A diversified tetrapod ichnite fauna from the Lower Cretaceous Hekou Group of Gansu Province, China

References

• Alexander RM. 1976. Estimates of speeds of dinosaurs. Nature. 261:129–130. doi:10.1038/261129a0.
   Web of Science ®Google Scholar
• Belvedere M, Farlow JO. 2016. A numerical scale for quantifying the quality of preservation of vertebrate tracks. In: Falkingham PL, Marty D, Richter A, editors. Dinosaur tracks: the next steps. Bloomington and Indianapolis: Indiana University Press; p. 92–99.
   Google Scholar
• Cai XF, Li CA, Gu YS. 1999. First record of dinosaur footprints from Lanzhou-Minhe Basin. Earth Sci J China Univ Geosci. 24:216. (in Chinese).
   Google Scholar
• Calvo JO. 1991. Huellas de dinosaurios en la Formación Rio Limay (Albiano-Cenomaniano?), Picun Leufu, Provincia de Neuquén, República Argentina. (Ornithischia-Saurischia: Sauropoda–Theropoda). Ameghiniana. 28:241–258.
   Google Scholar
• Calvo JO 1999. Dinosaurs and other vertebrates of the Lake Ezequiel Ramos Mexia area, Neuquén-Patagonia, Argentina. In: Tomida Y, Rich TH, Vickers-Rich P, editors. Proceedings of the Second Gondwanan Dinosaur symposium. Vol. 15. Tokyo: National Science Museum Monographs; p. 13–45.
   Google Scholar
• Casamiquela RM, Fasola A. 1968. Sobre pisadas de dinosaurios del Cretácico Inferior de Colchagua (Chile). Univ Chile Depart Geol. 30:1–24.
   Google Scholar
• Chen J 2013. Sedimentary Characteristics and Paleogeography of the Hekou Group in Lanzhou-Minhe Basin in the Early Cretaceous. Master Thesis, China University of Geosciences; p. 73.
   Google Scholar
• Chen J, Liu YQ, Kuang HW, Liu YX, Peng N, Xu H, Dong C, Liu H, Xue PL, Xu JL. 2013. Sedimentary characteristics and their basin analysis significance of the Lower Cretaceous Hekou Group in Zhongpu area of Lanzhou-Minhe Basin, Gansu Province. J Palaeogeogr. 15(2):155–168.
   Google Scholar
• Cobos A, Royo-Torres R, Luque L, Alcalá L, Mampel L. 2010. An Iberian stegosaurs paradise: the Villar del Arbispo Formation (Tithonian-Berriasian) in Teruel (Spain). Palaeogeogr Palaeoclimatol Palaeoecol. 293:223–236. doi:10.1016/j.palaeo.2010.05.024.
   Web of Science ®Google Scholar
• Díaz-Martínez I, Pereda-Suberbiola X, Perez-Lorente F, Canudo JI. 2015. Ichnotaxonomic review of large ornithopod dinosaur tracks: temporal and geographic implications. PloS One. 10(2):e0115477. doi:10.1371/journal.pone.0115477.
   PubMed Web of Science ®Google Scholar
• Farlow JO, Pittman JG, Hawthorne JM. 1989. Brontopodus birdi, lower cretaceous dinosaur footprints from the U.S. Gulf Coastal Plain. In: Gillette DD, Lockley MG, editors. Dinosaur tracks and traces. Cambridge: Cambridge University Press; p. 371–394.
   Google Scholar
• González Riga, B.J., 2011. Speeds and stance of titanosaur sauropods: analysis of Titanopodus tracks from the Late Cretaceous of Mendoza, Argentina. Anais da Academia Brasileira de Ciências, 83(1):279–290. doi:10.1590/S0001-37652011005000002
   PubMed Web of Science ®Google Scholar
• Hayashi K. 2006. Nonmarine ostracode zonation and longdistance correlationbased on analysis of regional ostracode successions in China, Korea, Japan, and Mongolia. Cretaceous Res. 27:168–188. doi:10.1016/j.cretres.2005.11.007.
