Anatomy and phylogenetic affinities of a new silesaurid assemblage from the Carnian beds of south Brazil

LITERATURE CITED

• Agnolin, F.L. & Ezcurra, M.D. (2019). The validity of Lagosuchus talampayensis Romer, 1971 (Archosauria, Dinosauriformes), from the Late Triassic of Argentina. Breviora, 565(1), 1–21. https://doi.org/10.3099/0006-9698-565.1.1.
   Google Scholar
• Agnolín, F.L., & Rozadilla, S. (2018). Phylogenetic reassessment of Pisanosaurus mertii Casamiquela, 1967, a basal dinosauriform from the Late Triassic of Argentina. Journal of Systematic Palaeontology, 16(10), 853–879. https://doi.org/10.1080/14772019.2017.1352623.
   Web of Science ®Google Scholar
• Amprino, R. (1947). La structure du tissu osseux envisagée comme expression de différences dans la vitesse de l’accroissement. Archives de biologie, 58(4), 317–330.
   Google Scholar
• Andreis, R., Bossi, G.E., & Montardo, D.K. (1980). O Grupo Rosário do Sul (Triássico) no Rio Grande do Sul. 659–673. In Anais do XXXI Congresso Brasileiro de Geologia, Camboriú, Santa Catarina. Sociedade Brasileira de Geologia, Sao Paulo, Brazil.
   Google Scholar
• Baron, M., Norman, D. B. & Barrett, P. (2017a). A new hypothesis of dinosaur relationships and early dinosaur evolution. Nature 543, 501–506. https://doi.org/10.1038/nature21700.
   PubMed Web of Science ®Google Scholar
• Baron, M. G., Norman, D. B., & Barrett, P. M. (2017b). Postcranial anatomy of Lesothosaurus diagnosticus (Dinosauria: Ornithischia) from the Lower Jurassic of southern Africa: implications for basal ornithischian taxonomy and systematics. Zoological Journal of the Linnean Society, 179(1), 125–168. https://doi.org/10.1111/zoj.12434.
   Web of Science ®Google Scholar
• Benton, M. J., Forth J., & Langer, M. C. (2014). Models for the rise of the dinosaurs. Current Biology, 24(2), R87–R95. https://doi.org/10.1016/j.cub.2013.11.063.
   PubMed Web of Science ®Google Scholar
• Benton, M.J., & Walker, A. D. (2011). Saltopus, a dinosauriform from the Upper Triassic of Scotland. Earth and Environmental Science Transactions of the Royal Society of Edinburgh, 101(3-4), 285–299. https://doi.org/10.1017/S1755691011020081.
   Google Scholar
• Bittencourt, J. D. S., Arcucci A. B., Marsicano C. A., & Langer, M. C. (2015). Osteology of the Middle Triassic archosaur Lewisuchus admixtus Romer (Chañares Formation, Argentina), its inclusivity, and relationships amongst early dinosauromorphs. Journal of Systematic Palaeontology, 13(3), 189–219. https://doi.org/10.1080/14772019.2013.878758.
   Web of Science ®Google Scholar
• Bittencourt, J. D. S., & Kellner, A, W. A. (2009). The anatomy and phylogenetic position of the Triassic dinosaur Staurikosaurus pricei Colbert, 1970. Zootaxa, 2079(1), 1–56. https://doi.org/10.11646/zootaxa.2079.1.1.
   Google Scholar
• Brusatte, S. L., Benton, M. J., Ruta, M., & Lloyd, G. T. (2008). Superiority, competition, and opportunism in the evolutionary radiation of dinosaurs. Science, 321(5895), 1485–1488. https://doi.org/10.1126/science.1161833.
   PubMed Web of Science ®Google Scholar
• Brusatte, S. L., Nesbitt, S. J., Irmis, R. B., Butler, R, J., Benton, M. J., & Norell, M. A. (2010). The origin and early radiation of dinosaurs. Earth-Science Reviews, 101(1-2), 68–100. https://doi.org/10.1016/j.earscirev.2010.04.001.
   Web of Science ®Google Scholar
• Butler, R. J. (2010). The anatomy of the basal ornithischian dinosaur Eocursor parvus from the lower Elliot Formation (Late Triassic) of South Africa. Zoological Journal of the Linnean Society, 160(4), 648–684. https://doi.org/10.1111/j.1096-3642.2009.00631.x.
