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Memoir Article

Anatomy, Relationships, and Paleobiology of Cambaytherium (Mammalia, Perissodactylamorpha, Anthracobunia) from the lower Eocene of western India

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Pages 1-147 | Received 12 Nov 2018, Accepted 03 Jan 2020, Published online: 05 Nov 2020

LITERATURE CITED

  • Adatte, K. H., H. Khozyem, J. E. Spangenberg, B. Samant, and G. Keller. 2014. Response of terrestrial environment to the Paleocene-Eocene Thermal Maximum (PETM), new insights from India and NE Spain. Rendiconti Online della Società Geologica Italiana 31:5–6.
  • Ali, J. R., and J. C. Aitchison. 2008. Gondwana to Asia: plate tectonics, paleogeography, and the biological connectivity of the Indian sub-continent from the Middle Jurassic through the latest Eocene (165–35 Ma). Earth-Science Reviews 88:145–166.
  • Alimohammadian, H., A. Sahni, R. Patnaik, R. S. Rana, and H. Singh. 2005. First record of an exceptionally diverse and well preserved amber-embedded biota from Lower Eocene (∼52 Ma) lignites, Vastan, Gujarat. Current Science 89:1328–1330.
  • Argot, C. 2012. Postcranial analysis of a carnivoran-like archaic ungulate: the case of Arctocyon primaevus (Arctocyonidae, Mammalia) from the Late Paleocene of France. Journal of Mammalian Evolution 20:83–114.
  • Asher, R. J., M. J. Novacek, and J. H. Geisler. 2003. Relationships of endemic African mammals and their fossil relatives based on morphological and molecular evidence. Journal of Mammalian Evolution 10:131–194.
  • Bai, B., Y.-Q. Wang, and J. Meng. 2010. New craniodental materials of Litolophus gobiensis (Perissodactyla, “Eomoropidae”) from Inner Mongolia, China and phylogenetic analysis of Eocene chalicotheres. American Museum Novitates 3688:1–27.
  • Bai, B., Y.-Q. Wang, and J. Meng. 2018. The divergence and dispersal of early perissodactyls as evidenced by early Eocene equids from Asia. Communications Biology 115:1–10.
  • Bai, B., J. Meng, Y.-Q. Wang, H.-B. Wang, and L. Holbrook. 2017. Osteology of the middle Eocene ceratomorph Hyrachyus modestus (Mammalia, Perissodactyla). Bulletin of the American Museum of Natural History 413:1–68.
  • Bajpai, S., and J. J. Head. 2007. An early Eocene palaeopheid snake from Vastan lignite mine, Gujarat, India. Gondwana Geological Magazine 22(2):85–90.
  • Bajpai, S., and V. V. Kapur. 2004. Oldest known gobiids from Vastan Lignite Mine (early Eocene), District Surat, Gujarat. Current Science 87:433–435.
  • Bajpai, S., and V. V. Kapur. 2008. Earliest Cenozoic frogs from the Indian subcontinent: implications for out-of-India hypothesis. Journal of the Palaeontological Society of India 53:65–71.
  • Bajpai, S., V. V. Kapur, and J. G. M. Thewissen. 2009. Creodont and condylarth from the Cambay Shale (early Eocene, ∼55–54 ma), Vastan Lignite Mine, Gujarat, western India. Journal of the Palaeontological Society of India 54:103–109.
  • Bajpai, S., D. P. Das, V. V. Kapur, B. N. Tiwari, and S. S. Srivastava. 2007. Early Eocene rodents (Mammalia) from Vastan Lignite Mine, Gujarat, western India. Gondwana Geological Magazine 22(2):91–95.
  • Bajpai, S., V. V. Kapur, J. G. M. Thewissen, D. P. Das, and B. N. Tiwari. 2006. New Early Eocene cambaythere (Perissodactyla, Mammalia) from the Vastan Lignite Mine (Gujarat, India) and an evaluation of cambaythere relationships. Journal of the Palaeontological Society of India 51:101–110.
  • Bajpai, S., V. V. Kapur, J. G. M. Thewissen, B. N. Tiwari, and D. P. Das. 2005a. First fossil marsupials from India: early Eocene Indodelphis n. gen. and Jaegeria n. gen. from Vastan Lignite Mine, District Surat, Gujarat. Journal of the Palaeontological Society of India 50:147–151.
  • Bajpai, S., V. V. Kapur, D. P. Das, B. N. Tiwari, N. Saravanan, and R. Sharma. 2005b. Early Eocene land mammals from Vastan Lignite Mine, District Surat (Gujarat), western India. Journal of the Palaeontological Society of India 50:101–113.
  • Bajpai, S., R. F. Kay, B. A. Williams, D. P. Das, V. V. Kapur, and B. N. Tiwari. 2008. The oldest Asian record of Anthropoidea. Proceedings of the National Academy of Sciences of the United States of America 105:11093–11098.
  • Bajpai, S., V. V. Kapur, J. G. M. Thewissen, D. P. Das, B. N. Tiwari, R. Sharma, and N. Saravanan. 2005c. Early Eocene primates from Vastan Lignite Mine, Gujarat, western India. Journal of the Palaeontological Society of India 50:43–54.
  • Beard, K. C. 1993. Phylogenetic systematics of the Primatomorpha, with special reference to Dermoptera; pp. 129–150 in F. S. Szalay, M. J. Novacek, and M. C. McKenna (eds.), Mammal Phylogeny—Placentals. Springer, New York.
  • Beard, K. C. 1998. East of Eden: Asia as an important center of taxonomic origination in mammalian evolution; pp. 5–39 in K. C. Beard and M. R. Dawson (eds.), Dawn of the Age of Mammals in Asia. Bulletin of Carnegie Museum of Natural History 34. Pittsburgh, Pennsylvania.
  • Bemis, K. E., J. C. Tyler, W. E. Bemis, K. Kumar, R. S. Rana, and T. Smith. 2017. A gymnodont fish jaw with remarkable molariform teeth from the early Eocene of Gujarat, India (Teleostei, Tetraodontiformes). Journal of Vertebrate Paleontology 37:e1369422.
  • Bertram, J. E. A., and A. Gutmann. 2009. Motions of the running horse and cheetah revisited: fundamental mechanics of the transverse and rotary gallop. Journal of the Royal Society Interface 6:549–559.
  • Biswas, S. K. 1987. Regional tectonic framework, structure and evolution of the western marginal basins of India. Tectonophysics 135:307–327.
  • Boszczyk, B. M., A. A. Boszczyk, and R. Putz. 2001. Comparative and functional anatomy of the mammalian lumbar spine. Anatomical Record 264:157–168.
  • Bowen, G. J., W. C. Clyde, P. L. Koch, S. Ting, J. Alroy, T. Tsubamoto, Y.-Q. Wang, and Y. Wang. 2002. Mammalian dispersal at the Paleocene/Eocene boundary. Science 295:2062–2065.
  • Bronnert, C., E. Gheerbrant, M. Godinot, and G. Métais. 2018. A primitive perissodactyl (Mammalia) from the early Eocene of Le Quesnoy (MP7, France). Historical Biology 30:237–250.
  • Buckley, M. 2015. Ancient collagen reveals evolutionary history of the endemic South American ‘ungulates.’ Proceedings of the Royal Society B: Biological Sciences 282:20142671.
  • Butler, P. M. 1956. The skull of Ictops and a classification of the Insectivora. Proceedings of the Zoological Society of London 126:453–481.
  • Campbell, B. 1936. The comparative myology of the forelimb of the hippopotamus, pig and tapir. American Journal of Anatomy 59:201–247.
  • Chandra, P. K., and L. R. Chowdhary. 1969. Stratigraphy of the Cambay Basin. Bulletin Oil and National Gas Commission (ONGC), Dehradun, India 6:37–50.
  • Chatterjee, S., A. Goswami, and C. R. Scotese. 2013. The longest voyage: tectonic, magmatic, and paleoclimatic evolution of the Indian plate during its northward flight from Gondwana to Asia. Gondwana Research 23:238–267.
  • Chatterjee, S., C. R. Scotese, and S. Bajpai. 2017. Indian plate and its epic voyage from Gondwana to Asia: its tectonic, paleoclimatic and paleobiogeographic evolution. Geological Society of America 529:1–147.
  • Clementz, M., S. Bajpai, V. Ravikant, J. G. M. Thewissen, N. Saravanan, I. B. Singh, and V. Prasad. 2011. Early Eocene warming events and the timing of terrestrial faunal exchange between India and Asia. Geology 39:15–18.
