The equisetalean Iberisetum wegeneri gen. nov., sp. nov. from the Upper Pennsylvanian of Portugal

References

• Agardh CH. 1825. Aphorismi botanici. Classes plantarum, pars 1. Berling: Lund.
   Google Scholar
• Barthel M. 2004. Die Rotliegendflora des Thüringer Waldes. Teil 2: Calamiten und Lepidophyten Veröffentlichungen der Naturhistorisches Museum Schleusingen. 19:19–48.
   Google Scholar
• Boardman DR, Iannuzzi R. 2010. Presence of the genus Giridia, sphenophyte, in the Paraná Basin (lower Permian, Rio Bonito Formation). Revista Brasileira De Paleontologia. 13(1):5–12. doi:https://doi.org/10.4072/rbp.2010.1.01.
   Google Scholar
• Boardman DR, Iannuzzi R, Dutra TL. 2016. A new genus of Sphenopsida from the lower permian of the Paraná Basin, Southern Brazil. Rev Palaeobot Palynol. 233:44–55. doi:https://doi.org/10.1016/j.revpalbo.2016.07.004.
   Google Scholar
• Boureau E. 1964. Traité de Paléobotanique III: sphenophyta et Noeggerathiophyta. Paris (France): Masson et Cie Éditeurs.
   Google Scholar
• Brongniart A. 1828. Histoire des vegetaux fossiles, ou recherches botaniques et géologiques sur les vegetaux renfermes dans les diverses couches du globe. Paris (France): Dufour et d’Ocagne.
   Google Scholar
• Clark JW, Puttick MN, Donoghue PCJ. 2019. Origin of horsetails and the role of whole-genome duplication in plant macroevolution. Proc R Soci B. 286:1–10.
   Web of Science ®Google Scholar
• Cleal CJ, Thomas BA. 2010. Botanical nomenclature and plant fossils. Taxon. 59:261–268. doi:https://doi.org/10.1002/tax.591024.
   Web of Science ®Google Scholar
• Cocks LRM, Torsvik TH. 2011. The palaeozoic geography of Laurentia and Western Laurussia: A stable craton with mobile margins. Earth-Sci Rev. 106(1–2):1–51. doi:https://doi.org/10.1016/j.earscirev.2011.01.007.
   Web of Science ®Google Scholar
• Correia P, Bashforth AR, Šimůnek Z, Cleal CJ, Sá AA, Labandeira CC. 2020. The history of herbivory on sphenophytes: a new calamitalean with an insect gall from the upper Pennsylvanian of Portugal and a review of arthropod herbivory on an ancient lineage. Int J Plant Sci. 181(4):387–418. doi:https://doi.org/10.1086/707105.
   Web of Science ®Google Scholar
• Correia P, Murphy JB. 2020. Iberian-Appalachian connection is the missing link between Gondwana and Laurasia that confirms a Wegenerian Pangaea configuration. Sci Rep. 10(1):2498. doi:https://doi.org/10.1038/s41598-020-59461-x.
   PubMed Web of Science ®Google Scholar
• Correia P, Sá AA, Murphy JB, Šimůnek Z, Flores D. 2016. Lesleya Lesquereux from the Pennsylvanian of the Iberian Massif: part of a dryland megaflora from the Variscan orogen, northwestern Portugal. Can J Earth Sci. 53:883–895. doi:https://doi.org/10.1139/cjes-2015-0213.
   Web of Science ®Google Scholar
• Correia P, Šimůnek Z, Sá AA, Flores D. 2018. A new late Pennsylvanian floral assemblage from the Douro Basin, Portugal. Geol J. 53:2507–2531. doi:https://doi.org/10.1002/gj.3086.
   Web of Science ®Google Scholar
• Cúneo R, Escapa I. 2006. The Equisetalean genus Cruciaetheca nov. from the Lower Permian of Patagonia, Argentina. Int J Plant Sci. 167(1):167–177. doi:https://doi.org/10.1086/497652.