   Web of Science ®Google Scholar
• Butler, R. J., Galton, P. M., Porro, L. B., Chiappe, L. M., Henderson, D. M., & Erickson, G. M. (2010). Lower limits of ornithischian dinosaur body size inferred from a new Upper Jurassic heterodontosaurid from North America. Proceedings of the Royal Society B: Biological Sciences, 277(1680), 375–381. https://doi.org/10.1098/rspb.2009.1494.
   PubMed Web of Science ®Google Scholar
• Butler, R. J., Porro, L. B., & Norman, D. B. (2008). A juvenile skull of the primitive ornithischian dinosaur Heterodontosaurus tucki from the ‘Stormberg’ of southern Africa. Journal of Vertebrate Paleontology, 28, 702–711. https://doi.org/10.1671/0272.
   Web of Science ®Google Scholar
• Cabreira, S. F., Kellner, A. W. A., Dias-da-Silva, S., da Silva, L. R., Bronzati, M., Marsola, J. C., Müller, R. T., Bittencourt, J. S., Batista, B. J., Raugust, T., Carrilho, R., Brodt, A., & Langer, M. C. (2016). A unique Late Triassic dinosauromorph assemblage reveals dinosaur ancestral anatomy and diet. Current Biology, 26(22), 3090–3095. https://doi.org/10.1016/j.cub.2016.09.040.
   PubMed Web of Science ®Google Scholar
• Casamiquela, R. M. (1967). Un nuevo dinosaurio ornitisquio triásico (Pisanosaurus mertii; Ornithopoda) de la Formación Ischigualasto, Argentina. Ameghiniana, 5(2), 47–64.
   Google Scholar
• Cerda, I. A., Pol, D., & Chinsamy, A. (2014). Osteohistological insight into the early stages of growth in Mussaurus patagonicus (Dinosauria, Sauropodomorpha). Historical Biology, 26(1), 110–121. https://doi.org/10.1080/08912963.2012.763119.
   Web of Science ®Google Scholar
• Chatterjee, S. (1984). A new ornithischian dinosaur from the Triassic of North America. Naturwissenschaften, 71(12), 630–631. DOI10.1007/BF00377897.
   Google Scholar
• Chinsamy-Turan, A. (1990). Physiological implications of the bone histology of Syntarsus rhodesiensis (Saurischia: Theropoda). Palaeontologia africana, 27, 77–82.
   Google Scholar
• Chinsamy-Turan, A. (2008). The microstructure of dinosaur bone: deciphering biology with fine-scale techniques. Zoological Journal, 154(3), 631–632. https://doi.org/10.1111/j.1096-3642.2008.00476.x.
   Google Scholar
• Coddington, J. A., & Scharff, N. (1994). Problems with zero-length branches. Cladistic 10, 415–423.
   Web of Science ®Google Scholar
• Da Rosa, Á. A. S. (2004). Sítios fossilíferos de Santa Maria, RS. Ciência & Natura 26, 5–90.
   Google Scholar
• Da Rosa, Á. A. S. (2005). Paleoalterações em depósitos sedimentares deplanícies aluviais do Triássico Médio a Superior do sul do Brasil:caracterização, análise estratigráfica e preservação fossilífera. UNISINOS, São Leopoldo. 2005.
   Google Scholar
• Da Rosa, Á. A. S. (2015). Geological context of the dinosauriform-bearing outcrops from the Triassic of Southern Brazil. Journal of South American Earth Sciences 61, 108–119. https://doi.org/10.1016/j.jsames.2014.10.008.
   Web of Science ®Google Scholar
• Desojo, J. B., Fiorelli, L.E., Ezcurra, M. D., Martinelli, A.G., Ramezani, J., Da Rosa, Á. A., von Baczko, M. B, Trotteyn, M. J., Montefeltro, F. C., Ezpeleta, M., & Langer, M. C. (2020). The Late Triassic Ischigualasto Formation at Cerro Las Lajas (La Rioja, Argentina): fossil tetrapods, high-resolution chronostratigraphy, and faunal correlations. Scientific Reports, 10(1), 1–34. https://doi.org/10.1038/s41598-020-67854-1.