  • Colbert, M. 2005. The facial skeleton of the early Oligocene Colodon (Perissodactyla, Tapiroidea). Palaeontologia Electronica 8(1):12A. Available at https://palaeo-electronica.org/2005_1/colbert12/issue1_05.htm
  • Coombs, W. P., Jr., and M. C. Coombs. 1979. Pilgrimella, a primitive Asiatic perissodactyl. Zoological Journal of the Linnean Society 65:185–192.
  • Cooper, L. N., E. R. Seiffert, M. Clementz, S. I. Madar, S. Bajpai, S. T. Hussain, and J. G. M. Thewissen. 2014. Anthracobunids from the middle Eocene of India and Pakistan are stem perissodactyls. PLoS ONE 9:e109232.
  • Cope, E. D. 1884. The Vertebrata of the Tertiary formations of the West. Report of the U.S. Geological Survey of the Territories, Volume 3:1–1009.
  • Copley, A., J.-P. Avouae, and J.-Y. Royer. 2010. India-Asia collision and the Cenozoic slow down of the Indian plate: implications for the forces driving plate motions. Journal of Geophysical Research 115:B0310. doi: 10.1029/2009JB006634.
  • Court, N. 1992. The skull of Arsinoitherium (Mammalia, Embrithopoda) and the higher order interrelationships of ungulates. Palaeovertebrata 22:1–43.
  • Damuth, J. 1990. Problems in estimating body masses of archaic ungulates using dental measurements; pp. 229–253 in J. Damuth and B. J. MacFadden (eds.), Body Size in Mammalian Paleobiology: Estimation and Biological Implications. Cambridge University Press, Cambridge, U.K.
  • Dashzeveg, D., and J. J. Hooker. 1997. New ceratomorph perissodactyls (Mammalia) from the Middle and Late Eocene of Mongolia: their implications for phylogeny and dating. Zoological Journal of the Linnaean Society 120:105–138.
  • Davis, D. D. 1964. The giant panda. A morphological study of evolutionary mechanisms. Fieldiana Zoology Memoirs 3:1–339.
  • Dean, C. M. 1987. Growth layers and incremental markings in hard tissues; a review of the literature and some preliminary observations about enamel structure in Paranthropus boisei. Journal of Human Evolution 16:157–172.
  • Dehm, R., and T. Oettingen-Spielberg. 1958. Paläontologische und geologische Untersuchungen in Tertiär von Pakistan. 2. Die mitteleocänen Säugetiere von Ganda Kas bei Basal in Nordwest-Pakistan. Abhandlungen Bayerischen. Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse 91:1–54.
  • Ding, L., M. Qasim, I. A. K. Jadoon, M. A. Khan, Q. Xu, F. Cai, H. Wang, U. Baral, and Y. Yue. 2016. The India-Asia collision in north Pakistan: insight from the U–Pb detrital zircon provenance of Cenozoic foreland basin. Earth and Planetary Science Letters 455:49–61.
  • Dirks, W., R. L. Anemone, P. A. Holroyd, D. J. Reid, and P. Walton. 2009. Phylogeny, life history and the timing of molar crown formation in two archaic ungulates, Meniscotherium and Phenacodus (Mammalia, ‘Condylarthra’); pp. 3–8 in T. Koppe, G. Meyer, and K. W. Alt (eds.), Comparative Dental Morphology. Karger, Basel, Switzerland.
  • Domning, D. P., C. E. Ray, and M. C. McKenna. 1986. Two new Oligocene desmostylians and a discussion of tethytherian systematics. Smithsonian Contributions to Paleobiology 59:1–56.
  • Ducrocq, S., A. N. Soe, B. Bo, M. Benammi, Y. Chaimanee, T. Tun, T. Thein, and J.-J. Jaeger. 2000. First record of an Anthracobunidae (Mammalia, ?Tethytheria) from the Eocene of the Pondaung Formation, Myanmar. Comptes Rendus de l'Académie des Sciences, Paris, Sciences de la Terre et des Planètes 330:725–730.
  • Dunn, R. H., K. D. Rose, R. S. Rana, K. Kumar, A. Sahni, and T. Smith. 2016. New euprimate postcrania from the early Eocene of Gujarat, India, and the strepsirrhine-haplorhine divergence. Journal of Human Evolution 99:25–51.
  • Dutta, S., S. M. Tripathi, M. Mallick, R. P. Matthews, P. F. Greenwood, M. R. Rao, and R. E. Summons. 2011. Eocene out-of-India dispersal of Asian dipterocarps. Review of Palaeobotany and Palynology 166:63–68.
  • Eriksson, T. 2001. AutoDecay ver. 5.0 (program distributed by the author). Bergius Foundation, Royal Swedish Academy of Sciences, Stockholm, Sweden.
  • Evans, H. E., and G. C. Christensen. 1979. Miller’s Anatomy of the Dog, second edition. W. B. Saunders, Philadelphia, Pennsylvania, 1181 pp.
  • Fischer, M. S. 1986. Die Stellung der Schliefer (Hyracoidea) im phylogenetischen System der Eutheria. Zugleich ein Beitrag zur Anpassungsgeschichte der Procaviidae. Courier Forschungsinstitut Senckenberg 84:1–132.
  • Fischer, M. S. 1989. Hyracoids, the sister-group of perissodactyls; pp. 37–56 in D. R. Prothero and R. M. Schoch (eds.), The Evolution of Perissodactyls. Oxford University Press, New York.
  • Fischer, M. S., and P. Tassy. 1993. The interrelation between Proboscidea, Sirenia, Hyracoidea, and Mesaxonia: the morphological evidence; pp. 217–234 in F. S. Szalay, M. J. Novacek, and M. C. McKenna (eds.), Mammal Phylogeny: Placentals. Springer, New York.
  • Fisher, R. E., K. M. Scott, and V. L. Naples. 2007. Forelimb myology of the pygmy hippopotamus (Choeropsis liberiensis). The Anatomical Record 290:673–693.
  • Fisher, R. E., B. Adrian, M. Barton, J. Holmgren, and S. Y. Tang. 2009. The phylogeny of the red panda (Ailurus fulgens): evidence from the forelimb. Journal of Anatomy 215:611–635.
  • Folie, A., R. S. Rana, K. D. Rose, A. Sahni, K. Kumar, L. Singh, and T. Smith. 2013. Early Eocene frogs from Vastan Lignite Mine, Gujarat, India. Acta Palaeontologica Polonica 58:511–524.
  • Franzen, J. L. 1989. Origin and systematic position of the Palaeotheriidae; pp. 102–108 in D. R. Prothero and R. M. Schoch (eds.), The Evolution of Perissodactyls. Oxford University Press, New York.
  • Franzen, J. L. 2007. Eozäne Equoidea (Mammalia, Perissodactyla) aus der Grube Messel bei Darmstadt (Deutschland). Funde der Jahre 1969–2000. Schweizerische Paläontologische Abhandlungen 127:1–245.
  • Franzen, J. L., and J. Habersetzer. 2017. Complete skeleton of Eurohippus messelensis (Mammalia, Perissodactyla, Equoidea) from the early middle Eocene of Grube Messel (Germany). Palaeobiodiversity and Palaeoenvironments 97:807–832.
  • Frischauf, C., R. Gockert, N. Kavcik-Graumann, and G. Rabeder. 2016. “Kiskevély Knives” indicate the menu of Alpine cave bears: comparative studies on wedge shaped defects of canines and incisors. International Cave Bear Symposium Proceedings 2016:14–17.
  • Froehlich, D. J. 1999. Phylogenetic systematics of basal perissodactyls. Journal of Vertebrate Paleontology 19:140–159.
  • Froehlich, D. J. 2002. Quo vadis eohippus? The systematics and taxonomy of the early Eocene equids (Perissodactyla). Zoological Journal of the Linnean Society 134:141–256.
  • Garg, R., K. Ateequzzaman, V. Prasad, S. K. M. Tripathi, I. B. Singh, A. K. Jauhri, and S. Bajpai. 2008. Age-diagnostic dinoflagellate cysts from lignite-bearing sediments of the Vastan lignite mine, Surat District, Gujarat, western India. Journal of the Palaeontological Society of India 53:99–105.
  • Gazin, C. L. 1953. The Tillodontia: an early Tertiary order of mammals. Smithsonian Miscellaneous Collections 121(10):1–110.
  • Gazin, C. L. 1965. A study of the early Tertiary condylarthran mammal Meniscotherium. Smithsonian Miscellaneous Collections 149(2):1–98.
  • Getty, R. 1975. Sisson and Grossman’s The Anatomy of the Domestic Animals. W. B. Saunders, Philadelphia, Pennsylvania, 2,130 pp.
  • Gheerbrant, E. 2009. Paleocene emergence of elephant relatives and the rapid radiation of African ungulates. Proceedings of the National Academy of Sciences of the United States of America 106:10717–10721.