   Web of Science ®Google Scholar
• Ishigaki S, Matsumoto Y. 2009. “Off-tracking”-like phenomenon observed in the turning sauropod trackway from the Upper Jurassic of Morocco. Mem Fukui Prefectural Dinosaur Mus. 8:1–10.
   Google Scholar
• Kim JY, Lockley MG, Kim HM, Lim JD, Kim SH, Lee SJ, Woo JO, Park HJ, Kim HS, Kim KS. 2009. New dinosaur tracks from Korea, Ornithopodichnus masanensis ichnogen. et ichnosp. nov. (Jindong Formation, Lower Cretaceous): implications for polarities in ornithopod foot morphology. Cretaceous Res. 30:1387–1397. doi:10.1016/j.cretres.2009.08.003.
   Web of Science ®Google Scholar
• Kuban G. 1989. Elongate dinosaur tracks. In: Gillette DD, Lockley MG, editors. Dinosaur tracks and traces, 1989. Cambridge: Cambridge University Press; p. 57–72.
   Google Scholar
• Leonardi G. 1984. Le impronte fossili di dinosauri. In: Bonaparte JF, Colbert EH, Currie PJ, de Rocles A, Kielan-Jaworowska Z, Leonardi G, Morello N, Taquet P, editors. Sulle ormi de dinosauri. Venice: Editio Editrice; p. 165–186.
   Google Scholar
• Leonardi G. 1987. Glossary and manual of tetrapod footprint palaeoichnology. Brazil: Departamento Nacional de Producao Mineral; p. 75.
   Google Scholar
• Li D, Xing L, Lockley MG, Pinuela L, Zhang J, Dai H, Kim JY, Scott Persons W IV, Kong D. 2015. A manus dominated pterosaur track assemblage form Gansu, China: implications for behavior. China Sci Bull. 60:264–272.
   Google Scholar
• Li LG, Li DQ, You HL, Dodson P. 2014. A new titanosaurian sauropod from the Hekou Group (Lower Cretaceous) of the Lanzhou-Minhe Basin, Gansu Province, China. PLoS ONE. 9:e85979. doi:10.1371/journal.pone.0085979.
   PubMed Web of Science ®Google Scholar
• Lockley MG. 1991. Tracking dinosaurs: a new look at an ancient world. Cambridge, NY: Cambridge University Press; p. 238.
   Google Scholar
• Lockley MG, Hunt AP, Meyer C. 1994a. Vertebrate tracks and the ichnofacies concept: implications for paleoecology and palichnostratigraphy. In: Donovan S, editor. The paleobiology of trace fossils. New York (USA): Wiley and Sons; p. 241–268.
   Google Scholar
• Lockley MG. 1999. The eternal trail: a tracker looks at evolution. Cambridge: Perseus Books; p. 1–334.
   Google Scholar
• Lockley MG. 2001. Trackways and dinosaur locomotion. In: Briggs DEG, Crowther P, editors. Palaeobiology II: a synthesis. Oxford: Blackwell Science; p. 412–416.
   Google Scholar
• Lockley MG, Farlow JO, Meyer CA. 1994b. Brontopodus and Parabrontopodus ichnogen. nov. and the significance of wide- and narrow-gauge sauropod trackways. Gaia. 10:135–146.
   Google Scholar
• Lockley MG, Huh M, Kim BS. 2012. Ornithopodichnus and pes-only sauropod trackways from the Hwasun tracksite Cretaceous of Korea. Ichnos. 19:93–100. doi:10.1080/10420940.2011.625726.
   Web of Science ®Google Scholar
• Lockley MG, Hunt AP. 1995. Dinosaur tracks and other fossil footprints of the western United States. New York: Columbia University Press; p. 338.
   Google Scholar
• Lockley MG, Li JJ, Li RH, Matsukawa M, Harris JD, Xing LD. 2013. A review of the tetrapod track record in China, with special reference to type ichnospecies: implications for ichnotaxonomy and paleobiology. Acta Geol Sin. 87(1):1–20. English edition. doi:10.1111/1755-6724.12026.
   Web of Science ®Google Scholar
• Lockley MG, Nadon G, Currie PJ. 2003. A diverse dinosaur-bird footprint assemblage from the Lance Formation, Upper Cretaceous, Eastern Wyoming: implications for ichnotaxonomy. Ichnos. 11:229–249. doi:10.1080/10420940490428625.