   PubMed Web of Science ®Google Scholar
• Dzik, J. (2003). A beaked herbivorous archosaur with dinosaur affinities from the early Late Triassic of Poland. Journal of Vertebrate Paleontology 23(3), 556–574. https://doi.org/10.1671/A1097.
   Web of Science ®Google Scholar
• Eltink, E., Da-Rosa, Á. A. S., & Dias-da-Silva, S. (2017). A capitosauroid from the Lower Triassic of South America (Sanga do Cabral Supersequence: Paraná Basin), its phylogenetic relationships and biostratigraphic implications. Historical Biology, 29(7), 863–874. https://doi.org/10.1080/08912963.2016.1255736.
   Web of Science ®Google Scholar
• Ezcurra, M. D. (2006). A review of the systematic position of the dinosauriform archosaur Eucoelophysis baldwini Sullivan & Lucas, 1999 from the Upper Triassic of New Mexico, USA. Geodiversitas, 28(4), 649–684.
   Web of Science ®Google Scholar
• Ezcurra, M. D., Nesbitt, S. J., Fiorelli, L. E., & Desojo, J. B. (2019). New specimen sheds light on the anatomy and taxonomy of the early Late Triassic dinosauriforms from the Chañares Formation, NW Argentina. The Anatomical Record, 303(5), 1393–1438. https://doi.org/10.1002/ar.24243.
   Google Scholar
• Ferigolo, J., & Langer, M. C. (2007). A Late Triassic dinosauriform from south Brazil and the origin of the ornithischian predentary bone. Historical Biology, 19(1), 23–33. https://doi.org/10.1080/08912960600845767.
   Google Scholar
• Fostowicz-Frelik, Ł., & Sulej, T. (2010). Bone histology of Silesaurus opolensis Dzik, 2003 from the Late Triassic of Poland. Lethaia, 43(2), 137–148. https://doi.org/10.1111/j.1502-3931.2009.00179.x.
   Web of Science ®Google Scholar
• Francillon-Vieillot, H., Buffrénil, V., Castanet, J., Géraudie, J., Meunier, F. J, Sire, J. Y, Zylberberg, L., & Ricqlès, A. (1990). Microstructure and mineralization of vertebrate skeletal tissues. Skeletal biomineralization: patterns, processes and evolutionary trends, 1, 471–530.
   Google Scholar
• Gauthier, J. (1986). Saurischian monophyly and the origin of birds. Memoirs of the California Academy of sciences, 8, 1–55. https://doi.org/10.1017/S247526300000091X.
   Google Scholar
• Goloboff, P. A., & Catalano, S. A. (2016). TNT version 1.5, including a full implementation of phylogenetic morphometrics. Cladistics, 32(3), 221–238. https://doi.org/10.1111/cla.12160.
   PubMed Web of Science ®Google Scholar
• Goloboff, P. A., Farris, J. S., Källersjö, M., Oxelman, B., Ramírez, M. J., & Szumik, C.A. (2003). Improvements to resampling measures of group support. Cladistics, 19(4), 324–32. https://doi.org/10.1111/j.1096-0031.2003.tb00376.x.
   Web of Science ®Google Scholar
• Goloboff, P. A., Farris, J. S., & Nixon, K.C. (2008). TNT, a free program for phylogenetic analysis. Cladistics, 24(5), 774–786. https://doi.org/10.1111/j.1096-0031.2008.00217.x.
   Web of Science ®Google Scholar
• Gordon, M. (1947). Classificação das formações gondwânicas do Paraná, Santa Catarina e Rio Grande do Sul. Notas Preliminares e Estudos, DNPM/DGM, Rio de Janeiro 38, 1–20.
   Google Scholar
• Griffin, C. T., & Nesbitt, S. J. (2016). The femoral ontogeny and long bone histology of the Middle Triassic (? late Anisian) dinosauriform Asilisaurus kongwe and implications for the growth of early dinosaurs. Journal of Vertebrate Paleontology, 36(3), e1111224. https://doi.org/10.1080/02724634.2016.1111224.
   Web of Science ®Google Scholar
• Hendrickx, C., Mateus, O., & Araújo, R. (2015). A proposed terminology of theropod teeth (Dinosauria, Saurischia). Journal of Vertebrate Paleontology, 35(5), e982797. https://doi.org/10.1080/02724634.2015.982797.