   Web of Science ®Google Scholar
• Dijkstra SJ, van Amerom HWJ. 1995. Fossilium Catalogus II: plantae. In: van Amerom HWJ, editor. Acrobulbillites – fuctibus, Vol. 96, Sphenophyta (1). 2. Supplement. Amsterdan: Kugler Publications; p. 1–156.
   Google Scholar
• Domingos LCG, Freire JLS, Gomes da Silva F, Gonçalves F, Pereira E, Ribeiro A. 1983. The structure of the intramontane Upper Carboniferous basins in Portugal. In: Lemos de Sousa MJ, Oliveira JT, editors. The Carboniferous of Portugal (Vol. 29). (Lisboa): Memórias dos Serviços Geológicos de Portugal; p. 187–194.
   Google Scholar
• Doweld AB. 2001. Prosyllabus Tracheophytorum. Tentamen Systematis Plantarum Vascularium (Tracheophyta) GEOS. 80:33–110.
   Google Scholar
• Doweld AB. 2014. (2287) Proposal to conserve the name Asterocalamites against Archaeocalamites and Stigmatocanna (fossil Equisetophyta: asterocalamitopsida). Taxon. 63:442–444.
   Web of Science ®Google Scholar
• Doweld AB. 2017. Proposals to conserve the names Autophyllites against Bryon and A. furcatus against B. crispatum (fossil Sphenophyllophyta: autophyllitales). Taxon. 66(6):1479–1480. (2573–2574). https://doi.org/10.12705/666.24.
   Web of Science ®Google Scholar
• Eagar RMC. 1983. The non-marine bivalve fauna of the Stephanian C of the North Portugal. In: Lemos de Sousa MJ, Oliveira JT, editors. The Carboniferous of the Portugal (Vol. 29). (Lisboa): Memórias dos Serviços Geológicos de Portugal; p. 179–185.
   Google Scholar
• Elgorriaga A, Escapa IH, Bomfleur B, Cúneo R, Ottone EG. 2015. Reconstruction and phylogenetic significance of a new Equisetum Linnaeus Species from the lower Jurassic of Cerro Bayo (Chubut Province, Argentina). Ameghiniana. 52(1):135–152. doi:https://doi.org/10.5710/AMGH.15.09.2014.2758.
   Web of Science ®Google Scholar
• Elgorriaga A, Escapa IH, Rothwell GW, Tomescu AMF, Cúneo NR. 2018. Origin of Equisetum: evolution of horsetails (Equisetales) within the major euphyllophyte clade Sphenopsida. Am J Bot. 105(8):1286–1303. doi:https://doi.org/10.1002/ajb2.1125.
   PubMed Web of Science ®Google Scholar
• Gillespie WH, Clendening JA. 1966. West Virginia plant fossils, I. Dolerotheca and Daubreeia. Proc West Virginia Acad Sci. 38:159–168.
   Google Scholar
• Grauvogel-Stamm L 1978. La flore du Grès à Voltzia (Buntsandstein supérieur) des Vosges du nord (France). Morphologie, anatomie, interprétations phylogénique et paléogéographique [monographie]. Sciences Géologiques, Université Louis Pasteur de Strasbourg, Institut de Géologie, Memoir 50:1–225. https://books.google.fr/books?id=esOXdvbRiMoC&printsec=frontcover&hl=pt-PT&source=gbs_ge_summary_r&cad=0#v=onepage&q&f=false
   Google Scholar
• Hilton J, Cleal CJ. 2007. The relationship between Euramerican and Cathaysian tropical floras in the late Palaeozoic: palaeobiogeographical and palaeogeographical implications. Earth-Sci Rev. 85(3–4):85–116. doi:https://doi.org/10.1016/j.earscirev.2007.07.003.
   Google Scholar
• Hirmer M. 1927. Handbuch der Paläobotanik, 1, Thallophyta, Bryophyta, Pteridophyta. München (Berlin): Druck und Verlag von R. Oldenbourg.
