Restudy of the original and new materials of Stromatoolithus pinglingensis and discussion on some Spheroolithidae eggs

References

• Agnolin FL, Powell JE, Novas FE, Kundrát M. 2012. New alvarezsaurid (Dinosauria, Theropoda) from uppermost Cretaceous of north-western Patagonia with associated eggs. Cretac Res. 35:33–56. doi:https://doi.org/10.1016/j.cretres.2011.11.014.
   Web of Science ®Google Scholar
• Amo-Sanjuán O, Canudo JI, Cuenca-Bescós G. 2000. First record of elongatoolithid eggshells from the Lower Barremian (Lower Cretaceous) of Europe (Cuesta Corrales 2, Galve Basin, Teruel, Spain). In: Bravo AM, Reyes T, editors. First international symposium on dinosaur eggs and babies (Extended abstracts). Catalonia (Spain): Isona i Conca Dellà; p. 7–14.
   Google Scholar
• Araújo R, Castanhinha R, Martins RMS, Mateus O, Hendrickx C, Beckmann F, Schell N, Alves LC. 2013. Filling the gaps of dinosaur eggshell phylogeny: late Jurassic theropod clutch with embryos from Portugal. Sci Rep. 3(1). doi:https://doi.org/10.1038/srep01924.
   Google Scholar
• Botfalvai G, Csiki-Sava Z, Grigorescu D, Vasile Ş. 2017. Taphonomical and palaeoecological investigation of the Late Cretaceous (Maastrichtian) Tuştea vertebrate assemblage (Romania; Haţeg Basin) - insights into a unique dinosaur nesting locality. Palaeogeogr Palaeoclimatol Palaeoecol. 468:228–262. doi:https://doi.org/10.1016/j.palaeo.2016.12.003.
   Web of Science ®Google Scholar
• Cheng Y-N, Ji Q, Wu XC, Shan H. 2008. Oviraptorosaurian eggs (dinosauria) with embryonic skeletons discovered for the first time in China. Acta Geol Sin-Engl. 82(6):1089–1094. doi:https://doi.org/10.1111/j.1755-6724.2008.tb00708.x.
   Web of Science ®Google Scholar
• Chiappe LM, Coria RA, Dingus L, Jackson F, Chinsamy A, Fox M. 1998. Sauropod dinosaur embryos from the Late Cretaceous of Patagonia. Nature. 396(6708):258–261. doi:https://doi.org/10.1038/24370.
   Web of Science ®Google Scholar
• Choi S, Han S, Lee YN. 2019. Electron backscatter diffraction (EBSD) analysis of maniraptoran eggshells with important implications for microstructural and taphonomic interpretations. Palaeontology. 62(5):777–803. doi:https://doi.org/10.1111/pala.12427.
   Web of Science ®Google Scholar
• Clark JM, Norell MA, Chiappe LM. 1999. An oviraptorid skeleton from the late Cretaceous of Ukhaa Tolgod, Mongolia, preserved in an avianlike brooding position over an oviraptorid nest. Am Mus Novit. 3265:1–36.
   Google Scholar
• Dong ZM, Currie PJ. 1996. On the discovery of an oviraptorid skeleton on a nest of eggs at Bayan Mandahu, Inner Mongolia, People’s Republic of China. Can J Earth Sci. 33(4):631–636. doi:https://doi.org/10.1139/e96-046.
   Web of Science ®Google Scholar
• Erben HK, Hoefs J, Wedepohl KH. 1979. Paleobiological and isotopic studies of eggshells from a declining dinosaur species. Paleobiology. 5(4):380–414. doi:https://doi.org/10.1017/S0094837300016900.
   Web of Science ®Google Scholar
• Fang XS, Lu LW, Jiang YG, Yang LF. 2003. Cretaceous fossil eggs from the Tiantai basin of Zhejiang, with a discussion on the extinction of dinosaurs. Geol. Bull. Of China. 22(7):512–520. Chinese with English Abstract.
   Google Scholar
• Fang XS, Wang YZ, Jiang YG. 2000. On the Late Cretaceous fossil eggs of Tiantai, Zhejiang. Geol. Rev 46(1):105–112. Chinese with English Summary.
   Google Scholar
• Fanti F, Currie PJ, Badamgarav D. 2012. New specimens of Nemegtomaia from the Baruungoyot and Nemegt Formations (late Cretaceous) of Mongolia. PLoS One. 7(2):e31330. doi:https://doi.org/10.1371/journal.pone.0031330.
