The Chalicotheriidae (Mammalia, Perissodactyla) from the upper Miocene Nakali Formation, Kenya

References

• Aguirre E, Guérin C. 1974. Première découverte d’un Iranotheriinae (Mammalia, Perissodactyla, Rhinocerotidae) en Afrique: kenyatherium bishopi nov. gen. sp. de la formation vallésienne (Miocene supérieur) de Nakali (Kenya) [First discovery of an Iranotheriinae (Mammalia, Perissodactyla, Rhinocerotidae) in Africa: kenyatherium bishopi nov. gen. sp. of the Vallesian formation (Upper Miocene) of Nakali (Kenya)]. Estud Geológ. 30:229–233.
   Google Scholar
• Aguirre E, Leakey P. 1974. Nakali, nueva fauna de Hipparion del Rift Valley de Kenya [Nakali, New Hipparion fauna in the Rift Valley of Kenya]. Estud Geológ. 30:219–227.
   Google Scholar
• Borissiak AA. 1946. [A new Chalicothere from the Tertiary of Kazakhstan]. Trudy Paleontologicheskovo Instituta, Akademia Nayk USSR. 13:1–134.(in Russian)
   Google Scholar
• Butler PM. 1965. Fossil mammals of Africa No. 18: east African Miocene and Pleistocene chalicohteres. Bull Brit Mus (Nat Hist) Geol. 24:165–237.
   Google Scholar
• Chen S. 2008. A review on Chinese Neogene chalicotheres. In: Dong W, editor. Proceedings of the eleventh annual meeting of the Chinese society of vertebrate paleontology. Beijing: China Ocean Press; p. 31–41.
   Google Scholar
• Coombs MC. 1975. Sexual dimorphism in chalicotheres (Mammalia, Perissodactyla). System Zool. 24(1):55–62. doi:https://doi.org/10.2307/2412697.
   Google Scholar
• Coombs MC. 1978. Reevaluation of early Miocene North American Moropus (Perissodactyla, Chalicotheriidae, Schizotheriinae). Bull Carnegie Mus Nat Hist. 4:1–62.
   Google Scholar
• Coombs MC. 1979. Tylocephalonyx, a new genus of North American dome-skulled chalicotheres (Mammalia, Perissodactyla). Bull Am Mus Nat Hist. 164:1–64.
   Google Scholar
• Coombs MC. 1989. Interrelationships and diversity in the Chalicotheriidae. In: Prothero DR, Schoch RM, editors. The evolution of perissodactyls. Oxford/New York: Clarendon/Oxford University Press; p. 438–457.
   Google Scholar
• Coombs MC. 1998. Chalicotherioidea. In: Janis CM, Scott KM, Jacobs LL, editors. Evolution of tertiary mammals of North America vol. 1: terrestrial carnivores, ungulates and ungulate-like mammals. Cambridge: Cambridge University Press; p. 560–568.
   Google Scholar
• Coombs MC. 2009. The chalicothere Metaschizotherium bavaricum (Perissodactyla, Chalicotheriidae, Schizotheriinae) from the Miocene (MN5) Lagerstätte of Sandelzhausen (Germany): description, comparison, and paleoecological significance. Paläontol Zeitschrift. 83(1):85–129. doi:https://doi.org/10.1007/s12542-009-0004-x.
   Google Scholar
• Coombs MC, Cote S. 2010. Chalicotheriidae. In: Werdelin L, Sanders WJ, editors. Cenozoic mammals of Africa. Berkeley: University of California Press; p. 659–667.
   Google Scholar
• Coombs MC, Hunt RM Jr., Stepleton E, Albright III LB, Fremd TJ. 2001. Stratigraphy, chronology, biogeography, and taxonomy of early Miocene small chalicotheres in North America. J Vert Paleontol. 21(3):607–620. doi:https://doi.org/10.1671/0272-4634(2001)021[0607:SCBATO]2.0.CO;2.