   Google Scholar
• Jongmans WJ. 1911. Anleitung zur Bestimmung der Karbonpflanzen West-Europas. Thallophytae, Equisetales, Sphenophyllales. 3: 1–482. Mededelingen Rijksopsporing Delfstoffen.
   Google Scholar
• Kelber K-P. 2015. 5. Die Makroflora des Lettenkeupers. In: Hagdorn H, Schoch R, Schweigert G, editors. Der Lettenkeuper – ein Fenster in die Zeit vor den Dinosauriern. Palaeodiversity Supplement. Staatliches Museum für Naturkunde. Germany: Stuttgart; p. 51–100.
   Google Scholar
• Kelber K-P, van Konijnenburg-van Cittert JHA. 1998. Equisetites arenaceus from the Upper Triassic of Germany with evidence for reproductive strategies. Rev Palaeobot Palynol. 100(1–2):1–26. doi:https://doi.org/10.1016/S0034-6667(97)00061-4.
   Web of Science ®Google Scholar
• Langiaux J. 1984. Flores et faunes des formations supérieures du Stéphanien de Blanzy-Montceau (Massif Central francais), Stratigraphie et Paléoécologie. Revue périodique de «La Physiophile», Société Etudes Sciences Naturelles et Historiques de Montceau-les-Mines. 60(Supplement au No. 100):1–270.
   Google Scholar
• Lemos de Sousa MJ, Wagner RH. 1983. General description of the terrestrial Carboniferous basins in Portugal and history of investigations. In: Lemos de Sousa MJ, Oliveira JT, editors. The Carboniferous of Portugal (Vol. 29). (Lisboa): Memórias dos Serviços Geológicos de Portugal; p. 117–126.
   Google Scholar
• Linnaeus C. 1753. Species Plantarum. Stockholm (Sweden): Imprensis Laurentii Salvii.
   Google Scholar
• Mcloughlin S, Larsson K, Lindström S. 2005. Permian plant macrofossils from Fossilryggen, Vestfjella, Dronning Maud Land. Antarct Sci. 17(1):73–86. doi:https://doi.org/10.1017/S0954102005002464.
   Google Scholar
• McNeill J, Barrie FR, Buck WR, Demoulin V, Greuter W, Hawksworth DL, Herendeen PS, Knapp S, Marhold K, Prado J, et al. 2011. International Code of Nomenclature for algae, fungi, and plants (Melbourne, Australia, 2011). Koeltz: Oberreifenberg, Germany.
   Google Scholar
• Meyen SV 1967. New data on the relationship between Angara and Gondwana late Paleozoic floras. Gondwana Stratigraphy, IUGS 1st Gondwana Symposium, Buenos Aires (Argentina) 1967, UNESCO Earth Sciences 2, p. 144–152.
   Google Scholar
• Meyen SV. 1969. The Angara members of the Gondwana genus Barakaria and its systematical position. Argumenta Palaeobotanica. 3:1–14.
   Google Scholar
• Meyen SV. 1971. Phyllotheca-like plants from the Upper Palaeozoic flora of Angaraland. Palaeontographica. 133B:1–33.
   Google Scholar
• Meyen SV. 1982. The Carboniferous and Permian floras of Angaraland (A synthesis). Biol Mem. 7:1–110.
   Google Scholar
• Meyen SV. 1987. Fundamentals of Palaeobotany. New York (USA): Chapman & Hall.
   Google Scholar
• Morris JL, Puttick MN, Clark JW, Edwards D, Kenrick P, Pressel S, Wellman CH, Yang Z, Schneider H, Donoghue PCJ. 2018. The timescale of early land plant evolution. Proc Natl Acad Sci U S A. 115(10):E2274–E2283. doi:https://doi.org/10.1073/pnas.1719588115.
