Leg bones of a new penguin species from the Waipara Greensand add to the diversity of very large-sized Sphenisciformes in the Paleocene of New Zealand

References

• Acosta Hospitaleche, C., 2014. New giant penguin bones from Antarctica: systematic and paleobiological significance. Comptes Rendus Palevol 13, 555–560.
   Web of Science ®Google Scholar
• Ando, T. & Fordyce, R.E., 2014. Evolutionary drivers for flightless, wing-propelled divers in the Northern and Southern Hemispheres. Palaeogeography, Palaeoclimatology, Palaeoecology 400, 50–61.
   Web of Science ®Google Scholar
• Baumel, J.J. & Witmer, L.M., 1993. Osteologia. In Handbook of Avian Anatomy: Nomina Anatomica Avium, 2nd edition. Baumel, J.J., King, A.S., Breazile, J.E., Evans, H.E. & Vanden Berge, J.C., eds, Publications of the Nuttall Ornithological Club, Cambridge, Massachusetts, 23, 45–132.
   Google Scholar
• ChÁvez Hoffmeister, M., 2014. Phylogenetic characters in the humerus and tarsometatarsus of penguins. Polish Polar Research 35, 469–496.
   Web of Science ®Google Scholar
• Clarke, J.A., Ksepka, D.T., Salas-Gismondi, R., Altamirano, A.J., Shawkey, M.D., D’Alba, L., Vinther, J., DeVries, T.J. & Baby, P., 2010. Fossil evidence for evolution of the shape and color of penguin feathers. Science 330, 954–957.
   PubMed Web of Science ®Google Scholar
• Elzanowski, A., 2008. The avian femur: morphology and terminology of the lateral condyle. Oryctos 7, 1–5.
   Google Scholar
• Fordyce, R.E. & Jones, C.M., 1986. The world’s oldest penguin? Geological Society of New Zealand Newsletter 74, 56–57.
   Google Scholar
• Fordyce, R.E. & Thomas, D.B., 2011. Kaiika maxwelli, a new Early Eocene archaic penguin (Sphenisciformes, Aves) from Waihao Valley, South Canterbury, New Zealand. New Zealand Journal of Geology and Geophysics 54, 43–51.
   Web of Science ®Google Scholar
• Jadwiszczak, P., 2006. Eocene penguins of Seymour Island, Antarctica: taxonomy. Polish Polar Research 27, 3–62.
   Google Scholar
• Jadwiszczak, P., 2012. Partial limb skeleton of a ‘giant penguin’ Anthropornis from the Eocene of Antarctic Peninsula. Polish Polar Research 33, 259–274.
   Web of Science ®Google Scholar
• Jadwiszczak, P., Acosta Hospitaleche, C. & Reguero, M., 2013. Redescription of Crossvallia unienwillia: the only Paleocene Antarctic penguin. Ameghiniana 50, 545–553.
   Web of Science ®Google Scholar
• Ksepka, D.T. & Ando, T., 2011. Penguins past, present, and future: Trends in the evolution of the Sphenisciformes. In Living Dinosaurs. The Evolutionary History of Modern Birds. Dyke, G. & Kaiser, G., eds, John Wiley and Sons, Chichester, 155–186.
   Google Scholar
• Ksepka, D.T. & Clarke, J.A., 2010. The basal penguin (Aves: Sphenisciformes) Perudyptes devriesi and a phylogenetic evaluation of the penguin fossil record. Bulletin of the American Museum of Natural History 337, 1–77.
   Google Scholar
• Ksepka, D.T., Clarke, J.A., DeVries, T.J. & Urbina, M., 2008. Osteology of Icadyptes salasi, a giant penguin from the Eocene of Peru. Journal of Anatomy 213, 131–147.
   PubMed Web of Science ®Google Scholar
• Ksepka, D.T., Fordyce, R.E., Ando, T. & Jones, C.M., 2012. New fossil penguins (Aves, Sphenisciformes) from the Oligocene of New Zealand reveal the skeletal plan of stem penguins. Journal of Vertebrate Paleontology 32, 235–254.
   Web of Science ®Google Scholar
• Mayr, G., 2009. Paleogene Fossil Birds. Springer, Heidelberg, 262 pp.
   Google Scholar
• Mayr, G., 2017. Avian Evolution: The Fossil Record of Birds and Its Paleobiological Significance. Wiley-Blackwell, Chichester, 293 pp.
   Google Scholar
• Mayr, G., Goedert, J.L. & Vogel, O., 2015. Oligocene plotopterid skulls from western North America and their bearing on the phylogenetic affinities of these penguin-like seabirds. Journal of Vertebrate Paleontology 35, e943764.
   Web of Science ®Google Scholar
• Mayr, G., De Pietri, V.L., Love, L., Mannering, A.A. & Scofield, R.P., 2017a. A well-preserved new mid-Paleocene penguin (Aves, Sphenisciformes) from the Waipara Greensand in New Zealand. Journal of Vertebrate Paleontology 37, e139816915.
   Web of Science ®Google Scholar
• Mayr, G., De Pietri, V.L. & Scofield, R.P., 2017b. A new fossil from the mid-Paleocene of New Zealand reveals an unexpected diversity of world’s oldest penguins. The Science of Nature 104, 9.
   Google Scholar
• Mayr, G., Scofield, R.P., De Pietri, V.L. & Tennyson, A.J.D., 2017c. A Paleocene penguin from New Zealand substantiates multiple origins of gigantism in fossil Sphenisciformes. Nature Communications 8, 1927.
   PubMed Web of Science ®Google Scholar
• Mayr, G., De Pietri, V.L., Love, L., Mannering, A.A. & Scofield, R.P., in press. Oldest, smallest, and phylogenetically most basal pelagornithid, from the early Paleocene of New Zealand, sheds light on the evolutionary history of the largest flying birds. Papers in Palaeontology.
   Web of Science ®Google Scholar
• Myrcha, A., Jadwiszczak, P., Tambussi, C.P., Noriega, J.I., Gazdzicki, A., Tatur, A. & Del Valle, R., 2002. Taxonomic revision of Eocene Antarctic penguins based on tarsometatarsal morphology. Polish Polar Research 23, 5–46.
   Google Scholar
• Simpson, G.G., 1971. Review of fossil penguins from Seymour Island. Proceedings of the Royal Society of London, Series B, Biological Sciences 178, 357–387.
   Web of Science ®Google Scholar
• Simpson, G.G., 1975. Fossil penguins. In The Biology of Penguins. Stonehouse, B., ed., The Macmillan Press, London, 19–41.
   Google Scholar
• Slack, K.E., Jones, C.M., Ando, T., Harrison, G.L., Fordyce, R.E., Arnason, U. & Penny, D., 2006. Early penguin fossils, plus mitochondrial genomes, calibrate avian evolution. Molecular Biology and Evolution 23, 1144–1155.
   PubMed Web of Science ®Google Scholar
• Stephan, B., 1979. Vergleichende Osteologie der Pinguine. Mitteilungen des Zoologischen Museums in Berlin 55, Supplement Annalen für Ornithologie 3, 3–98.
   Google Scholar
• Tambussi, C.P., Reguero, M.A., Marenssi, S.A. & Santillana, S.N., 2005. Crossvallia unienwillia, a new Spheniscidae (Sphenisciformes, Aves) from the late Paleocene of Antarctica. Geobios 38, 667–675.
   Web of Science ®Google Scholar