   Google Scholar
• Lockley MG, Wright J, White D, Li JJ, Feng L, Li H. 2002. The first sauropod trackways from China. Cretaceous Res. 23:363–381. doi:10.1006/cres.2002.1005.
   Web of Science ®Google Scholar
• Lockley MG, Xing LD, Lockwood JAF, Pond S. 2014. A review of large Cretaceous ornithopod tracks, with special reference to their ichnotaxonomy. Biol J Linn Soc. 113:721–736. doi:10.1111/bij.12294.
   Web of Science ®Google Scholar
• Lucas SG. 2001. Taphotaxon. Lethaia. 34:30. doi:10.1080/002411601300068198.
   Web of Science ®Google Scholar
• Lull RS. 1904. Fossil footprints of the Jura-Trias of North America. Mem Boston Soc Nat Hist. 5:461–557.
   Google Scholar
• Marty D 2008 . Sedimentology, taphonomy, and ichnology of Late Jurassic dinosaur tracks from the Jura carbonate platform (ChevenezeCombe Ronde tracksite, NW Switzerland): insights into the tidal-flat palaeoenvironment and dinosaur diversity, locomotion, and palaeoecology. PhD dissertation, University of Fribourg; p. 278.
   Google Scholar
• Marty D, Belvedere M, Meyer CA, Mietto P, Paratte G, Lovis C, Thüring B. 2010. Comparative analysis of Late Jurassic sauropod trackways from the Jura mountains (NW Switzerland) and the central High Atlas mountains (Morocco): implications for sauropod ichnotaxonomy. Hist Biol. 22(1–3):109–133. doi:10.1080/08912960903503345.
   Web of Science ®Google Scholar
• Matsukawa M, Lockley MG, Hunt AP. 1999. Three age groups of ornithopods inferred from footprints in the mid Cretaceous Dakota Group, eastern Colorado, North America. Palaeogeogr Palaeoclimatol Palaeogeogr. 147:39–51. doi:10.1016/S0031-0182(98)00154-0.
   Web of Science ®Google Scholar
• Matsukawa M, Shibata K, Kukihara R, Koarai K, Lockley MG. 2005. Review of Japanese dinosaur track localities, implications for ichno-taxonomy, paleogeography and stratigraphic correlation. Ichnos. 12(3):201–222. doi:10.1080/10420940591009231.
   Google Scholar
• Olsen PE, Smith JB, McDonald NG. 1998. Type material of the type species of the classic theropod footprint genera Eubrontes, Anchisauripus and Grallator (Early Jurassic, Hartford and Deerfield basins, Connecticut and Massachusetts, U.S.A.). J Vertebr Paleontol. 18(3):586‒601. doi:10.1080/02724634.1998.10011086.
   Web of Science ®Google Scholar
• Pascual-Arribas C, Hernández-Medrano N, Latorre-Macarrón P, Sanz-Pérez E. 2009. El icnogénero Iguanodontipus en el yacimiento de “Las Cuestas I” (Santa Cruz de Yanguas, Soria, España). Studia Geologica Salmanticensia. 45:105–128.
   Google Scholar
• Santos VF, Moratalla JJ, Royo-Torres R. 2009. New sauropod trackways from the Middle Jurassic of Portugal. Acta Palaeontol Pol. 54(3):409–422. doi:10.4202/app.2008.0049.
   Web of Science ®Google Scholar
• Shibata M, You HL, Azuma Y. 2017. Recent advance in dinosaur research in Japan: comparison of Early Cretaceous dinosaur faunas in East and Southeast Asia. Fossils Palaeontol Soc Jpn. 101:23‒41.
   Google Scholar
• Suarez C, Trieschmann B, Suarez M, You H, Li D 2014. Mean annual precipitation and composition interpreted from vertebrate bioapatite and bulk organic carbon form the Early Cretaceous Hekou Group, Gansu Province, China. 2014 GSA Annual Meeting in Vancouver, British Columbia, Paper No. 251‒11.