   Web of Science ®Google Scholar
• Horn, B. L. D., Melo, T. M., Schultz, C. L., Philipp, R. P., Kloss, H. P., & Goldberg, K. (2014). A new third-order sequence stratigraphic framework applied to the Triassic of the Paraná Basin, Rio Grande do Sul, Brazil, based on structural, stratigraphic and paleontological data. Journal of South American Earth Sciences, 55, 123–132. https://doi.org/10.1016/j.jsames.2014.07.007.
   Web of Science ®Google Scholar
• Huene, F. V. (1942). Die fossilen Reptilien des Südamerikanischen Gondwanalandes. Ergebnisse der Sauriergrabungen in Südbrasilien, 1928/1929. C. H. Beck’sche Verlagsbuchhandlung, Munich, 161–332.
   Google Scholar
• Huene, F. V. (1928). Ein Cynodontier aus der Trias Brasiliens. Centralblatt für Mineralogie, Geologie und Paläontologie B, 251–270.
   Google Scholar
• Irmis, R. B., Nesbitt, S. J., Padian, K., Smith, N. D., Turner, A. H., Woody, D., & Downs, A. (2007). A late Triassic Dinosauromorph assemblage from New Mexico and the rise of dinosaurs. Science, 317 (5836), 358–361. DOI: 10.1126/science.1143325.
   PubMed Web of Science ®Google Scholar
• Kammerer, C. F., Nesbitt, S.J ., & Shubin, N. H. (2011). The first silesaurid dinosauriform from the Late Triassic of Morocco. Acta Palaeontologica Polonica, 57(2), 277–284. https://doi.org/10.4202/app.2011.0015.
   Web of Science ®Google Scholar
• Klein, N., & Sander, P. M. (2007). Bone histology and growth of the prosauropod Plateosaurus engelhardti Meyer, 1837 from the Norian bonebeds of Trossingen (Germany) and Frick (Switzerland). Special Papers in Palaeontology 77, 169–206.
   Google Scholar
• Langer, M. C. (2003). The pelvic and hind limb anatomy of the stem sauropodomorph Saturnalia tupiniquim (Late Triassic, Brazil). PaleoBios, 23, 1–40.
   Google Scholar
• Langer, M. C. (2005a). Studies on continental Late Triassic tetrapod biochronology. I. The type locality of Saturnalia tupiniquim and the faunal succession in south Brazil. Journal of South American Earth Sciences, 19:205–218. https://doi.org/10.1016/j.jsames.2005.04.003.
   Web of Science ®Google Scholar
• Langer, M. C. (2005b). Studies on continental Late Triassic tetrapod biochronology. II. The Ischigualastian and a Carnian global correlation. Journal of South American Earth Sciences, 19, 219–239. https://doi.org/10.1016/j.jsames.2005.04.002.
   Web of Science ®Google Scholar
• Langer, M. C. (2014). The origins of Dinosauria: much ado about nothing. Palaeontology, 57(3), 469–478. https://doi.org/10.1111/pala.12108.
   Web of Science ®Google Scholar
• Langer, M. C., Abdala, F., Richter, M., & Benton, M. J. (1999). A sauropodomorph dinosaur from the Upper Triassic (Carnian) of southern Brazil. Comptes Rendus de l’Academie des Sciences, 329, 511–517. https://doi.org/10.1016/S1251-8050(00)80025-7
   Web of Science ®Google Scholar
• Langer, M. C., & Benton, M. J. (2006). Early dinosaurs: a phylogenetic study. Journal of Systematic Palaeontology, 4(4), 309–358. https://doi.org/10.1017/S1477201906001970.
   Web of Science ®Google Scholar
• Langer, M. C., Ezcurra, M. D., Bittencourt, J. S., & Novas, F. E. (2010). The origin and early evolution of dinosaurs. Biological Reviews, 85(1), 55–110. https://doi.org/10.1111/j.1469-185X.2009.00094.x.
   PubMed Web of Science ®Google Scholar
• Langer, M. C., Ezcurra, M. D., Rauhut, O. W., Benton, M. J., Knoll, F., McPhee, B. W., Novas, F. E., Pol, D., & Brusatte, S. L. (2017). Untangling the dinosaur family tree. Nature, 551(7678), E1–E3. https://doi.org/10.1038/nature21700.