  • Gheerbrant, E., B. Bouya, and M. Amaghzaz. 2012. Dental and cranial anatomy of Eritherium azzouzorum from the Paleocene of Morocco, earliest known proboscidean mammal. Palaeontographica, Abteilung A: Paläozoologie–Stratigraphie 297:151–183.
  • Gheerbrant, E., D. P. Domning, and P. Tassy. 2005b. Paenungulata (Sirenia, Proboscidea, Hyracoidea, and relatives); pp. 84–105 in K. D. Rose and J. D. Archibald (eds.), The Rise of Placental Mammals: Origins and Relationships of the Major Extant Clades. Johns Hopkins University Press, Baltimore, Maryland.
  • Gheerbrant, E., A. Filippo, and A. Schmitt. 2016. Convergence of afrotherian and laurasiatherian ungulate-like mammals: first morphological evidence from the Paleocene of Morocco. PLoS ONE 11:e0157556.
  • Gheerbrant E., J. Sudre, and H. Cappetta. 1996. A Palaeocene proboscidean from Morocco. Nature 383:68–71.
  • Gheerbrant, E., J. Sudre, P. Tassy, M. Amaghzaz, B. Bouya, and M. Iarochène. 2005a. Nouvelles données sur Phosphatherium escuilliei (Mammalia, Proboscidea) de l’Éocène inférieur du Maroc, apports à la phylogénie des Proboscidea et des ongulés lophodontes. Geodiversitas 27:239–333.
  • Gingerich, P. D. 1974. Size variability of the teeth in living mammals and the diagnosis of closely related sympatric fossil species. Journal of Paleontology 48:895–903.
  • Gingerich, P. D. 1976. Cranial anatomy and evolution of early Tertiary Plesiadapidae (Mammalia, Primates). University of Michigan Papers on Paleontology 15:1–141.
  • Gingerich, P. D. 1977. A small collection of fossil vertebrates from the middle Eocene Kuldana and Kohat formations of Punjab (Pakistan). Contributions from the Museum of Paleontology, The University of Michigan 24:190–203.
  • Gingerich, P. D. 1986. Early Eocene Cantius torresi—oldest primate of modern aspect from North America. Nature 320:319–321.
  • Gingerich, P. D. 1989. New earliest Wasatchian mammalian fauna from the Eocene of northwestern Wyoming: composition and diversity in a rarely sampled high-floodplain assemblage. University of Michigan Papers on Paleontology 28:1–97.
  • Gingerich, P. D. 1991. Systematics and evolution of early Eocene Perissodactyla (Mammalia) in the Clarks Fork Basin, Wyoming. Contributions from the Museum of Paleontology, The University of Michigan 28:181–213.
  • Gingerich, P. D. 2006. Environment and evolution through the Paleocene-Eocene thermal maximum. Trends in Ecology and Evolution 21:246–253.
  • Gingerich, P. D. 2010. Mammalian faunal succession through the Paleocene-Eocene Thermal Maximum (PETM) in western North America. Vertebrata PalAsiatica 48:308–327.
  • Gingerich, P. D., S. G. Abbas, and M. Arif. 1997. Early Eocene Quettacyon parachai (Condylarthra) from the Ghazij Formation of Baluchistan (Pakistan): oldest Cenozoic land mammal from South Asia. Journal of Vertebrate Paleontology 17:629–637.
  • Gingerich, P. D., D. E. Russell, and N. A. Wells. 1990. Astragalus of Anthracobune (Mammalia, Proboscidea) from the early–middle Eocene of Kashmir. Contributions from the Museum of Paleontology, The University of Michigan 28:71–77.
  • Gingerich, P. D., M. Arif, I. H. Khan, and S. G. Abbas. 1998. First early Eocene land mammals from the upper Ghazij Formation of the Sor Range, Baluchistan; pp. 1–17 in M. I. Ghaznavi, S. M. Raza, and M. T. Hasan (eds.), Siwaliks of South Asia. Proceedings of the Third GEOSAS Workshop held at Islamabad, Pakistan, March 1997. Geological Survey of Pakistan, Islamabad, Pakistan.
  • Gingerich, P. D., M. Arif, I. H. Khan, W. C. Clyde, and J. I. Bloch. 1999. Machocyon abbasi, a new early Eocene quettacyonid (Mammalia, Condylarthra). Contributions from the Museum of Paleontology, The University of Michigan 30:233–250.
  • Gingerich, P. D., M. Arif, I. H. Khan, M. Ul-Haq, J. I. Bloch, W. C. Clyde, and G. F. Gunnell. 2001. Gandhera Quarry, a unique mammalian faunal assemblage from the early Eocene of Baluchistan (Pakistan); pp. 251–262 in G. F. Gunnell (ed.), Eocene Biodiversity: Unusual Occurrences and Rarely Sampled Habitats. Kluwer Academic/Plenum Publishers, New York.
  • Ginsburg, L., K. H. Durrani, A. M. Kassi, and J.-L. Welcomme. 1999. Discovery of a new Anthracobunidae (Tethytheria, Mammalia) from the lower Eocene lignite of the Kach-Harnai area in Baluchistan (Pakistan). Comptes Rendus de l’Académie des Sciences. Série 2, Sciences de la Terre et Planètes 328:209–213.
  • Goloboff, P., J. S. Farris, and K. Nixon. 2008. TNT: a free program for phylogenetic analysis. Cladistics 24:774–786.
  • Granger, W. 1908. American Eocene horses. Bulletin of the American Museum of Natural History 24:221–264.
  • Halpert, A. P., F. A. Jenkins Jr., and H. Franks. 1987. Structure and scaling of the lumbar vertebrae in African bovids (Mammalia, Artiodactyla). Journal of Zoology 211:239–258.
  • Hermanson, J. W., and B. J. MacFadden. 1992. Evolutionary and functional morphology of the shoulder region and stay-apparatus in fossil and extant horses (Equidae). Journal of Vertebrate Paleontology 12:377–386.
  • Hildebrand, M. 1959. Motions of the running cheetah and horse. Journal of Mammalogy 40:481–495.
  • Holbrook, L. T. 1999. The phylogeny and classification of tapiromorph perissodactyls (Mammalia). Cladistics 15:331–350.
  • Holbrook, L. T. 2001. Comparative osteology of early Tertiary tapiromorphs (Mammalia, Perissodactyla). Zoological Journal of the Linnean Society 132:1–54.
  • Holbrook, L. T. 2009. Osteology of Lophiodon Cuvier, 1822 (Mammalia, Perissodactyla) and its phylogenetic implications. Journal of Vertebrate Paleontology 29:212–230.
  • Holbrook, L. T. 2014. On the skull of Radinskya (Mammalia) and its phylogenetic position. Journal of Vertebrate Paleontology 34:1203–1215.
  • Holbrook, L. T. 2015. The identity and homology of the postprotocrista and its role in molarization of upper premolars of Perissodactyla (Mammalia). Journal of Mammalian Evolution 22:259–269.
  • Holbrook, L. T., and J. B. LaPergola. 2011. A new genus of perissodactyl (Mammalia) from the Bridgerian of Wyoming, with comments on basal perissodactyl phylogeny. Journal of Vertebrate Paleontology 31:895–901.
  • Holland, W. J., and O. A. Peterson. 1914. The osteology of the Chalicotheroidea, with special reference to a mounted skeleton of Moropus elatus Marsh, now installed in the Carnegie Museum. Memoirs of the Carnegie Museum 3:189–406.
  • Hooker, J. J. 1984. A primitive ceratomorph (Perissodactyla, Mammalia) from the early Tertiary of Europe. Zoological Journal of the Linnean Society 82:229–244.
  • Hooker, J. J. 1989. Character polarities in early perissodactyls and their significance for Hyracotherium and infraordinal relationships; pp. 79–101 in D. R. Prothero and R. M. Schoch (eds.), The Evolution of Perissodactyls. Oxford University Press, New York.
  • Hooker, J. J. 1994. The beginning of the equoid radiation. Zoological Journal of the Linnean Society 112:29–63.
  • Hooker, J. J. 1996. Mammalian biostratigraphy across the Paleocene-Eocene boundary in the Paris, London and Belgian basins; pp. 205–218 in R. W. O’B. Knox, R. M. Corfield, and R. E. Dunay (eds.), Correlation of the Early Paleogene in Northwest Europe. Geological Society Special Publication 101. The Geological Society, London.
  • Hooker, J. J. 2005. Perissodactyla; pp. 199–214 in K. D. Rose and J. D. Archibald (eds.), The Rise of Placental Mammals: Origins and Relationships of the Major Extant Clades. Johns Hopkins University Press, Baltimore, Maryland.
  • Hooker, J. J. 2010. The Mammal Fauna of the Early Eocene Blackheath Formation of Abbey Wood, London. Monograph of the Palaeontographical Society, London, 162 pp. + 4 pls.