   Web of Science ®Google Scholar
• Cote SM. 2004. Origins of the African hominoids: an assessment of the palaeobiogeographical evidence. Compt Rendus Palevol. 3(4):323–340. doi:https://doi.org/10.1016/j.crpv.2004.03.006.
   Web of Science ®Google Scholar
• de Bonis L. 1995. Le Garouillas et les sites contemporains (Oligocene, MP 25) des phosphorites du Quercy (Lot, Tarn-et-Garonne, France) et leurs faunes de vertebres. Palaeontogr Abteilung A. 236:191–204.
   Google Scholar
• Dietrich WO. 1942. Ältestquartäre Säugetiere aus der südlichen Serengeti, Deutsch-Ostafrika. Palaeontograp Abt A. 94A:43–130.
   Google Scholar
• Fahlke JM, Coombs MC. 2009. Dentition and first postcranial description of Metaschizotherium fraasi Koenigswald, 1932 (Perissodactyla: chalicotheriidae) and its occurrence on a karstic plateau – new insights into schizotheriine morphology, relationships, and ecology. Palaeontograp Abt A. 290(1–3):65–129. doi:https://doi.org/10.1127/pala/290/2009/65.
   Google Scholar
• Fischer MS. 1986. Die Stellung der Schliefer (Hyracoidea) im phylogenetischen System der Eutheria. Zugleich ein Beitrag zur Anpassungsgeschichte der Procaviidae. Cour Forschung Sencken. 84:1–132.
   Google Scholar
• Forster-Cooper C. 1920. Chalicotheroidea from Baluchistan. Procee Zool Soc London. 3:357–366.
   Google Scholar
• Gaudry A. 1862-67. Animaux fossiles et géologie de l’Attique. Paris: Savy. [The chapter on Ancylotherium was published in 1863.].
   Google Scholar
• Gaudry A, Lartet E. 1856. Résultats des recherches paléontologiques entreprises dans l’Attique sous les auspices de l’Académie. C R Acad Sci Paris. 43:271–274.
   Google Scholar
• George M. 1950. A chalicothere from the Limeworks Quarry of the Makapan Valley, Potgietersrust District. South Afr J Sci. 46:241–242.
   Google Scholar
• Geraads D, Alemseged Z, Bellon H. 2002. The late Miocene mammalian fauna of Chorora, Awash basin, Ethiopia: systematics, biochronology and the 40K-40Ar ages of the associated volcanics. Tert Res. 21:113–122.
   Google Scholar
• Geraads D, Mcrosson M, Benefit B. 2012. A New rhinoceros, Victoriaceros kenyensis gen. et sp. nov., and other Perissodactyla from the Middle Miocene of Maboko, Kenya. J Mammalian Evol. 19(1):57–75. doi:https://doi.org/10.1007/s10914-011-9183-9.
   Web of Science ®Google Scholar
• Geraads D, Spassov N, Kovchev D. 2001. New Chalicotheriidae (Perissodactyla,Mammalia) from the late Miocene of Bulgaria. J Vert Paleontol. 21(3):596–606. doi:https://doi.org/10.1671/0272-4634(2001)021[0596:NCPMFT]2.0.CO;2.
   Web of Science ®Google Scholar
• Geraads D, Tsoukala E, Spassov N. 2007. A skull of Ancylotherium (Chalicotheriidae, Mammalia) from the late Miocene of Thermopigi (Serres, N. Greece) and the relationships of the genus. J Vert Paleontol. 27(2):461–466. doi:https://doi.org/10.1671/0272-4634(2007)27[461:ASOACM]2.0.CO;2.
   Web of Science ®Google Scholar
• Gill T. 1872. Arrangement of the families of mammals with analytical tables. Smithsonian Miscellan Collect. 11:1–98.