   PubMed Web of Science ®Google Scholar
• Langer, M. C., & Ferigolo, J. (2013). The Late Triassic dinosauromorph Sacisaurus agudoensis (Caturrita Formation; Rio Grande do Sul, Brazil): anatomy and affinities. Geological Society, London, Special Publications, 379(1), 353–392. https://doi.org/10.1144/SP379.16.
   Google Scholar
• Langer, M. C., Nesbitt, S. J., Bittencourt, J. S., & Irmis, R. B. (2013). Non-dinosaurian dinosauromorpha. Geological Society, London, Special Publications, 379(1), 157–186. https://doi.org/10.1144/SP379.9.
   Google Scholar
• Langer, M. C., Ramezani, J., & Da Rosa, A. S. (2018). U-Pb age constraints on dinosaur rise from South Brazil. Gondwana Research, 57, 133–140. https://doi.org/10.1016/j.gr.2018.01.005.
   Web of Science ®Google Scholar
• Langer, M. C., Ribeiro, A. M., Schultz, C. L., & Ferigolo, J. (2007). The continental tetrapod-bearing Triassic of South Brazil. New Mexico Museum of Natural History and Science Bulletin, 41, 201–218.
   Google Scholar
• LeBlanc, A. R. H., Brink, K. S., Cullen, T. M., & Reisz, R. R. (2017). Evolutionary implications of tooth attachment versus tooth implantation: a case study using dinosaur, crocodilian, and mammal teeth. Journal of Vertebrate Paleontology, 37(5), e1354006. https://doi.org/10.1080/02724634.2017.1354006
   Web of Science ®Google Scholar
• Luca, S., & Luca, A. P. (1980). The postcranial skeleton of Heterodontosaurus tucki (Reptilia, Ornithischia) from the Stormberg of South Africa. Annals of the South African Museum, 79, 159–211.
   Google Scholar
• Margerie, E., Cubo, J., & Castanet, J. (2002). Bone typology and growth rate: testing and quantifying ‘Amprino’s rule’in the mallard (Anas platyrhynchos). Comptes rendus biologies, 325(3), 221–230. https://doi.org/10.1016/S1631-0691(02)01429-4.
   PubMed Web of Science ®Google Scholar
• Marsà, J. A. G., Agnolín, F. L., & Novas, F. (2017). Bone microstructure of Lewisuchus admixtus Romer, 1972 (Archosauria, Dinosauriformes). Historical Biology, 31(2), 157–162. https://doi.org/10.1080/08912963.2017.1347646.
   Web of Science ®Google Scholar
• Marsola, J. C., Bittencourt, J. S., Butler, R. J., Da Rosa Á. A., Sayão, J. M., & Langer, M. C. (2019a). A new dinosaur with theropod affinities from the Late Triassic Santa Maria Formation, South Brazil. Journal of Vertebrate Paleontology, 38(5), e1531878. https://doi.org/10.1080/02724634.2018.1531878.
   Web of Science ®Google Scholar
• Marsola, J. C., Bittencourt, J. S., Da Rosa, Á. A., Martinelli, A. G., Ribeiro, A. M., Ferigolo, J., & Langer, M. C. (2018). New sauropodomorph and cynodont remains from the late Triassic Sacisaurus site in Southern Brazil and its stratigraphic position in the Norian Caturrita Formation. Acta Palaeontologica Polonica, 63(4), 653–669. https://doi.org/10.4202/app.00492.2018.
   Web of Science ®Google Scholar
• Marsola, J. C., Ferreira, G. S., Langer, M. C., Button, D. J., & Butler, R. J. (2019b). Increases in sampling support the southern Gondwanan hypothesis for the origin of dinosaurs. Palaeontology, 62(3), 473–482. https://doi.org/10.5061/dryad.6t5m946.
   Web of Science ®Google Scholar
• Martinelli, A. G., Eltink, E., Da Rosa, Á. A., & Langer, M. C. (2017). A new cynodont from the Santa Maria Formation, south Brazil, improves Late Triassic probainognathian diversity. Papers in Palaeontology 3, 401–423. https://doi.org/10.5061/dryad.q2m2v.
   Web of Science ®Google Scholar
• Martínez, R. N., Apaldetti, C., Alcober, O. A., Colombi, C. E., Sereno, P. C., Fernandez, E., Malnis, P. S., Correa, G. A. & Abelin, D. (2012). Vertebrate succession in the Ischigualasto Formation. Journal of Vertebrate Paleontology, 32, 10–30. https://doi.org/10.1080/02724634.2013.818546.