   PubMed Web of Science ®Google Scholar
• Naugolnykh SV. 2002. Paracalamitina striata—a newly reconstructed equisetophyte from the Permian of Angaraland. J Paleontol. 76(2):377–385. doi:https://doi.org/10.1017/S0022336000041755.
   Google Scholar
• Naugolnykh SV. 2004. On some aberrations of extant horsetails (Equisetum L.) and the origin of the family Equisetaceae. Paleontol J. 38:335–342.
   Web of Science ®Google Scholar
• Naugolnykh SV. 2009. A new fertile Neocalamites from the Upper Permian of Russia and equisetophyte evolution. Geobios. 42(4):513–523. doi:https://doi.org/10.1016/j.geobios.2008.12.003.
   Web of Science ®Google Scholar
• Němejc F. 1963. Paleobotanika, Vol. II. Systematická část: rostliny mechovité, psilofytové a kapraďorosty. Praha: Nakladatelství Československé Akademie Věd.
   Google Scholar
• Novikoff A, Barabasz-Krasny B 2015. Modern plant systematics. General issues. Liga-Press, Lviv. [in Ukrainian]
   Google Scholar
• Oshurkova MV. 1996. Paleoecological parallelism between the Angaran and Euramerican phytogeographic provinces. Rev Palaeobot Palynol. 90(1–2):99–111. doi:https://doi.org/10.1016/0034-6667(95)00026-7.
   Google Scholar
• Pfefferkorn HW, Wang J. 2016. Paleoecology of Noeggerathiales, an enigmatic, extinct plant group of Carboniferous and Permian times. Palaeogeogr Palaeoclimatol Palaeoecol. 448:141–150. doi:https://doi.org/10.1016/j.palaeo.2015.11.022.
   Google Scholar
• Pinto de Jesus A. 2001. Génese e Evolução da Bacia Carbonífera do Douro (Estefaniano C inferior, NW de Portugal); Um Modelo [unpublished Ph.D. dissertation]. University of Porto, Portugal.
   Google Scholar
• Pinto de Jesus A. 2003. Evolução sedimentar e tectónica da Bacia Carbonífera do Douro (Estefaniano C inferior, NW de Portugal). Caderno Lab Xeolóxico Laxe Coruña. 28:107–125.
   Google Scholar
• Pott C, Kerp H, Krings M. 2008. Sphenophytes from the Carnian (Upper Triassic) of Lunz am See (Lower Austria). Jahrbuch der Geologischen Bundesanstalt. 148:183–199.
   Google Scholar
• Prevec R, Gastaldo RA, Neveling J, Samuel BR, Looy CV. 2010. An autochthonous glossopterid flora with latest Permian palynomorphs and its depositional setting in the Dicynodon Assemblage Zone of the southern Karoo Basin, South Africa. Palaeogeogr Palaeoclimatol Palaeoecol. 292(3–4):391–408. doi:https://doi.org/10.1016/j.palaeo.2010.03.052.
   Web of Science ®Google Scholar
• Purkyňová E. 1970. Die Unternamurflora des Beckens von Horní Slezsko (ČSSR). Paläont Abh B, Berlin. 3(2):131–268.
   Google Scholar
• Radczenko GP. 1955. Index-fossil of Upper Paleozoic flora of Sayan-Altai region. Atlas Rukovod Form Iskop Fauny I Flory Zap Sibiri. 2:42–153.
   Google Scholar
• Radczenko GP 1956. Familles et genres nouveaux de plantes. Matériel de Paléontologie, n.s., série 12, V.S.E.G.E.I.
   Google Scholar
• Radczenko GP 1962. Vazhnejshie pozdnepermskie rastenija Altae-Sajanskoj gornoj oblasti. [Most important Late Permian plants of Altaj-Sajan mountain region] Trudy Vsesoyuzn. Nauchno-Issled. Geological Institute. Novaja Series: 62:39–77. [in Russian]
   Google Scholar
• Rigby JF. 1970. The Notocalamitaceae, a new family of upper Palaeozoic equisetaleans. Palaeobotanist. 19:161–163.