  • Hooker, J. J. 2015. A two-phase mammalian dispersal event across the Paleocene-Eocene transition. Newsletters on Stratigraphy 48:201–220.
  • Hooker, J. J., and M. E. Collinson. 2012. Mammalian faunal turnover across the Paleocene-Eocene boundary in NW Europe: the roles of displacement, community evolution and environment. Austrian Journal of Earth Sciences 105:17–28.
  • Hooker, J. J., and D. Dashzeveg. 2003. Evidence for direct mammalian faunal interchange between Europe and Asia near the Paleocene-Eocene boundary; pp. 479–500 in S. L. Wing, P. D. Gingerich, B. Schmitz, and E. Thomas (eds.), Causes and Consequences of Globally Warm Climates in the Early Paleogene. Geological Society of America Special Paper 369. Boulder, Colorado.
  • Hooker, J. J., and D. Dashzeveg. 2004. The origin of chalicotheres (Perissodactyla, Mammalia). Palaeontology 47:1363–1386.
  • Howell, A. B. 1944. Speed in Animals. University of Chicago Press, Chicago, Illinois, 270 pp.
  • Hu, X., E. Garzanti, T. Moore, and I. Raffi. 2015. Direct stratigraphic dating of India-Asia collision onset at the Selandian (middle Paleocene, 59 ± 1 Ma). Geology 43:859–862.
  • Hu, X., E. Garzanti, J. Wang, W. Huang, W. An, and A. Webb. 2016. The timing of India-Asia collision onset—facts, theories, controversies. Earth-Science Reviews 160:264–299.
  • Hussain, S. T. 1975. Evolutionary and functional anatomy of the pelvic limb in fossil and recent Equidae (Perissodactyla, Mammalia). Anatomia, Histologia, Embryologia 4:179–222.
  • Iakovleva, A. I., H. Brinkhuis, and C. Cavagnetto. 2001. Late Palaeocene–early Eocene dinoflagellate cysts from the Turgay Strait, Kazakhstan; correlations across ancient seaways. Palaeogeography, Palaeoclimatology, Palaeoecology 172:243–268.
  • IBM Corp. 2017. IBM SPSS Statistics for Windows, version 25.0. IBM Corp., Armonk, New York.
  • International Commission on Zoological Nomenclature. 2012. International Code of Zoological Nomenclature, fourth edition. Available at www.iczn.org/the-code/the-international-code-of-zoological-nomenclature/the-code-online. Accessed August 15, 2018.
  • International Committee on Veterinary Gross Anatomical Nomenclature. 2012. Nomina Anatomica Veterinaria, fifth edition, revised. Editorial Committee of ICVGAN, Hannover, Germany.
  • Jenkins, F. A., Jr., and S. M. Camazine. 1977. Hip structure and locomotion in ambulatory and cursorial carnivores. Journal of Zoology, London 181:351–370.
  • Jones, K. E. 2015a. Evolutionary allometry of lumbar shape in Felidae and Bovidae. Biological Journal of the Linnean Society London 116:721–740.
  • Jones, K. E. 2015b. Preliminary data on the effect of osseous anatomy on ex vivo joint mobility in the equine thoracolumbar region. Equine Veterinary Journal 48:502–508.
  • Jones, K. E. 2016. New insights on equid locomotor evolution from the lumbar region of fossil horses. Proceedings of the Royal Society B: Biological Sciences 283:20152947.
  • Jones, K. E., and L. T. Holbrook. 2016. The evolution of lateral accessory articulations in the lumbar region of perissodactyls. Journal of Vertebrate Paleontology. doi: 10.1080/02724634.2016.1224892.
  • Kapur, V. V., and S. Bajpai. 2015. Oldest south Asian tapiromorph (Perissodactyla, Mammalia) from the Cambay Shale Formation, western India, with comments on its phylogenetic position and biogeographic implications. The Paleobotanist 64:95–103.
  • Kapur, V. V., D. P. Das, S. Bajpai, and G. V. R. Prasad. 2017a. First mammal of Gondwanan lineage in the early Eocene of India. Comptes Rendus Palevol 16:721–737.
  • Kapur, V. V., D. P. Das, S. Bajpai, and G. V. R. Prasad. 2017b. Corrigendum to “First Mammal of Gondwanan lineage in the early Eocene of India” [C. R. Palevol, Kapur et al. 16 (2017)]. Comptes Rendus Palevol 16:820.
  • Kitts, D. B. 1956. American Hyracotherium (Perissodactyla, Equidae). Bulletin of the American Museum of Natural History 110:1–60.
  • Kneepkens, A. F. L. M., D. M. Badoux, and A. A. MacDonald. 1989. Descriptive and comparative myology of the forelimb of the Babirusa (Babyrousa babyrussa L., 1758). Anatomia, Histologia, Embryologia 18:349–365.
  • Koenigswald, W. von. 1997. Brief survey of the enamel diversity at the schmelzmuster level in Cenozoic placental mammals; pp. 137–161 in W. von Koenigswald and P. Sander (eds.), Tooth Enamel Microstructure. Balkema, Rotterdam, The Netherlands.
  • Koenigswald, W. von, and K. D. Rose. 2005. The enamel microstructure of the early Eocene pantodont Coryphodon and the nature of the zigzag-enamel. Journal of Mammalian Evolution 12:419–432.
  • Koenigswald, W. von, L. T. Holbrook, and K. D. Rose. 2011. Diversity and evolution of Hunter-Schreger band configuration in tooth enamel of perissodactyl mammals. Acta Palaeontologica Polonica 56:11–32.
  • Koenigswald, W. von, K. D. Rose, L. T. Holbrook, K. Kumar, R. S. Rana, and T. Smith. 2018. Mastication and enamel structure in Cambaytherium, a perissodactyl-like ungulate from the early Eocene of India. Palaeontologische Zeitschrift 92:671–680.
  • Kondrashov, P., and S. G. Lucas. 2012. Nearly complete skeleton of Tetraclaenodon (Mammalia, Phenacodontidae) from the early Paleocene of New Mexico: morpho-functional analysis. Journal of Paleontology 86:25–43.
  • Koob, T. J., and J. H. Long. 2000. The vertebrate body axis: evolution and mechanical function. American Zoologist 40:1–18.
  • Krause, D. W., and M. C. Maas. 1990. The biogeographic origins of late Paleocene–early Eocene mammalian immigrants to the Western Interior of North America; pp. 71–105 in T. M. Bown and K. D. Rose (eds.), Dawn of the Age of Mammals in the Northern Part of the Rocky Mountain Interior, North America. Geological Society of America Special Paper 243. Boulder, Colorado.
  • Kumar, K. 1991. Anthracobune aijiensis nov. sp. (Mammalia: Proboscidea) from the Subathu Formation, Eocene from NW Himalaya, India. Geobios 24:221–239.
  • Kumar, K., H. Singh, and R. S. Rana. 2011. Ichnospecies Teredolites longissimus and teredinid body fossils from the Early Eocene of India—taphonomic and palaeoenvironmental implications. Ichnos 18:57–71.
  • Kumar, K., K. D. Rose, R. S. Rana, L. Singh, T. Smith, and A. Sahni. 2010. Early Eocene artiodactyls (Mammalia) from western India. Journal of Vertebrate Paleontology 30:1245–1274.
  • Ladevèze, S., P. Missiaen, and T. Smith. 2010. First skull of Orthaspidotherium edwardsi (Mammalia, Condylarthra) from the late Paleocene of Berru (France) and phylogenetic affinities of the enigmatic European family Pleuraspidotheriidae. Journal of Vertebrate Paleontology 30:1559–1578.
  • Luckett, W. P. 1993. An ontogenetic assessment of dental homologies in therian mammals; pp. 182–204 in F. S. Szalay, M. J. Novacek, and M. C. McKenna (eds.), Mammal Phylogeny: Mesozoic Differentiation, Multituberculates, Monotremes, Early Therians, and Marsupials. Springer, New York.
  • Macdonald, A. A., and A. F. L. M. Kneepkens. 1995. Descriptive and comparative myology of the hindlimb of the babirusa (Babyrousa babyrussa L. 1758). Anatomia, Histologia, Embryologia 24:197–207.
  • MacFadden, B. J. 1976. Cladistic analysis of primitive equids, with notes on other perissodactyls. Systematic Zoology 25:1–14.
  • MacPhee, R. D. E. 1994. Morphology, adaptations, and relationships of Plesiorycteropus, and a diagnosis of a new order of eutherian mammals. Bulletin of the American Museum of Natural History 220:1–214.