   Web of Science ®Google Scholar
• Martínez, R. N., Sereno, P. C., Alcober, O. A., Colombi, C. E., Renne, P. R., Montañez, I. P., & Currie, B. S. (2011). A basal dinosaur from the dawn of the dinosaur era in southwestern Pangaea. Science, 331(6014), 206–210. https://doi.org/10.1126/science.119846.
   PubMed Web of Science ®Google Scholar
• Martz, J. W., & Small, B. J. (2019). Non-dinosaurian dinosauromorphs from the Chinle Formation (Upper Triassic) of the Eagle Basin, northern Colorado: Dromomeron romeri (Lagerpetidae) and a new taxon, Kwanasaurus williamparkeri (Silesauridae). PeerJ, 7, e7551. https://doi.org/10.7717/peerj.7551.
   PubMed Web of Science ®Google Scholar
• Mestriner, G., LeBlanc, A. R. H., Nesbitt, S. J., Marsola, J. C. A., Irmis, R. B., Da Rosa Á. A., Ribeiro, A. M., Ferigolo, J., & Langer, M. C. (2022). Histological analysis of ankylothecodonty in Silesauridae (Archosauria: Dinosauriformes) and its implications for the evolution of dinosaur tooth attachment. The Anatomical Record, 305(2), 393–423. https://doi.org/10.1002/ar.24679.
   Google Scholar
• Müller, R. T., & Garcia, M. S. (2023) A new silesaurid from Carnian beds of Brazil fills a gap in the radiation of avian line archosaurs. Scientific Reports, 13(1), 4981. https://doi.org/10.1038/s41598-023-32057-x.
   PubMed Web of Science ®Google Scholar
• Müller, R. T., & Garcia, M. S. (2020). A paraphyletic ‘Silesauridae’ as an alternative hypothesis for the initial radiation of ornithischian dinosaurs. Biology Letters, 16(8), 20200417. https://doi.org/10.1098/rsbl.2020.0417.
   PubMed Web of Science ®Google Scholar
• Nesbitt, S. J. (2011). The early evolution of archosaurs: relationships and the origin of major clades. Bulletin of the American Museum of Natural History, 352(1), 1–292. http://doi.org/10.1206/352.1.
   Google Scholar
• Nesbitt, S. J., Butler, R. J., Ezcurra, M. D., Barrett, P. M., Stocker, M. R., Angielczyk, K.D., Smith, R. M. H., Sidor, C. A., Niedzwiedzki, G., Sennikov, A. G., & Charig, A. J. (2017a). The earliest bird-line archosaurs and the assembly of the dinosaur body plan. Nature, 544(7651), 484–487. https://doi.org/10.1038/nature22037.
   PubMed Web of Science ®Google Scholar
• Nesbitt, S .J., Butler, R .J., Ezcurra, M. D., Charig, A. J., & Barrett P. M. (2017b). The anatomy of Teleocrater rhadinus, an early avemetatarsalian from the lower portion of the Lifua Member of the Manda Beds (Middle Triassic). Journal of Vertebrate Paleontology, 37, 142–177. https://doi.org/10.1080/02724634.2017.1396539.
   Web of Science ®Google Scholar
• Nesbitt, S. J., Irmis, R. B., & Parker, W. G. (2007). A critical re-evaluation of the Late Triassic dinosaur taxa of North America. Journal of Systematic Palaeontology, 5(2), 209–243. https://doi.org/10.1017/S1477201907002040.
   Web of Science ®Google Scholar
• Nesbitt, S. J., Irmis, R. B., Parker, W. G., Smith, N. D., Turner, A. H., & Rowe, T. (2009). Hindlimb osteology and distribution of basal dinosauromorphs from the Late Triassic of North America. Journal of Vertebrate Paleontology, 29(2), 498–516. https://doi.org/10.1671/039.029.0218.
   Web of Science ®Google Scholar
• Nesbitt, S. J., Langer, M. C., & Ezcurra, M. D. (2019). The anatomy of Asilisaurus kongwe, a dinosauriform from the Lifua Member of the Manda Beds (∼ Middle Triassic) of Africa. The Anatomical Record, 303(4), 813–873. https://doi.org/10.1002/ar.24287.