   Google Scholar
• Guérin C, Pickford M. 2003. Ougandatherium napakense nov. gen. nov. sp., le plus ancien Rhinocerotidae Iranotheriinae d’Afrique. Annal De Paléontol. 89(1):1–35. doi:https://doi.org/10.1016/S0753-3969(03)00004-1.
   Google Scholar
• Guérin C, Pickford M. 2005. Ancylotherium cheboitense nov. sp., nouveau Chalicotheriidae (Mammalia, Perissodactyla) du Miocène supérieur des Tugen Hills (Kénya). Compt Rendus Palevol. 4(3):225–234. doi:https://doi.org/10.1016/j.crpv.2004.10.005.
   Web of Science ®Google Scholar
• Handa N, Nakatsukasa M, Kunimatsu Y, Nakaya H. 2017a. A new Elasmotheriini (Perissodactyla, Rhinocerotidae) from the upper Miocene of Samburu Hills and Nakali, northern Kenya. Geobios. 50(3):197–209. doi:https://doi.org/10.1016/j.geobios.2017.04.002.
   Web of Science ®Google Scholar
• Handa N, Nakatsukasa M, Kunimatsu Y, Nakaya H. 2017b. Additional specimens of Diceros (Perissodactyla, Rhinocerotidae) from the Upper Miocene Nakali Formation in Nakali, central Kenya. Histolog Biolol:12. doi:https://doi.org/10.1080/08912963.2017.1362560.
   Google Scholar
• Handa N, Nakatsukasa M, Kunimatsu Y, Nakaya H. 2018. Brachypotherium (Perissodactyla, Rhinocerotidae) from the late Miocene of Samburu Hills, Kenya. Geobios. 51(5):391–399. doi:https://doi.org/10.1016/j.geobios.2018.08.003.
   Web of Science ®Google Scholar
• Handa N, Nakatsukasa M, Kunimatsu Y, Tsubamoto T, Nakaya H. 2015. New specimens of Chilotheridium (Perissodactyla, Rhinocerotidae) from the Upper Miocene Namurungule and Nakali Formations, Northern Kenya. Paleontol Res. 19(3):181–194. doi:https://doi.org/10.2517/2014PR035.
   Web of Science ®Google Scholar
• Heissig K. 1999. Family Chalicotheriidae. In: Rössner GE, Heissig K, editors. The Miocene Land Mammals of Europe. Munich: Pfeil; p. 189–192.
   Google Scholar
• Heissig K. 2012. Les Rhinocerotidae (Perissodactyla) de Sansan. In: Peigné S, Sen S, editors. Mammifè res de Sansan. Mémoires du Muséum National d’Histoire Naturelle 203. Paris: Muséum National d’Histoire Naturelle; p. 317–485.
   Google Scholar
• Heissig K, Fejfar O. 2013. Die Säugetiere aus dem Untermiozän des Chomutov Beckens - I. Chalicotheriidae (Mammalia, Perissodactyla). Acta Musei Nat Pragae Ser B Hist Nat. 69:7–65. doi:https://doi.org/10.14446/AMNP.2013.007.
   Google Scholar
• Holland WJ, Peterson OA. 1914. The osteology of the Chalicotheroidea with special reference to a mounted skeleton of Moropus elatus Marsh, now installed in the Carnegie Museum. Mem Carnegie Mus. 3:89–406.
   Google Scholar
• Katoh S, Beyene Y, Itaya T, Hyodo H, Hyodo M, Yagi K, Gouzu C, Woldegabriel G, Hart WK, Ambrose SH, et al. 2016. New geological and palaeontological age constraint for the gorilla-human lineage split. Nature. 530(7589):215–218. doi:https://doi.org/10.1038/nature16510.
   PubMed Web of Science ®Google Scholar
• Kingston JD, Jacobs BF, Hill A, Deino A. 2002. Stratigraphy, age and environments of the late Miocene Mpesida Beds, Tugen Hills, Kenya. J Human Evol. 42(1–2):95–116. doi:https://doi.org/10.1006/jhev.2001.0503.