   Google Scholar
• Tang YH, Dai S, Huang YB, Zhu Q, Fang XM, Hu HF, Liu JW, Kong L, Zhao J, Liu X. 2008. The early Cretaceous tectonic uplift of Qilian mountains, evidence from the sedimentary facies and susceptibility of rocks of the Hekou Group, Lanzhou-Minhe basin. Earth Sci Front. 15(2):261–271.
   Google Scholar
• Thulborn RA. 1990. Dinosaur tracks. London: Chapman and Hall; p. 410.
   Google Scholar
• Xing LD, Buckley LG, Lockley MG, Zhang JP, Marty D, Wang Y, Li JJ, McCrea RT, Peng C. 2016a. A new bird track, Koreanaornis lii ichnosp. nov., from the Lower Cretaceous Hekou Group in the Lanzhou-Minhe Basin, Gansu, Northwest China, and implications for Early Cretaceous avian diversity. Cretaceous Res. 66:141–154. doi:10.1016/j.cretres.2016.06.005.
   Web of Science ®Google Scholar
• Xing LD, Li DQ, Harris JD, Bell PR, Azuma Y, Fujita M, Lee Y, Currie PJ. 2013. A new deinonychosaurian track from the Lower Cretaceous Hekou Group, Gansu Province, China. Acta Palaeontol Pol. 58(4):723–730.
   Web of Science ®Google Scholar
• Xing LD, Li DQ, Klein H, Lockley MG, Liang QQ, Yang JT, Li LF, You WZ. 2019. Lower Cretaceous turtle tracks from Hekou Group of Northwest China. Cretaceous Res. 99:269–274. doi:10.1016/j.cretres.2019.03.009.
   Web of Science ®Google Scholar
• Xing LD, Li DQ, Lockley MG, Marty D, Zhang JP, Persons WSIV, You HL, Peng C, Kümmell SB. 2015a. Dinosaur natural track casts from the Lower Cretaceous Hekou Group in the Lanzhou–Minhe Basin, Gansu, Northwest China: ichnology track formation, and distribution. Cretaceous Res. 52:194–205. doi:10.1016/j.cretres.2014.10.001.
   Web of Science ®Google Scholar
• Xing LD, Li DQ, Lockley MG, Zhang JP, You HL, Klein H, Marty D, Persons WSIV, Peng C. 2014b. Theropod and sauropod track assemblages from the Lower Cretaceous Hekou Group at Zhongpu, Gansu Province, China. Acta Palaeontol Sin. 53(3):381–391.
   Google Scholar
• Xing LD, Liu YQ, Marty D, Kuang HW, Klein H, Persons WS, Lyu Y. 2017b. Sauropod trackway reflecting an unusual walking pattern from the Early Cretaceous of Shandong Province, China. Ichnos. 24(1):27–36. doi:10.1080/10420940.2015.1126583.
   Web of Science ®Google Scholar
• Xing LD, Lockley MG, Kim KS, Klein H, Matsukawa M, Lim JD, Persons WS, Xu X. 2017a. Mid-Cretaceous dinosaur track assemblage from the Tongfosi Formation of China: comparison with the track assemblage of South Korea. Cretaceous Res. 74:155–164. doi:10.1016/j.cretres.2017.02.019.
   Web of Science ®Google Scholar
• Xing LD, Lockley MG, Marty D, Klein H, Yang G, Zhang JP, Peng GZ, Ye Y, Persons WSIV, Yin XY, et al. 2016c. A diverse saurischian (theropod-sauropod) dominated footprint assemblage from the Lower Cretaceous Jiaguan Formation in the Sichuan Basin, southwestern China: A new ornithischian ichnotaxon, pterosaur tracks and an unusual sauropod walking pattern. Cretaceous Res. 60:176–193. doi:10.1016/j.cretres.2015.12.005.
   Web of Science ®Google Scholar
• Xing LD, Lockley MG, Marty D, Piñuela L, Klein H, Zhang JP, Persons WSIV. 2015b. Re–description of the partially collapsed Early Cretaceous Zhaojue dinosaur tracksite (Sichuan Province, China) by using previously registered video coverage. Cretaceous Res. 52:138–152. doi:10.1016/j.cretres.2014.09.010.