   Google Scholar
• Nesbitt, S, J., Sidor, C. A., Irmis, R. B., Angielczyk, K. D., Smith, R. M., & Tsuji, L. A. (2010). Ecologically distinct dinosaurian sister group shows early diversification of Ornithodira. Nature, 464(7285), 95. https://doi.org/10.1038/nature08718.
   PubMed Web of Science ®Google Scholar
• Norman, D. B., Baron, M. G., Garcia, M. S., & Müller, R. T. (2022). Taxonomic, palaeobiological and evolutionary implications of a phylogenetic hypothesis for Ornithischia (Archosauria: Dinosauria). Zoological Journal of the Linnean Society, 196(4), 1273–1309. https://doi.org/10.1093/zoolinnean/zlac062.
   Web of Science ®Google Scholar
• Novas, F. E., Agnolin, F. L., Ezcurra, M. D., Müller, R. T., Martinelli, A., & Langer, M. C. (2021). Review of the fossil record of early dinosaurs from South America, and its phylogenetic implications. Journal of South American Earth Sciences, 110, 103341. https://doi.org/10.1016/j.jsames.2021.103341.
   Web of Science ®Google Scholar
• Novas, F. E. (1996). Dinosaur monophyly. Journal of Vertebrate Paleontology, 16(4), 723–741. https://doi.org/10.1080/02724634.1996.10011361.
   Web of Science ®Google Scholar
• Novas, F. E., 1992. La evolución de los dinosaurios carnívoros. In Los dinosaurios y su entorno biótico: II Curso de Paleontología, 10 a 12 de julio de 1990. Actas, 125–163.
   Google Scholar
• Novas, F. E. (1994). New information on the systematics and postcranial skeleton of Herrerasaurus ischigualastensis (Theropoda: Herrerasauridae) from the Ischigualasto Formation (Upper Triassic) of Argentina. Journal of Vertebrate Paleontology, 13(4), 400–423. https://doi.org/10.1080/02724634.1994.10011523.
   Google Scholar
• Padian, K., Horner, J. R., & Ricqlès, A. (2004). Growth in small dinosaurs and pterosaurs: the evolution of archosaurian growth strategies. Journal of Vertebrate Paleontology, 24(3), 555–571. https://doi.org/10.1671/0272-4634.
   Web of Science ®Google Scholar
• Padian, K., & Lamm, E.T. (2013). Bone histology of fossil tetrapods: advancing methods, analysis, and interpretation. University of California Press.
   Google Scholar
• Peecook, B. R., Sidor, C. A., Nesbitt, S. J., Smith, R. M., Steyer, J. S., & Angielczyk, K. D. (2013). A new silesaurid from the upper Ntawere Formation of Zambia (Middle Triassic) demonstrates the rapid diversification of Silesauridae (Avemetatarsalia, Dinosauriformes). Journal of Vertebrate Paleontology, 33(5), 1127–1137. https://doi.org/10.1080/02724634.2013.755991.
   Web of Science ®Google Scholar
• Pretto, F. A., Müller, R. T., Moro, D., Garcia, M. S., Neto, V. D. P. & Da Rosa, Á. A. S., (2022). The oldest South American silesaurid: New remains from the Middle Triassic (Pinheiros-Chiniquá Sequence, Dinodontosaurus Assemblage Zone) increase the time range of silesaurid fossil record in southern Brazil. Journal of South American Earth Sciences, 120, 104039. https://doi.org/10.1016/j.jsames.2022.104039.
   Web of Science ®Google Scholar
• Prondvai, E., Stein, K. H., De Ricqlès, A., & Cubo, J. (2014). Development-based revision of bone tissue classification: the importance of semantics for science. Biological Journal of the Linnean Society, 112(4), 799–816. https://doi.org/10.1111/bij.12323.
   Web of Science ®Google Scholar
• Qvarnström, M., Wernström, J. V., Piechowski, R., Talanda, M., Ahlberg, P. E., & Niedzwiedzki, G. (2019). Beetle-bearing coprolites possibly reveal the diet of a Late Triassic dinosauriform. Royal Society Open Science, 6(3), 181042. https://doi.org/10.1098/rsos.181042.
   PubMed Web of Science ®Google Scholar
• Rauhut, O. W. M. (2003). The interrelationships and evolution of basal theropod dinosaurs. Special Papers in Palaeontology, 69, 1–213. https://doi.org/10.1017/S0016756803248834.