  • Mallick, M., S. Dutta, P. F. Greenwood, and N. Bertram. 2009. Pyrolytic and spectroscopic studies of Eocene resin from Vastan Lignite Mine, Cambay Basin, western India. Journal Geological Society of India 74:16–22.
  • Mao, F.-Y., Y.-Q. Wang, Q. Li, and X. Jin. 2016. New records of archaic ungulates from the Lower Eocene of Sanshui Basin, Guangdong, China. Historical Biology 28:787–802.
  • Marsh, O. C. 1884. Dinocerata. A Monograph of an Extinct Order of Gigantic Mammals. Monographs of the United States Geological Survey 10, Washington, D.C., 237 pp.
  • Matthew, W. D. 1915. A revision of the Lower Eocene Wasatch and Wind River faunas. Part I: Order Ferae (Carnivora). Suborder Creodonta. Bulletin of the American Museum of Natural History 34:1–103.
  • Matthew, W. D. 1917. Absence of the pollex in Perissodactyla. Bulletin of the American Museum of Natural History 37:573–577.
  • Matthew, W. D. 1937. Paleocene faunas of the San Juan Basin, New Mexico. Transactions of the American Philosophical Society (new series) 30:1–510.
  • Mayr, G., R. S. Rana, A. Sahni, and T. Smith. 2007. Oldest fossil avian remains from the Indian subcontinental plate. Current Science 92:1266–1269.
  • Mayr, G., R. S. Rana, K. D. Rose, A. Sahni, K. Kumar, and T. Smith. 2013. New specimens of the early Eocene bird Vastanavis and the interrelationships of stem group Psittaciformes. Paleontological Journal 47:1308–1314.
  • Mayr, G., R. S. Rana, K. D. Rose, A. Sahni, K. Kumar, L. Singh, and T. Smith. 2010. Quercypsitta-like birds from the early Eocene of India (Aves, ?Psittaciformes). Journal of Vertebrate Paleontology 30:467–478.
  • McCann, T. 2010. Chenier plain sedimentation in the Palaeogene of western India. Zeitschrift der Deutschen Gesellschaft fur Geowissenschaften 161:335–351.
  • McKenna, M. C. 1975. Fossil mammals and early Eocene North Atlantic land continuity. Annals of the Missouri Botanical Garden 62:335–353.
  • McKenna, M. C., and S. K. Bell. 1997. Classification of Mammals above the Species Level. Columbia University Press, New York, 631 pp.
  • McKenna, M. C., M. Chow, S. Ting, and Z.-X. Luo. 1989. Radinskya yupingae, a perissodactyl-like mammal from the late Paleocene of China; pp. 24–36 in D. R. Prothero and R. M. Schoch (eds.), The Evolution of Perissodactyls. Oxford University Press, New York.
  • Mead, J. G., and R. E. Fordyce. 2009. The therian skull: a lexicon with emphasis on the odontocetes. Smithsonian Contributions to Zoology 627:1–249.
  • Meng, J., C. Wang, and X. Zhao. 2012. India-Asia collision was at 24° N and 50 Ma: palaeomagnetic proof from southernmost Asia. Scientific Reports 2:925.
  • Meng, J., R. Zhai, and A. Wyss. 1998. The late Paleocene Bayan Ulan fauna of Inner Mongolia, China; pp. 148–185 in K. C. Beard and M. R. Dawson (eds.), Dawn of the Age of Mammals in Asia. Bulletin of Carnegie Museum of Natural History 34. Pittsburgh, Pennsylvania.
  • Meredith, R. W., J. Janecka, J. Gatesy, O. A. Ryder, C. A. Fisher, E. C. Teeling, A. Goodbla, E. Eizirik, T. L. L. Simão, T. Stadler, D. L. Rabosky, R. L. Honeycutt, J. J. Flynn, C. M. Ingram, C. Steiner, T. L. Williams, T. J. Robinson, A. Burk-Herrick, M. Westerman, N. A. Ayoub, M. S. Springer, and W. J. Murphy. 2011. Impacts of the Cretaceous terrestrial revolution and KPg extinction on mammal diversification. Science 334:521–524.
  • Missiaen, P., and P. D. Gingerich. 2012. New early Eocene tapiromorph perissodactyls from the Ghazij Formation of Pakistan, with implications for mammalian biochronology in Asia. Acta Palaeontologica Polonica 57:21–34.
  • Missiaen, P., and P. D. Gingerich. 2014. New basal Perissodactyla (Mammalia) from the Lower Eocene Ghazij Formation of Pakistan. Contributions from the Museum of Paleontology, The University of Michigan 32:139–160.
  • Missiaen, P., G. F. Gunnell, and P. D. Gingerich. 2011a. New Brontotheriidae (Mammalia, Perissodactyla) from the early and middle Eocene of Pakistan, with implications for mammalian paleobiogeography. Journal of Paleontology 85:665–677.
  • Missiaen, P., F. Quesnel, C. Dupuis, J.-Y. Storme, and T. Smith. 2013. The earliest Eocene mammal fauna of the Erquelinnes Sand Member near the French-Belgian border. Geologica Belgica 16:262–273.
  • Missiaen, P., K. D. Rose, R. S. Rana, K. Kumar, and T. Smith. 2011b. Revision of Indobune and Cambaytherium from the early Eocene of Vastan (India), and their affinities with anthracobunid and perissodactyl mammals. Journal of Vertebrate Paleontology 31 (Program and Abstracts):159.
  • Mohan, M. 1995. Cambay basin—a promise of oil and gas potential. Journal of the Palaeontological Society of India 40:41–47.
  • Mones, A. 1989. Nomen dubium versus nomen vanum. Journal of Vertebrate Paleontology 9:232–234.
  • Muizon, C. de, and R. L. Cifelli. 2000. The “condylarths” (archaic Ungulata, Mammalia) from the early Paleocene of Tiupampa (Bolivia): implications on the origin of South American ungulates. Geodiversitas 22:47–150.
  • Muller, J. 1935. The orbitotemporal region of the skull of the Mammalia. Archives Néerlandaises de Zoologie 1:118–259.
  • Murie, J. 1872. On the Malayan tapir, Rhinochoerus sumatranus (Gray). Journal of Anatomy and Physiology 6:131–169.
  • Murphy, W. J., E. Eizirik, W. E. Johnson, Y. P. Zhang, O. A. Ryder, and S. J. O’Brien. 2001. Molecular phylogenetics and the origins of placental mammals. Nature 409:614–618.
  • Nagori, M. L., S. C. Khosla, and S. R. Jakhar. 2013. Middle Eocene Ostracoda from the Tadkeshwar Lignite Mine, Cambay Basin, Gujarat. Journal of the Geological Society of India 81:514–520.
  • Nolf, D., R. S. Rana, and H. Singh. 2006. Fish otoliths from the Ypresian (early Eocene) of Vastan, Gujarat, India. Bulletin de l’Institut Royal des Sciences Naturelles de Belgique, Sciences de la Terre, Belgium 76:105–118.
  • Novacek, M. J. 1986. The skull of leptictid insectivorans and the higher-level classification of eutherian mammals. Bulletin of the American Museum of Natural History 183:1–112.
  • Novacek, M. J., and A. R. Wyss. 1986. Higher-level relationships of the Recent eutherian orders: morphological evidence. Cladistics 2:257–287.
  • O’Leary, M. A., and K. D. Rose. 1995. Postcranial skeleton of the early Eocene mesonychid Pachyaena (Mammalia: Mesonychia). Journal of Vertebrate Paleontology 15:401–430.
  • O’Leary, M. A., J. I. Bloch, J. J. Flynn, T. J. Gaudin, A. Giallombardo, N. P. Giannini, S. L. Goldberg, B. P. Kraatz, Z.-X. Luo, J. Meng, X. Ni, M. J. Novacek, F. A. Perini, Z. S. Randall, G. W. Rougier, E. J. Sargis, M. T. Silcox, N. B. Simmons, M. Spaulding, P. M. Velazco, M. Weksler, J. R. Wible, and A. L. Cirranello. 2013. The placental mammal ancestor and the post-K-Pg radiation of placentals. Science 339:662–667.
  • Osborn, H. F. 1898. Remounted skeleton of Phenacodus primaevus. Comparison with Euprotogonia. Bulletin of the American Museum of Natural History 10:159–164.
  • Otts, C. 1991. Postural and locomotor capabilities in the phenacodontid condylarths (Mammalia). Ph.D. dissertation, University of Arizona, Tucson, Arizona, 566 pp.
  • Owen, R. 1848. Description of teeth and portions of jaws of two extinct Anthracotherioid quadrupeds (Hyopotamus vectianus and Hyop. bovinus) discovered by the Marchioness of Hastings in the Eocene deposits on the NW coast of the Isle of Wight: with an attempt to develope Cuvier’s idea of the Classification of Pachyderms by the number of their toes. Quarterly Journal of the Geological Society of London 4:103–141.