   Google Scholar
• Roesler GA, Iannuzzi R. 2012. Nova espécie de Sphenophyta no Eopermiano do Rio Grande do Sul (Grupo Itararé, Bacia do Paraná). Revista Brasileira De Paleontologia. 15(2):141–152. doi:https://doi.org/10.4072/rbp.2012.2.03.
   Google Scholar
• Rozefelds AC, Dettmann ME, Milroy AK, Hammond A, Clifford HT, Ekins M. 2019. The unexpected, recent history of horsetails in Australia. Aust Syst Bot. 32:255–268.
   Web of Science ®Google Scholar
• Shute CH, Cleal CJ. 1996. Palaeobotany in museums. Geolo Curator. 4:553–559.
   Google Scholar
• Srivastava AK, Agnihotri D. 2010. Dilemma of late Palaeozoic mixed floras in Gondwana. Palaeogeogr Palaeoclimatol Palaeoecol. 298(1–2):54–69. doi:https://doi.org/10.1016/j.palaeo.2010.05.028.
   Web of Science ®Google Scholar
• Sunderlin D. 2010. Evidence for a western extension of the Angaran phytogeographic province in the Early Permian. Int J Coal Geol. 83(2–3):266–275. doi:https://doi.org/10.1016/j.coal.2009.07.010.
   Google Scholar
• Taylor TN, Taylor EL, Krings M. 2009. Paleobotany: the biology and evolution of fossil plants. Amsterdam: Academic Press.
   Google Scholar
• Tenchov YG. 1977. Flora und Biostratigraphie des Oberkarbons im Svoge-becken (VR Burgarien). Schriftenreihe für geologische Wissenschaften, Academie, Berlin. 7:1–163.
   Google Scholar
• Thomas BA, Cleal CJ. 2020. The nomenclature of fossil-taxa representing different preservational states: lepidodendron as a case-study. Taxon. 59(1):261–268.
   Google Scholar
• Townrow JA. 1955. On some species of Phyllotheca. J Proc R Soc N S W. 83:39–63.
   Google Scholar
• Trapl S. 1927. O listových útvarech čel. Calamariaceae (r. Asterophyllites a Annularia). Sborník Prírodovedecký. 4:1–40.
   Google Scholar
• Uhl D, Raisch M. 2009. Die fossile Schachtelhalmbeblätterung Calamariophyllum im Rotliegend des Saar-Nahe Beckens (SW-Deutschland)–Fundmitteilung. Mitteilungen der POLLICHIA. 94:1–4.
   Google Scholar
• Vaez-Javadi F. 2011. Middle Jurassic flora from the Dansirit formation of the Shemshak Group, Alborz, north Iran. Alcheringa. 35(1):77–102. doi:https://doi.org/10.1080/03115518.2010.489424.
   Web of Science ®Google Scholar
• Vai GB. 2003. Development of the palaeogeography of Pangaea from late Carboniferous to Early Permian. Palaeogeogr Palaeoclimatol Palaeoecol. 196(1–2):125–155. doi:https://doi.org/10.1016/S0031-0182(03)00316-X.
   Google Scholar
• van Amerom HWJ, Gillespie WH, Wagner RH. 2003. Daubreeia pateraeformis (Germar) Zeiller, a Stephanian sphenopsid from western Europe and North America. Revista Española de Paleontología. 18:159–170.
   Google Scholar
• van Amerom HWJ, Kabon H. 2001. Neue Daubreeia-Funde in Kärnten (Ö sterreich). Wulfenia. 8:47–59.
   Google Scholar
• Verbitskaya NG, Radchenko GP. 1968. New Siberian arthropods. In: Zanina IE, Kiparisova LD, Markovsky BP, Lokrovskiy IM, Miklouho-Maclay KV, Radchenko GP, editors. New species of ancient plants and invertebrates of the USSR. Publishing House Nedra, Moscow, Russia [in Russian]; p. 13–18.