   Google Scholar
• Romer, A. S. (1972). The Chañares (Argentina) Triassic reptile fauna. XIV. Lewisuchus admixtus a further thecodont from the Chamares Bed. Breviora 390, 1–13.
   Google Scholar
• Sarigul, V., Agnolin, F., & Chatterjee, S. (2018). Description of a multitaxic bone assemblage from the Upper Triassic Post Quarry of Texas (Dockum Group), including a new small basal dinosauriform taxon. Historia Natural, Tercera Serie Volumen 8 (1), 5–24.
   Google Scholar
• Schultz, C. L., Martinelli, A. G., Soares, M. B., Pinheiro, F. L., Kerber, L., Horn, B. L., Pretto, F. A., Müller, R. T., & Melo, T. P. (2020). Triassic faunal successions of the Paraná Basin, southern Brazil. Journal of South American Earth Sciences, 104, 102846. https://doi.org/10.1016/j.jsames.2020.102846.
   Web of Science ®Google Scholar
• Sereno, P. C., & Arcucci, A. B. (1994). Dinosaurian precursors from the Middle Triassic of Argentina: Marasuchus lilloensis, gen. nov. Journal of Vertebrate Paleontology, 14(1), 53–73. https://doi.org/10.1080/02724634.1994.10011538
   Web of Science ®Google Scholar
• Stein, K. H. W. (2011). Long bone histology of basalmost and derived Sauropodomorpha: the convergence of fibrolamellar bone and the evolution of giantism and nanism. [Unpublished doctoral’s dissertation]. Universitäts-und Landesbibliothek Bonn.
   Google Scholar
• Sullivan, R. M., & Lucas, S. G. (1999). Eucoelophysis baldwini a new theropod dinosaur from the Upper Triassic of New Mexico, and the status of the original types of Coelophysis. Journal of Vertebrate Paleontology, 19(1), 81–90. https://doi.org/10.1080/02724634.1999.10011124.
   Web of Science ®Google Scholar
• Tykoski, R. (2005). Osteology, ontogeny, and relationships of the coelophysoid theropods. [Unpublished doctoral’s dissertation]; The University of Texas, Austin.
   Google Scholar
• Veiga, F. H., Botha-Brink, J., Ribeiro, A. ., Ferigolo, J., & Soares, M.B. (2018). Osteohistology of the silesaurid Sacisaurus agudoensis from southern Brazil (Late Triassic) and implications for growth in early dinosaurs. Anais da Academia Brasileira de Ciências, 91. https://doi.org/10.1590/0001-3765201920180643.
   Web of Science ®Google Scholar
• Wilson, J. A. (1999). A nomenclature for vertebral laminae in sauropods and other saurischian dinosaurs. Journal of vertebrate Paleontology, 19(4), 639–653.
   Web of Science ®Google Scholar
• Wilson, J. A., D'Emic, M. D., Ikejiri, T., Moacdieh, E. M. & Whitlock, J. A. (2011). A nomenclature for vertebral fossae in sauropods and other saurischian dinosaurs. PLoS ONE, 6(2), e17114.
   PubMed Web of Science ®Google Scholar
• Witmer, L. M. (1997). The evolution of the antorbital cavity of archosaurs: a study in soft-tissue reconstruction in the fossil record with an analysis of the function of pneumaticity. Society of Vertebrate Paleontology, Memoir, 3, 1–73. https://doi.org/10.1080/02724634.1997.10011027.
   Google Scholar
• Yates, A. M. (2003). A new species of the primitive dinosaur Thecodontosaurus (Saurischia: Sauropodomorpha) and its implications for the systematics of early dinosaurs. Journal of Systematic Palaeontology, 1(1), 1–4. https://doi.org/10.1017/S1477201903001007.
   Google Scholar
• Zerfass, H., Lavina, E. L., Schultz, C. L., Garcia, A. J. V., Faccini, U. F., & Chemale, J. F. (2003). Sequence stratigraphy of continental Triassic strata of Southernmost Brazil: a contribution to Southwestern Gondwana palaeogeography and palaeoclimate. Sedimentary Geology, 161(1-2), 85–105. https://doi.org/10.1016/S0037-0738(02)00397-4.
   Web of Science ®Google Scholar