  • Pandey, J., N. P. Singh, B. R. Krishna, D. D. Sharma, A. K. Parakh, and S. S. Nath. 1993. Lithostratigraphy of Cambay Basin. Document III, Keshav Dev Malviya Institute of Petroleum Exploration Publication, Oil and Natural Gas Commission, Dehradun 1:1–166.
  • Paul, S., J. Sharma, B. D. Singh, P. K. Saraswati, and S. Dutta. 2015. Early Eocene equatorial vegetation and depositional environment: biomarker and palynological evidences from a lignite-bearing sequence of Cambay Basin, western India. International Journal of Coal Geology 149:77–92.
  • Pilgrim, G. E. 1940. Middle Eocene mammals from northwest Pakistan. Proceedings of the Zoological Society of London, Series B 110:127–152.
  • Prasad, G. V. R., and S. Bajpai. 2008. Agamid lizards from the early Eocene of western India: oldest Cenozoic lizards from South Asia. Palaeontologia Electronica 11.1.4A.
  • Prasad, V., A. Farooqui, S. K. M. Tripathi, and B. Thakur. 2009. Evidence of Late Palaeocene-Early Eocene equatorial rain forest refugia in southern Western Ghats, India. Journal of Biosciences 34:777–797.
  • Prasad, V., I. B. Singh, S. Bajpai, R. Garg, B. Thakur, A. Singh, N. Saravanan, and V. V. Kapur. 2013. Palynofacies and sedimentology-based high-resolution sequence stratigraphy of the lignite-bearing muddy coastal deposits (early Eocene) in the Vastan Lignite Mine, Gulf of Cambay, India. Facies 59:737–761.
  • Prothero, D. R., E. M. Manning, and M. Fischer. 1988. The phylogeny of the ungulates; pp. 201–234 in M. J. Benton (ed.), The Phylogeny and Classification of the Tetrapods. 2: Mammals. Clarendon Press, Oxford, U.K.
  • Punekar, J., and P. K. Saraswati. 2010. Age of the Vastan lignite in context of some oldest Cenozoic fossil mammals from India. Journal of the Geological Society of India 76:63–68.
  • Radinsky, L. B. 1963a. Origin and early evolution of North American Tapiroidea. Bulletin Yale Peabody Museum 17:1–106.
  • Radinsky, L. B. 1963b. The perissodactyl hallux. American Museum Novitates 2145:1–8.
  • Radinsky, L. B. 1965a. Evolution of the tapiroid skeleton from Heptodon to Tapirus. Bulletin of the Museum of Comparative Zoology 134:69–106.
  • Radinsky, L. B. 1965b. Early Tertiary Tapiroidea of Asia. Bulletin of the American Museum of Natural History 129:181–264.
  • Radinsky, L. B. 1966. The adaptive radiation of the phenacodontid condylarths and the origin of the Perissodactyla. Evolution 20:408–417.
  • Radinsky, L. B. 1969. The early evolution of the Perissodactyla. Evolution 23:308–328.
  • Rage, J.-C., A. Folie, R. S. Rana, H. Singh, K. D. Rose, and T. Smith. 2008. A diverse snake fauna from the early Eocene of Vastan Lignite Mine, Gujarat, India. Acta Palaeontologica Polonica 53:391–403.
  • Rana, R. S., K. Kumar, and H. Singh. 2004. Vertebrate fauna from the subsurface Cambay Shale (Lower Eocene), Vastan Lignite Mine, Gujarat, India. Current Science 87:1726–1733.
  • Rana, R. S., M. Augé, A. Folie, K. D. Rose, K. Kumar, L. Singh, A. Sahni, and T. Smith. 2013. High diversity of acrodontan lizards in the Early Eocene Vastan Lignite Mine of India. Geologica Belgica 16:290–301.
  • Rana, R. S., K. Kumar, G. Escarguel, A. Sahni, K. D. Rose, T. Smith, H. Singh, and L. Singh. 2008. An ailuravine rodent from the lower Eocene Cambay Formation at Vastan, western India, and its palaeobiogeographic implications. Acta Palaeontologica Polonica 53:1–14.
  • Rana, R. S., K. Kumar, S. P. Zack, F. Solé, K. D. Rose, P. Missiaen, L. Singh, A. Sahni, and T. Smith. 2015. Craniodental and postcranial morphology of Indohyaenodon raoi from the early Eocene of India, and its implications for ecology, phylogeny, and biogeography of hyaenodontid mammals. Journal of Vertebrate Paleontology. doi: 10.1080/02724634.2015.965308.
  • Rangarajan, S. 2008. A case for revisiting the evolutionary history of pericratonic Cambay basin, India; pp. 123–129 in Seventh International Conference and Exposition on Petroleum Geophysics, Hyderabad, 2008. Society of Petroleum Geologists of India.
  • Rao, K. L. N. 1969. Lithostratigraphy of the Paleogene succession of Southern Cambay Basin. Bulletin Oil and National Gas Commission (ONGC), Dehradun, India 6:24–37.
  • Rao, M. R., A. Sahni, R. S. Rana, and P. Verma. 2013. Palynostratigraphy and depositional environment of Vastan Lignite Mine (Early Eocene), Gujarat, western India. Journal Earth System Science 122:289–307.
  • Rasmussen, D. T., M. Gagnon, and E. L. Simons. 1990. Taxeopody in the carpus and tarsus of Oligocene Pliohyracidae (Mammalia: Hyracoidea) and the phyletic position of hyraxes. Proceedings of the National Academy of Sciences of the United States of America 87:4688–4691.
  • Ray, C. E., D. P. Domning, and M. C. McKenna. 1994. A new specimen of Behemotops proteus (Order Desmostylia) from the marine Oligocene of Washington. Proceedings of the San Diego Museum of Natural History 29:205–222.
  • Richards, M. W., and A. G. Watson. 1991. Development and variation of the lateral vertebral foramen of the atlas in dogs. Anatomia, Histologia, Embryologia 20:363–368.
  • Rose, K. D. 1981. The Clarkforkian Land-Mammal Age and mammalian faunal composition across the Paleocene-Eocene boundary. University of Michigan Papers on Paleontology 26:1–197.
  • Rose, K. D. 1990. Postcranial skeletal remains and adaptations in early Eocene mammals from the Willwood Formation, Bighorn Basin, Wyoming; pp. 107–133 in T. M. Bown and K. D. Rose (eds.), Dawn of the Age of Mammals in the Northern Part of the Rocky Mountain Interior. Geological Society of America Special Paper 243. Boulder, Colorado.
  • Rose, K. D. 1996. Skeleton of early Eocene Homogalax and the origin of Perissodactyla. Palaeovertebrata 25:243–260.
  • Rose, K. D., R. H. Dunn, and L. Grande. 2014a. A new pantolestid skeleton from the early Eocene Fossil Butte Member, Green River Formation (Wyoming), and skeletal ontogeny in Pantolestidae (Mammalia, Pantolesta). Journal of Vertebrate Paleontology 34:932–940.
  • Rose, K. D., L. T. Holbrook, and W. P. Luckett. 2018a. Deciduous premolars of Eocene Equidae and their phylogenetic significance. Historical Biology 30:89–118.
  • Rose, K. D., R. S. Rana, A. Sahni, and T. Smith. 2007. A new adapoid primate from the early Eocene of India. Contributions from the Museum of Paleontology, The University of Michigan 31:379–385.
  • Rose, K. D., K. Kumar, R. S. Rana, A. Sahni, and T. Smith. 2013. New hypsodont tillodont (Mammalia: Tillodontia) from the early Eocene of India. Journal of Paleontology 87:842–853.
  • Rose, K. D., A. E. Chew, R. H. Dunn, M. J. Kraus, H. C. Fricke, and S. P. Zack. 2012. Earliest Eocene mammalian fauna from the Paleocene-Eocene Thermal Maximum at Sand Creek Divide, southern Bighorn Basin, Wyoming. University of Michigan Papers on Paleontology 36:1–122.
  • Rose, K. D., R. S. Rana, A. Sahni, K. Kumar, L. Singh, and T. Smith. 2009a. First tillodont from India: additional evidence for an early Eocene faunal connection between Europe and India? Acta Palaeontologica Polonica 54:351–355.
  • Rose, K. D., V. B. DeLeon, P. Missiaen, R. S. Rana, A. Sahni, L. Singh, and T. Smith. 2008. Early Eocene lagomorph (Mammalia) from western India and the early diversification of Lagomorpha. Proceedings of the Royal Society London B: Biological Sciences 275:1203–1208.