   PubMed Web of Science ®Google Scholar
• Koufos GD. 2012. New material of Chalicotheriidae (Perissodactyla, Mammalia) from the Late Miocene of Axios Valley, Macedonia (Greece) with the description of a new species. Annal De Paléontol. 98(3):203–224. doi:https://doi.org/10.1016/j.annpal.2012.06.002.
   Web of Science ®Google Scholar
• Koufos GD, Kostopolous DS. 2016. Palaeontology of the upper Miocene vertebrate localities of Nikiti (Chalkidiki Peninsula, Macedonia, Greece), Chalicotheriidae. Giobios. 49(1–2):75–83. doi:https://doi.org/10.1016/j.geobios.2016.01.014.
   Web of Science ®Google Scholar
• Kunimatsu Y, Nakatsukasa M, Sakai T, Saneyoshi M, Sawada Y, Nakaya H. 2017a. A newly discovered galagid fossil from Nakali, an early Late Miocene locality of East Africa. J Human Evol. 105:123–126. doi:https://doi.org/10.1016/j.jhevol.2017.02.003.
   PubMed Web of Science ®Google Scholar
• Kunimatsu Y, Nakatsukasa M, Sawada Y, Sakai T, Hyodo M, Hyodo H, Itaya T, Nakaya H, Saegusa H, Mazurire A, et al. 2007. A new Late Miocene great ape from Kenya and its implications for the origins of African great apes and humans. PNAS. 104(49):19220–19225. doi:https://doi.org/10.1073/pnas.0706190104.
   PubMed Web of Science ®Google Scholar
• Kunimatsu Y, Nakatsukasa M, Sawada Y, Sakai T, Saneyoshi M, Nakaya H, Yamamoto A, Mbua E. 2016. A second hominoid species in the early Late Miocene fauna of Nakali (Kenya). Anthropol Sci. 124(2):75–83. doi:https://doi.org/10.1537/ase.160331.
   Web of Science ®Google Scholar
• Kunimatsu Y, Sawada Y, Sakai T, Saaneyoshi M, Nakaya H, Yamamoto A, Nakatsukasa M. 2017b. The latest occurrence of the nyanzapithecines from the early Late Miocene Nakali Formation in Kenya, East Africa. Anthropol Sci. 125(2):45–51. doi:https://doi.org/10.1537/ase.170126.
   Web of Science ®Google Scholar
• Leidy J 1873. Contributions to the Extinct Vertebrate Fauna of the Western Territories. Report of the United States Geological Survey of the Territories Volume 1 (Fossil Vertebrates). Washington (D.C.): Government Printing Office
   Google Scholar
• Marsh OC. 1877. Notice of some new vertebrate fossils. Am J Sci. 14(81):249–256. doi:https://doi.org/10.2475/ajs.s3-14.81.249.
   Google Scholar
• Mein P, Ginsburg L. 2002. Sur l’âge relatif des différents karstiques miocènes de La Grive-Saint-Alban (Isère). Cahiers scientifiques, Mus Hist natur. Lyon. 2:7–47.
   Google Scholar
• Morales J, Nieto M, Köhler M, Moyà Solà S. 1999. Large mammals from the Vallesian of Spain. In: Agusti J, Rook L, Andrews P, editors. Hominoid evolution and climatic change in Europe: the evolution of the neogene terrestrial ecosystems in Europe. Cambridge: Cambridge University Press; p. 113–126.
   Google Scholar
• Morales J, Pickford M. 2006. A large percrocutid carnivore from the Late Miocene (ca. 10-9 Ma) of Nakali, Kenya. Annal De Paléontol. 92(4):359–366. doi:https://doi.org/10.1016/j.annpal.2006.07.004.