   Google Scholar
• von Berchtold FW, Presl JS. 1820. O Přirozenosti Rostlin. Krause, Prague, Czech Republic, Krala Wiljma Endersa. https://en.m.wikipedia.org/wiki/O_P%C5%99irozenosti_Rostlin#CITEREFBerchtoldPresl1820
   Google Scholar
• von Sternberg KM 1820–1838. Versuch einer geognostisch-botanischen Darstellung der Flora der Vorwelt. C.E. Brenck’s widow, Leipzig (tent). XLII pp. (Vol. I)/J. Spurny, Prague (Vol. II).
   Google Scholar
• Wagner RH. 1983. The palaeogeographical and age relationships of the Portuguese Carboniferous floras with those of other parts of the Western Iberian Peninsula. In: Lemos de Sousa MJ, Oliveira JT, editors. The Carboniferous of Portugal (Vol. 29). Lisboa: Memórias dos Serviços Geológicos de Portugal; p. 153–177.
   Google Scholar
• Wagner RH. 2004. Climatic changes as mirrored by Carboniferous and Permian floral distributions. Monografías del Jardín Botánico de Córdoba. 39:29–39.
   Google Scholar
• Wagner RH, Lemos de Sousa MJ. 1983. The Carboniferous Megafloras of Portugal - A revision of identifications and discussion of stratigraphic ages. In: Lemos de Sousa MJ, Oliveira JT, editors. The Carboniferous of Portugal (Vol. 29). (Lisboa): Memórias dos Serviços Geológicos de Portugal; p. 127–152.
   Google Scholar
• Walton J. 1936. On the factors which influence the external form of fossil plants; with descriptions of the foliage of some species of the Palaeozoic equisetalean genus Annularia Sternberg. Philosl Trans R Soci London, Ser B–Biol Sci. 226:219–237.
   Google Scholar
• Weber R. 2005. Equisetites aequecaliginosus sp. nov., ein Riesenschachtelhalm aus der spättriassischen Formation Santa Clara, Sonora, Mexiko [Equisetites aequecaliginosus sp. nov., a tall horsetail from the Late Triassic Santa Clara Formation, Sonora, Mexico]. Revue de Paléobiologie. 24:331–364.
   Google Scholar
• Weber R. 2008. Homomorfismo en Equisetaceae del Triásico: asinisetum gen. nov., Equisetites aequecaliginosus Weber y conos asociados de Sonora, México. In: Weber R, editor. Plantas triásicas y jurásicas de México (Vol. 115). Boletín: Universidad Nacional Autónoma de México, Instituto de Geología; p. 1–83.
   Google Scholar
• Wnuk C. 1996. The development of floristic provinciality during the Middle and Late Palaeozoic. Review of Palaeobotany and Palynology. 90(1–2):5–40. doi:https://doi.org/10.1016/0034-6667(95)00022-4.
   Google Scholar
• Yamada T, Uemura K. 2008. The plant fossils from the Kaizara Formation (Callovian, Jurassic) of the Tetori Group in the Izumi district, Fukui Prefecture, Central Japan. Paleontol Res. 12(1):1–17. doi:https://doi.org/10.2517/1342-8144(2008)12[1:TPFFTK]2.0.CO;2.
   Google Scholar
• Zalessky MD. 1934. Observations sur les végétaux permiens du bassin de la Petchora, I. Bulletin of Academy of Sciences, URSS, Mathematics and Natural Sciences. 2–3:241–290.
   Google Scholar
• Zalessky MD. 1939. Végétaux Permiens du Bardien de l’Oural. Probl Paleontol. 5:329–374.
   Google Scholar
• Zeiller R. 1888/1890. Études sur le terrain houiller de Commentry. Flore fossile. Première Partie, Texte (1890): 366 pp.; Atlas ( 1888): 42 pls. . Saint-Étienne, France.
   Google Scholar