  • Rose, K. D., R. H. Dunn, K. Kumar, J. M. G. Perry, K. A. Prufrock, R. S. Rana, and T. Smith. 2018b. New fossils from Tadkeshwar Mine (Gujarat, India) increase primate diversity from the early Eocene Cambay Shale. Journal of Human Evolution 122:93–107.
  • Rose, K. D., R. S. Rana, A. Sahni, K. Kumar, P. Missiaen, L. Singh, and T. Smith. 2009b. Early Eocene primates from Gujarat, India. Journal of Human Evolution 56:366–404.
  • Rose, K. D., T. Smith, R. S. Rana, A. Sahni, H. Singh, P. Missiaen, and A. Folie. 2006. Early Eocene (Ypresian) continental vertebrate assemblage from India, with description of a new anthracobunid (Mammalia, Tethytheria). Journal of Vertebrate Paleontology 26:219–225.
  • Rose, K. D., L. T. Holbrook, R. S. Rana, K. Kumar, K. E. Jones, H. E. Ahrens, P. Missiaen, A. Sahni, and T. Smith. 2014b. Early Eocene fossils suggest that the mammalian order Perissodactyla originated in India. Nature Communications 5:5570.
  • Russell, D. E. 1964. Les mammifères Paléocènes d’Europe. Mémoires du Muséum National d’Histoire Naturelle Série C, Sciences de la Terre, tome XIII:1–324.
  • Rust, J., H. Singh, R. S. Rana, T. McCann, L. Singh, K. Anderson, N. Sarkar, P. C. Nascimbene, F. Stebner, J. C. Thomas, M. S. Kraemer, C. J. Williams, M. S. Engel, A. Sahni, and D. Grimaldi. 2010. Biogeographic and evolutionary implications of a diverse paleobiota in amber from the early Eocene of India. Proceedings of the National Academy of Sciences of the United States of America 107:18360–18365.
  • Sahni, A., and S. K. Khare. 1973. Additional Eocene mammals from the Subathu Formation of Jammu and Kashmir. Journal of the Palaeontological Society of India 17:31–49.
  • Sahni, A., and K. Kumar. 1980. Lower Eocene Sirenia, Ishatherium subathuensis, gen. et. sp. nov. from the type area, Subathu Formation, Subathu, Simla Himalayas, H.P. Journal of the Palaeontological Society of India 23–24:132–135.
  • Sahni, A., P. K. Saraswati, R. S. Rana, K. Kumar, H. Singh, H. Alimohammadian, N. Sahni, K. D. Rose, L. Singh, and T. Smith. 2006. Temporal constraints and depositional palaeoenvironments of the Vastan lignite sequence, Gujarat: analogy for the Cambay Shale hydrocarbon source rock. Indian Journal of Petroleum Geology 15:1–20.
  • Samanta, A., A. Sarkar, M. K. Bera, J. Rai, and S. S. Rathore. 2013. Late Paleocene-early Eocene carbon isotope stratigraphy from a near-terrestrial tropical section and antiquity of Indian mammals. Journal of Earth System Science 122:163–171.
  • Scotese, C. R. 2006. Paleomap project. Available at www.scotese.com. Accessed November 15, 2018.
  • Scott, K. M. 1983. Prediction of body weight of fossil Artiodactyla. Zoological Journal of the Linnean Society 77:199–215.
  • Scott, K. M. 1990. Postcranial dimensions of ungulates as predictors of body mass; pp. 301–336 in J. Damuth and B. J. MacFadden (eds.), Body Size in Mammalian Paleobiology: Estimation and Biological Implications. Cambridge University Press, Cambridge, U.K.
  • Scott, W. B. 1888. On some new and little known creodonts. Journal of the Academy of Natural Sciences of Philadelphia 9:155–185.
  • Seiffert, E. R. 2007. A new estimate of afrotherian phylogeny based on simultaneous analysis of genomic, morphological, and fossil evidence. BMC Evolutionary Biology 7:224.
  • Serra-Kiel, J., L. Hottinger, E. Caus, K. Drobne, C. Ferrandez, A. K. Jauhri, G. Less, R. Pavlovec, J. Pignatti, J. M. Samsó, H. Schaub, E. Sirel, A. Strougo, Y. Tambareau, J. Tosquella, and E. Y. Zakrevskaya. 1998. Larger foraminiferal biostratigraphy of the Tethyan Paleocene and Eocene. Bulletin de la Société Géologique de France 169:281–299.
  • Shelley, S. L., T. E. Williamson, and S. L. Brusatte. 2018. The osteology of Periptychus carinidens: a robust, ungulate-like placental mammal (Mammalia: Periptychidae) from the Paleocene of North America. PLoS ONE 13:e0200132.
  • Silva, M., and J. A. Downing. 1995. CRC Handbook of Mammalian Body Masses. CRC Press, Boca Raton, Florida, 359 pp.
  • Simpson, G. G. 1940. Mammals and land bridges. Journal of the Washington Academy of Sciences 30:137–163.
  • Simpson, G. G. 1945. The principles of classification and a classification of mammals. Bulletin of the American Museum of Natural History 85:1–350.
  • Simpson, G. G. 1947. Holarctic mammalian faunas and continental relationships during the Cenozoic. Bulletin of the Geological Society of America 58:613–688.
  • Simpson, G. G., A. Roe, and R. C. Lewontin. 1960. Quantitative Zoology. Harcourt, Brace and Company, New York, 440 pp.
  • Singh, H., M. Prasad, K. Kumar, and S. K. Singh. 2011. Paleobotanical remains from the Paleocene–Lower Eocene Vagadkhol Formation, western India and their climatic and phytogeographic implications. Palaeoworld 20:332–356.
  • Singh, H., M. Prasad, K. Kumar, and S. K. Singh. 2015. Early Eocene macroflora and associated palynofossils from the Cambay Shale Formation, Western India: phytogeographic and palaeoclimatic implications. Palaeoworld 24:293–323.
  • Singh, H., B. Samant, T. Adatte, and H. Khozyem. 2014. Diverse palynoflora from amber and associated sediments of Tarkeshwar lignite mine, Surat district, Gujarat, India. Current Science 106:930–932.
  • Singh, H., M. Prasad, K. Kumar, R. S. Rana, and S. K. Singh. 2010. Fossil fruits from Early Eocene Vastan Lignite, Gujarat, India: taphonomic and phytogeographic implications. Current Science 98:1625–1632.
  • Slijper, E. J. 1946. Comparative biologic-anatomical investigations on the vertebral column and spinal musculature of mammals. Verhandelingen der Koninklijke Nederlandse Akademie van Wetenschappen, Afdeeling Natuurkunde, Tweede Reeks 42:1–128.
  • Sloan, R. E. 1969. Cretaceous and Paleocene terrestrial communities of western North America. Proceedings of the North American Paleontological Convention E:427–453.
  • Smith, T., K. D. Rose, and P. D. Gingerich. 2006. Rapid Asia-Europe-North America geographic dispersal of earliest Eocene primate Teilhardina during the Paleocene-Eocene Thermal Maximum. Proceedings of the National Academy of Sciences of the United States of America 103:11223–11227.
  • Smith, T., J. Habersetzer, N. B. Simmons, and G. F. Gunnell. 2012. Systematics and paleobiogeography of early bats; pp. 23–66 in G. F. Gunnell and N. B. Simmons (eds.), Evolutionary History of Bats: Fossils, Molecules and Morphology. Cambridge University Press, Cambridge, U.K.
  • Smith, T., R. S. Rana, P. Missiaen, K. D. Rose, A. Sahni, H. Singh, and L. Singh. 2007. High bat (Chiroptera) diversity in the Early Eocene of India. Naturwissenschaften 94:1003–1009.
  • Smith, T., F. Solé, P. Missiaen, R. S. Rana, K. Kumar, A. Sahni, and K. D. Rose. 2015. First early Eocene tapiroid from India and its implication for the paleobiogeographic origin of perissodactyls. Palaeovertebrata 39(2):e5.
  • Smith, T., K. Kumar, R. S. Rana, A. Folie, F. Solé, C. Noiret, T. Steeman, A. Sahni, and K. D. Rose. 2016. New early Eocene vertebrate assemblage from western India reveals a mixed fauna of European and Gondwanan affinities. Geoscience Frontiers 7:969–1001.
  • Solé, F., E. De Bast, H. Legendre, R. S. Rana, K. Kumar, K. D. Rose, and T. Smith. In press. New specimens of Frugivastodon (Mammalia: Apatotheria) from the early Eocene of India confirm its apatemyid status and elucidate dispersal of Apatemyidae; in G. V. R. Prasad and R. Patnaik (eds.), Biological Consequences of Plate Tectonics: New Perspectives on Post-Gondwanaland Break-up–A Tribute to Ashok Sahni, in Vertebrate Paleobiology and Paleoanthropology series. Springer.