   Google Scholar
• Nakatsukasa M, Mbua E, Sawada Y, Sakai T, Nakaya H, Yano W, Kunimatsu Y. 2010. Earliest colobine skeletons from Nakali, Kenya. Am J Physic Anthropol. 143(3):365–382. doi:https://doi.org/10.1002/ajpa.21327.
   PubMed Web of Science ®Google Scholar
• Nakaya H. 1994. Faunal change of late Miocene Africa and Eurasia: mammalian fauna from the Namurungule Formation, Samburu Hills, northern Kenya. Afr Study Monogr Suppl. 20:1–112.
   Google Scholar
• Nakaya H, Onodera M, Yamada E, Kunimatsu Y, Nakatsukasa M, Sakai T. 2012. Mammalian fauna and its paleoenvironments of the Late Miocene Samburu Hills and Nakali, Rift Valley, Kenya. Abst Japan Geosci Union Meet. 2012:BPT27 02.
   Google Scholar
• Nakaya H, Pickford M, Nakano Y, Ishida H. 1984. The late Miocene large mammal fauna from the Namurungule Formation, Samburu Hills, northern Kenya. Afr Study Monogr Suppl. 2:87–131.
   Google Scholar
• Onodera M, Kunimatsu Y, Nakatsukasa M, Yamada E, Nakaya H 2011. Late Miocene rodent fossils from the Nakali Formation, Kenya and its paleoenvironment. Abstracts with Programs, The 160th Regular Meeting, The Palaeontological Society of Japan, Kochi, Japan, p. 19. (in Japanese)
   Google Scholar
• Peterson OA. 1907. Preliminary notes on some American chalicotheres. Am Nat. 41(492):733–752. doi:https://doi.org/10.1086/278876.
   Google Scholar
• Pickford M. 1979. New evidence pertaining to the Miocene Chalicotheriidae (Mammalia, Perissodactyla) of Kenya. Tert Res. 2:83–91.
   Google Scholar
• Pickford M. 1983. On the origins of Hippopotamidae together with descriptions of two species, a new genus and a new subfamily from the Miocene of Kenya. Geobios. 16(2):193–217. doi:https://doi.org/10.1016/S0016-6995(83)80019-9.
   Web of Science ®Google Scholar
• Pickford M. 1986. A revision of the Miocene Suidae and Tayassuidae (Artiodactyla, Mammalia) of Africa. Tert Res Special Paper. 7:1–83.
   Google Scholar
• Pickford M. 1996. Pliohyracoids (Mammalia, Hyracoidea) from the upper Middle Miocene at Berg Aukas, Namibia. Compt rendus de l’Académie des sci Série. 2(322):501–505.
   Google Scholar
• Pilgrim GE. 1910. Notices of new mammalian genera and species from the Tertiaries of India. Records Geolog Survey India. 40:63–71.
   Google Scholar
• Saegusa H, Nakaya H, Kunimatsu Y, Nakatsukasa M, Tsujikawa H 2006. Earliest elephantid remains from Nakali, Kenya. Abstracts with Programs, The 2006 Annual Meeting, The Palaeontological Society of Japan, Shimane, Japan, 19. (in Japanese)
   Google Scholar
• Sakai T, Saneyoshi M, Sawada Y, Nakatsukasa M, Kunimatsu Y, Mbua E. 2013. Early continental rift basin stratigraphy, depositional facies and tectonics in volcaniclastic system: examples from the Miocene successions along the Japan Sea and in the East African Rift Valley (Kenya). In: Itoh Y, editor. Mechanism of Sedimentary Basin Formation—Multidisciplinary Approach on Active Plate Margin. Rijeka: InTech; p. 83–108.
   Google Scholar
• Saraç G, Sen S. 2005. Chalicotheriidae (Mammalia, Perissodactyla) from the late Miocene of Akkaşdağı, Turkey. Geodiversitas. 27:591–600.
