A., P. M. Duncan, and P. L. Handby. 1966. Early Cretaceous feathers from Victoria. Emu 64:81–86. doi: 10.1080/01584197.1966.11797177
  • Thulborn, R. A. 2013. Lark Quarry revisited: a critique of methods used to identify a large dinosaurian track-maker in the Winton Formation (Albian–Cenomanian), western Queensland, Australia. Alcheringa 37:312–330. doi: 10.1080/03115518.2013.748482
  • Thulborn, R. A., and M. Wade. 1979. Dinosaur stampede in the Cretaceous of Queensland. Lethaia 12:275–279. doi: 10.1111/j.1502-3931.1979.tb01008.x
  • Thulborn, R. A., and M. Wade. 1984. Dinosaur trackways in the Winton Formation (mid-Cretaceous) of Queensland. Memoirs of the Queensland Museum 21:413–517.
  • Thulborn, T. 1990. Dinosaur Tracks. Chapman and Hall, London, 410 pp.
  • Thulborn, T. 2000. Australia’s earliest theropods: footprint evidence in the Ipswich Coal Measures (Upper Triassic) of Queensland; pp. 301–311 in B. P. Pérez-Moreno, T. R. Holtz Jr., J. L. Sanz, and J. J. Moratalla (eds.), Aspects of Theropod Paleobiology. Museu Nacional de História Natural, Lisbon, Portugal.
  • Thulborn, T. 2017. Behaviour of dinosaurian track-makers in the Winton Formation (Cretaceous, Albian–Cenomanian) at Lark Quarry, western Queensland, Australia: running or swimming? Ichnos 24(1):1–18. doi: 10.1080/10420940.2015.1129326
  • Torsvik, T. H., R. Van der Voo, U. Preeden, C. Mac Niocaill, B. Steinberger, P. V. Doubrovine, D. J. J. Van Hinsbergen, M. Domeier, C. Gaina, E. Tohver, J. G. Meert, P. J. A. McCausland, and L. R. M. Cocks. 2012. Phanerozoic polar wander, palaeogeography and dynamics. Earth-Science Reviews 114:325–368. doi: 10.1016/j.earscirev.2012.06.007
  • Tosolini, A.-M., A. Vadala, and S. McLoughlin. 1998. The Early Cretaceous (Barremian–Aptian) flora of the Binns Road Quarry, Otway Ranges, Victoria. The Fossil Collector 53:3–11.
  • Van Hinsbergen, D. J. J., L. V. de Groot, S. J. Van Schaik, W. Spakman, P. K. Bijl, A. Sluijs, C. G. Langereis, and H. Brinkhuis. 2015. A paleolatitude calculator for paleoclimate studies. PLoS ONE 10:e0126946. doi: 10.1371/journal.pone.0126946
  • Veevers, J. J., J. G. Jones, and C. M. Powell. 1982. Tectonic framework of Australia’s sedimentary basins. The APPEA Journal 22:283–300 doi: 10.1071/AJ81023
  • Veevers, J. J., C. M. Powell, and S. R. Roots. 1991. Review of seafloor spreading around Australia. I. Synthesis of the patterns of spreading. Australian Journal of Earth Sciences 38:373–389. doi: 10.1080/08120099108727979
  • Vickers-Rich, P. 1991. The Mesozoic and Tertiary history of birds on the Australian plate; pp. 721–808 in P. Vickers-Rich, J. M. Monaghan, R. F. Baird, and T. H. Rich (eds.), Vertebrate Palaeontology of Australasia. Pioneer Design Studio, Melbourne, Australia.
  • Waldman, M. 1970. A third specimen of a Lower Cretaceous feather from Victoria, Australia. Condor 72:377. doi: 10.2307/1366033
  • Wagstaff, B. E., and J. McEwen Mason. 1989. Palynological dating of Lower Cretaceous coastal vertebrate localities, Victoria, Australia. National Geographic Research 5:54–63.
  • Wagstaff, B. E., S. J. Gallagher, and J. K. Trainor. 2012. A new subdivision of the Albian spore-pollen zonation of Australia. Review of Palaeobotany and Palynology 171:57–72 doi: 10.1016/j.revpalbo.2011.11.003
  • Welles, S. P. 1984. Dilophosaurus wetherilli (Dinosauria, Theropoda) osteology and comparisons. Palaeontographica Abteilung A 185:85–180.
  • White, M. A., A. G. Cook, A. J. Klinkhamer, and D. A. Elliott. 2016. The pes of Australovenator wintonensis (Theropoda: Megaraptoridae): analysis of the pedal range of motion and biological restoration. PeerJ 4:e2312. doi: 10.7717/peerj.2312
  • White, M. A., P. R. Bell, A. G. Cook, S. F. Poropat, and D. A. Elliott. 2015a. The dentary of Australovenator wintonensis (Theropoda, Megaraptoridae); implications for megaraptorid dentition. PeerJ 3:e1512. doi: 10.7717/peerj.1512
  • White, M. A., P. L. Falkingham, A. G. Cook, S. A. Hocknull, and D. A. Elliott. 2013a. Morphological comparisons of metacarpal I for Australovenator wintonensis and Rapator ornitholestoides: implications for their taxonomic relationships. Alcheringa 37:435–441. doi: 10.1080/03115518.2013.770221
  • White, M. A., A. G. Cook, S. A. Hocknull, T. Sloan, G. H. K. Sinapius, and D. A. Elliott. 2012. New forearm elements discovered of holotype specimen Australovenator wintonensis from Winton, Queensland, Australia. PLoS ONE 7:e39364. doi: 10.1371/journal.pone.0039364
  • White, M. A., P. R. Bell, A. G. Cook, D. G. Barnes, T. R. Tischler, B. J. Bassam, and D. A. Elliott. 2015b. Forearm range of motion in Australovenator wintonensis (Theropoda, Megaraptoridae). PLoS ONE 10:e0137709. doi: 10.1371/journal.pone.0137709
  • White, M. A., R. B. J. Benson, T. R. Tischler, S. A. Hocknull, A. G. Cook, D. G. Barnes, S. F. Poropat, S. J. Wooldridge, T. Sloan, G. H. K. Sinapius, and D. A. Elliott. 2013b. New Australovenator hind limb elements pertaining to the holotype reveal the most complete neovenatorid leg. PLoS ONE 8:e68649. doi: 10.1371/journal.pone.0068649
  • Xu, X., J. M. Clark, C. A. Forster, M. A. Norell, G. M. Erickson, D. A. Eberth, C. Jia, and Q. Zhao. 2006. A basal tyrannosauroid dinosaur from the Late Jurassic of China. Nature 439:715–718. doi: 10.1038/nature04511
  • Zanno, L. E. 2010. Osteology of Falcarius utahensis (Dinosauria: Theropoda): characterizing the anatomy of basal therizinosaurs. Zoological Journal of the Linnean Society 158:196–230. doi: 10.1111/j.1096-3642.2009.00464.x
  • Zanno, L. E., and P. J. Makovicky. 2013. Neovenatorid theropods are apex predators in the Late Cretaceous of North America. Nature Communications 4: Article 2827. doi: 10.1038/ncomms3827

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