  • Solounias, N., M. Danowitz, E. Stachtiaris, A. Khurana. M. Araim, M. Sayegh, and J. Natale. 2018. The evolution and anatomy of the horse manus with an emphasis on digit reduction. Royal Society Open Science 5:e171782.
  • Sondaar, P. Y. 1968. The osteology of the manus of fossil and recent Equidae with special reference to phylogeny and function. Verhandelingen der Koninklijke Nederlandse Akademie van Wetenschappen, Afdeeling Natuurkunde, Eerste Reeks 25:1–76.
  • Sperber, G. H. 2017. Dental wear: attrition, erosion, and abrasion—a palaeo-odontological approach. Dentistry Journal 5(2):e19.
  • Stefen, C. 1997. Differentiations in Hunter-Schreger bands of carnivores; pp. 123–136 in W. von Koenigswald and P. M. Sander (eds.), Tooth Enamel Microstructure. A. A. Balkema, Rotterdam, The Netherlands.
  • Sudhakar, R., and D. N. Basu. 1973. A reappraisal of the Paleogene stratigraphy of southern Cambay Basin. Bulletin of the Oil and Natural Gas Commission 10:55–76.
  • Swofford, D. L. 2003. PAUP*: Phylogenetic Analysis Using Parsimony (*and Other Methods), version 4.0b10a. Sinauer Associates, Sunderland, Massachusetts.
  • Szalay, F. S. 1969. Mixodectidae, Microsyopidae, and the insectivore-primate transition. Bulletin of the American Museum of Natural History 140:193–330.
  • Tassy, P. 1981. Le crâne de Moeritherium (Proboscidea, Mammalia) de l’Eocène de Dor el Talha (Libye) et le problème de la classification phylogénétique du genre dans les Tethytheria McKenna, 1975. Bulletin du Muséum National d’Histoire Naturelle, Paris 4:87–147.
  • Tassy, P., and J. Shoshani. 1988. The Tethytheria: elephants and their relatives; pp. 283–315 in M. J. Benton (ed.), The Phylogeny and Classification of the Tetrapods, Volume 2. Clarendon Press, Oxford, U.K.
  • Thewissen, J. G. M. 1990. Evolution of Paleocene and Eocene Phenacodontidae (Mammalia, Condylarthra). University of Michigan Papers on Paleontology 29:1–107.
  • Thewissen, J. G. M., and D. P. Domning. 1992. The role of phenacodontids in the origin of the modern orders of ungulate mammals. Journal of Vertebrate Paleontology 12:494–504.
  • Ting, S.-Y. 1993. A preliminary report on an early Eocene mammalian fauna from Hengdong, Hunan Province, China. Kaupia 3:201–207.
  • Tong, Y.-S., J.-W. Wang, and J. Meng. 2004. Olbitherium millenariusum, a new perissodactyl-like archaic ungulate (Mammalia) from the early Eocene Wutu Formation, Shandong. Vertebrata PalAsiatica 42:27–38.
  • Tsubamoto, T., S. T. Tun, N. Egi, M. Takai, N. Shigehara, A. N. Soe, A. K. Aung, and T. Thein. 2003. Reevaluation of some ungulate mammals from the Eocene Pondaung Formation, Myanmar. Paleontological Research 7:219–243.
  • Ungar, P. S. 2010. Mammalian Teeth: Origin, Evolution, and Diversity. The Johns Hopkins University Press, Baltimore, Maryland, 304 pp.
  • Van Valen, L. 1966. Deltatheridia, a new order of mammals. Bulletin of the American Museum of Natural History 132:1–126.
  • Welker, F., M. Collins, J. Thomas, M. Wadsley, S. Brace, E. Cappellini, S. Turvey, M. Reguero, J. Gelfo, A. Kramarz, J. Burger, J. Thomas-Oates, D. Ashford, P. Ashton, K. Rowsell, D. Porter, B. Kessler, R. Fischer, C. Baessmann, S. Kaspar, J. Olsen, P. Kiley, J. Elliott, C. Kelstrup, V. Mullin, M. Hofreiter, E. Willerslev, J.-J. Hublin, L. Orlando, I. Barnes, and R. MacPhee. 2015. Ancient proteins resolve the evolutionary history of Darwin’s South American ungulates. Nature 522:81–84.
  • Wells, N. A., and P. D. Gingerich. 1983. Review of Eocene Anthracobunidae (Mammalia, Proboscidea) with a new genus and species, Jozaria palustris, from the Kuldana Formation of Kohat (Pakistan). Contributions from the Museum of Paleontology, The University of Michigan 26:117–139.
  • West, R. M. 1971. Deciduous dentition of the early Tertiary Phenacodontidae (Condylarthra, Mammalia). American Museum Novitates 2461:1–37.
  • West, R. M. 1980. Middle Eocene large mammal assemblage with Tethyan affinities, Ganda Kas region, Pakistan. Journal of Paleontology 54:508–533.
  • West, R. M. 1983. South Asian Middle Eocene moeritheres (Mammalia: Tethytheria). Annals of Carnegie Museum 52:359–373.
  • Westbury, M., S. Baleka, A. Barlow, S. Hartmann, J. Paijmans, A. Kramarz, A. Forasiepi, M. Bond, J. Gelfo, M. Reguero, P. López-Mendoza, M. Taglioretti, F. Scaglia, A. Rinderknecht, W. Jones, F. Mena, G. Billet, C. de Muizon, J. Aguilar, R. MacPhee, and M. Hofreiter. 2015. A mitogenomic timetree for Darwin’s enigmatic South American mammal Macrauchenia patachonica. Nature Communications 8:15951.
  • Wible, J. R. 1987. The eutherian stapedial artery: character analysis and implications for superordinal relationships. Zoological Journal of the Linnean Society 91:107–135.
  • Wible, J. R., G. W. Rougier, M. J. Novacek, and R. J. Asher. 2009. The eutherian mammal Maelestes gobiensis from the Late Cretaceous of Mongolia and the phylogeny of Cretaceous Eutheria. Bulletin of the American Museum of Natural History 327:1–123.
  • Williams, P. L. (ed.). 1995. Gray’s Anatomy, thirty-eighth edition. Churchill Livingstone, Edinburgh, U.K., 2092 pp.
  • Williamson, T. E., and S. G. Lucas. 1992. Meniscotherium (Mammalia, “Condylarthra”) from the Paleocene-Eocene of western North America. Bulletin of the New Mexico Museum of Natural History and Science 1:1–75.
  • Wood, A. R., R. M. Bebej, C. L. Manz, D. L. Begun, and P. D. Gingerich. 2011. Postcranial functional morphology of Hyracotherium (Equidae, Perissodactyla) and locomotion in the earliest horses. Journal of Mammalian Evolution 18:1–32.
  • Wortman, J. L. 1894. Osteology of Patriofelis, a middle Eocene creodont. Bulletin of the American Museum of Natural History 6:129–165.
  • Wortman, J. L. 1896. Species of Hyracotherium and allied perissodactyls from the Wasatch and Wind River beds of North America. Bulletin of the American Museum of Natural History 8:81–111.
  • Wyss, A. R., and J. J. Flynn. 1993. A phylogenetic analysis and definition of the Carnivora; pp. 32–52 in F. S. Szalay, M. J. Novacek, and M. C. McKenna (eds.), Mammal Phylogeny: Placentals. Springer, New York.
  • Wyss, A., and J. Meng. 1996. Application of phylogenetic taxonomy to poorly resolved crown clades: a stem-modified node-based definition of Rodentia. Systematic Biology 45:559–568.
  • Youlatos, D. 2003. Osteological correlates of tail prehensility in carnivorans. Journal of Zoology, London 259:423–430.
  • Zack, S. P., K. D. Rose, L. T. Holbrook, K. Kumar, R. S. Rana, and T. Smith. 2019. An enigmatic new ungulate-like mammal from the early Eocene of India. Papers in Palaeontology. doi: 10.1002/spp2.1288.
  • Zhang, Y.-P. 1978. Two new genera of condylarthran phenacolophids from the Paleocene of Nanxiong Basin, Guangdong. Vertebrata PalAsiatica 16:267–274.
  • Zhou, X., W. J. Sanders, and P. D. Gingerich. 1992. Functional and behavioral implications of vertebral structure in Pachyaena ossifraga (Mammalia, Mesonychia). Contributions from the Museum of Paleontology, The University of Michigan 28:289–319.
  • Zhu, D. C., Q. Wang, Z. D. Zhao, S. L. Chung, P. A. Cawood, Y. Niu, S. A. Liu, F. Y. Wu, and X. X. Mo. 2015. Magmatic record of India-Asia collision. Scientific Reports 5:14289.

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