   Web of Science ®Google Scholar
• Semprebon GM, Sise PJ, Coombs MC. 2011. Potential bark and fruit browsing as revealed by stereomicrowear analysis of the peculiar clawed herbivores known as chalicotheres (Perissodactyla, Chalicotherioidea). J Mammalian Evol. 18(1):33–55. doi:https://doi.org/10.1007/s10914-010-9149-3.
   Web of Science ®Google Scholar
• Tanabe Y, Onodera M, Nakatsukasa M, Kunimatsu Y, Nakaya H. 2020. A new cane rat (Rodentia, Thryonomyidae) from the Upper Miocene Nakali Formation, northern Kenya. J Geol Soc Japan. 126(4):167–181. doi:https://doi.org/10.5575/geosoc.2020.0002.
   Google Scholar
• Thenius E. 1953. Studien über fossile Vertebraten Griechenlands. III. Das Maxillargebiss von Ancylotherium pentelicum Gaudry und Lartet. Annal géolog des pays hellén. 5:97–106.
   Google Scholar
• Tsubamoto T, Kunimatsu Y, Nakaya H, Sakai T, Saneyoshi M, Mbua E, Nakatsukasa M. 2015. New specimens of a primitive hippopotamus, Kenyapotamus coryndonae, from the Upper Miocene Nakali Formation, Kenya. J Geol Soc Japan. 121(4):153–159. doi:https://doi.org/10.5575/geosoc.2015.0004.
   Google Scholar
• Tsubamoto T, Kunimatsu Y, Sakai T, Saneyoshi M, Shimizu D, Morimoto N, Nakaya H, Handa N, Tanabe Y, Manthi FK, et al. 2020. A new species of Nyanzachoerus (Mammalia, Artiodactyla, Suidae, Tetraconodontinae) from the upper Miocene Nakali Formation, Kenya. Paleontol Res. 24(1):41–63. doi:https://doi.org/10.2517/2019PR004.
   Web of Science ®Google Scholar
• Tsubamoto T, Kunimatsu Y, Sakai T, Saneyoshi M, Shimizu D, Morimoto N, Nakaya H, Nakatsukasa M. 2017. Listriodontine suid and tragulid artiodactyls (Mammalia) from the upper Miocene Nakali Formation, Kenya. Paleontol Res. 21(4):347–357. doi:https://doi.org/10.2517/2016PR034.
   Web of Science ®Google Scholar
• Uno KT, Cerling TE, Harris JM, Kunimatsu Y, Leakey MG, Nakatsukasa M, Nakaya H. 2011. Late Miocene to Pliocene carbon isotope record of differential diet change among East African herbivores. PNAS. 108(16):6509–6514. doi:https://doi.org/10.1073/pnas.1018435108.
   PubMed Web of Science ®Google Scholar
• Von Koenigswald GHR. 1932. Metaschizotherium fraasi n. g. n. sp., ein neuer Chalicotheriide aus dem Obermiocän von Steinheim a. Albuch. Bemerkungen zur Systematik der Chalicotheriiden. Palaeontographica. 8:1–24.
   Google Scholar
• Webb GL. 1965. Notes on some chalicothere remains from Makapansgat. Palaeontol Afr. 9:49–73.
   Google Scholar
• Wood AR, Ridwell NM. 2015. The first Central American chalicothere (Mammalia, Perissodactyla) and the paleobiogeographic implications for small-bodied schizotheriines. J Vert Paleontol. 35(3):5. https://doi.org/10.1080/02724634.2014.923893,e923893.
   Web of Science ®Google Scholar
• Wu W, Chen G. 1976. A new schizotherine genus from the Neogene of Pingliang Gansu. Verte PalAsiat. 14:194–197.
   Google Scholar
• Zapfe H. 1979. Chalicotherium grande (Blainv.) aus der miozänen Spaltenfüllung von Neudorf an der March (Devinská Nová Ves), Tschechoslowakei. Neue-Denkschriften des Naturhistorischen Museums Wien. 2:1–282.
   Google Scholar