   Web of Science ®Google Scholar
• Xing LD, Lockley MG, Zhang JP. 2016b. Early Cretaceous dinosaur and other tetrapod tracks of southwestern China. Ningbo: Ningbo Publishing House; p. 1–410.
   Google Scholar
• Xing LD, Lockley MG, Zhang JP, Klein H, Persons WSIV, Dai H. 2014c. Diverse sauropod–, theropod–, and ornithopod–track assemblages and a new ichnotaxon Siamopodus xui ichnosp. nov. from the Feitianshan Formation, Lower Cretaceous of Sichuan Province, southwest China. Palaeogeogr Palaeoclimatol Palaeoecol. 414:79–97. doi:10.1016/j.palaeo.2014.08.011.
   Web of Science ®Google Scholar
• Xing LD, Marty D, Wang KB, Lockley MG, Chen SQ, Xu X, Liu YQ, Kuang HW, Zhang JP, Ran H, et al. 2015c. An unusual sauropod turning trackway from the Early Cretaceous of Shandong Province, China. Palaeogeogr Palaeoclimatol Palaeoecol. 437:74–84. doi:10.1016/j.palaeo.2015.07.036.
   Web of Science ®Google Scholar
• Xing LD, Niedźwiedzki G, Lockley MG, Zhang JP, Cai XF, Persons WSIV, Ye Y. 2014a. Asianopodus type footprints from the Hekou Group of Honggu District, Lanzhou City, Gansu, China and the “heel” of large theropod tracks. Palaeoworld. 23:304–313. doi:10.1016/j.palwor.2014.08.003.
   Web of Science ®Google Scholar
• Yang JT, You HL, Li DQ, Kong DL. 2013. First discovery of polacanthine ankylosaur dinosaur in Asia. Vertebrata PalAsiatica. 51(4):265–277.
   Google Scholar
• You HL, Ji Q, Li D. 2005. Lanzhousaurus magnidens gen. et sp. nov. from Gansu Province, China: the largest-toothed herbivorous dinosaur in the world. Geol Bull China. 24:785‒794.
   Google Scholar
• You HL, Li DQ, Zhou LQ, Ji Q. 2006. Huanghetitan liujiaxiaensis. a new sauropod dinosaur from the Lower Cretaceous Hekou Group of Lanzhou Basin, Gansu Province, China. Geol Rev. 52(5):668–674.
   Google Scholar
• You HL, Li DQ, Zhou LQ, Ji Q. 2008. Daxiatitan binglingi: a giant sauropod dinosaur from the Early Cretaceous of China. Gansu Geol. 17(4):1–10.
   Google Scholar
• Young CC. 1948. Fossil crocodiles in China, with notes on dinosaurian remains associated with the Kansu crocodiles. Bull Geol Soc China. 28(3–4):255–288. doi:10.1111/j.1755-6724.1948.mp283-4011.x.
   Google Scholar
• Young CC, Zhao XJ. 1972. Mamenchisaurus hochuanensis sp. nov. Inst Vertebr Paleontol Paleoanthropol Monogr Ser A. 8:1–30.
   Google Scholar
• Yu J, Zhang W, Zhao Q, Song S. 1982. Late Jurassic and early Cretaceous palynological assemblages from Minhe Basin, Qinghai and Gansu provinces. Bull Inst Geol Chin Acad Geol Sci. 5:111–126. ( in Chinese, with English summary).
   Google Scholar
• Zhang HF, Lin QX, Zhang ZY, Gu YS, Cai XF, Yan XQ. 2003. Study on the sedimentary sequence and sedimentary facies of the Early Cretaceous Hekou Group in Lanzhou-Minhe Basin. Geol Sci Technol Inf. 22(4):21–26.
   Google Scholar
• Zhang JP, Li DQ, Li ML, Lockley MG, Bai Z. 2006. Diverse dinosaur, pterosaur and bird-track assemblages from the Hakou Formation, Lower Cretaceous of Gansu Province, northwest China. Cretaceous Res. 27:44–55. doi:10.1016/j.cretres.2005.10.006.
   Web of Science ®Google Scholar