   PubMed Web of Science ®Google Scholar
• Fernández MS, Khosla A. 2015. Parataxonomic review of the Upper Cretaceous dinosaur eggshells belonging to the oofamily Megaloolithidae from India and Argentina. Hist Biol. 27(2):158–180. doi:https://doi.org/10.1080/08912963.2013.871718.
   Web of Science ®Google Scholar
• Garcia G, Vianey-Liaud M. 2001. Nouvelles données sur les coquilles d’ufs de dinosaures Megaloolithidae du Sud de la France: systématique et variabilité intraspécifique. Cr Acad Sci Ii B-mec. 332(3):185–191. French with English Abstract.
   Google Scholar
• Grellet-Tinner G, Chiappe L, Norell M, Bottjer D. 2006. Dinosaur eggs and nesting behaviors: a palaeobiological investigation. Palaeogeogr Palaeoclimatol Palaeoecol. 232(2–4):294–321. doi:https://doi.org/10.1016/j.palaeo.2005.10.029.
   Web of Science ®Google Scholar
• Grigorescu D. 2010. The ‘Tustea Puzzle’: hadrosaurid (Dinosauria, Ornithopoda) hatchlings associated with Megaloolithidae eggs in the Maastrichtian of the Hateg Basin (Romania). Ameghiniana. 47(1):89–97. doi:https://doi.org/10.5710/AMGH.v47i1.9.
   Web of Science ®Google Scholar
• Grigorescu D. 2016. The ‘Tuştea puzzle’ revisited: late Cretaceous (Maastrichtian) Megaloolithus eggs associated with Telmatosaurus hatchlings in the Haţeg Basin. Hist Biol. 29(5):627–640. doi:https://doi.org/10.1080/08912963.2016.1227327.
   Web of Science ®Google Scholar
• Grigorescu D, Garcia G, Csiki Z, Codrea Vlad BA. 2010. Uppermost Cretaceous megaloolithid eggs from the Haţeg Basin, Romania, associated with hadrosaur hatchlings: search for explanation. Palaeogeogr Palaeoclimatol Palaeoecol. 293(3–4):360–374. doi:https://doi.org/10.1016/j.palaeo.2010.03.031.
   Web of Science ®Google Scholar
• Hirsch KF. 1994. Upper Jurassic eggshells from the Western Interior of North America. In: Carpenter K, Hirsch KF, Horner JR, editors. Dinosaur eggs and babies. Cambridge: Cambridge University Press; p. 137–150.
   Google Scholar
• Hirsch KF, Quinn B. 1990. Eggs and eggshell fragments from the Upper Cretaceous Two Medicine Formation of Montana. J Vertebr Paleontol. 10(4):491–511. doi:https://doi.org/10.1080/02724634.1990.10011832.
   Google Scholar
• Horner JR. 1982. Evidence of colonial nesting and ‘site fidelity’ among ornithischian dinosaurs. Nature. 297(5868):675–676. doi:https://doi.org/10.1038/297675a0.
   Web of Science ®Google Scholar
• Horner JR. 1999. Egg clutches and embryos of two hadrosaur dinosaurs. J Vertebr Paleontol. 19(4):607–611. doi:https://doi.org/10.1080/02724634.1999.10011174.
   Web of Science ®Google Scholar
• Horner JR. 2000. Dinosaur reproduction and parenting. Annu Rev Earth Pl Sc. 28(1):1–62. doi:https://doi.org/10.1146/annurev.earth.28.1.19.
   Google Scholar
• Horner JR, Makela R. 1979. Nest of juveniles provides evidence of family structure among dinosaurs. Nature. 282(5736):296–298. doi:https://doi.org/10.1038/282296a0.
   Web of Science ®Google Scholar
• Horner JR, Weishampel DB. 1988. A comparative embryological study of two ornithischian dinosaurs. Nature. 332(6161):256–257. doi:https://doi.org/10.1038/332256a0.
   Web of Science ®Google Scholar
• Horner JR, Weishampel DB. 1996. A comparative embryological study of two ornithischian dinosaurs, a correction. Nature. 383(6595):103. doi:https://doi.org/10.1038/383103b0.
   PubMed Web of Science ®Google Scholar
• Hu CZ, Cheng ZW, Pang QQ, Fang XS. 2001. Shantungosaurus giganteus. Beijing. Chinese: Geological Publishing House.
   Google Scholar
• Jackson FD, Varricchio DJ. 2010. Fossil eggs and eggshell from the lowermost Two Medicine Formation of western Montana, Sevenmile Hill locality. J Vertebr Paleontol. 30(4):1142–1156. doi:https://doi.org/10.1080/02724634.2010.483537.
   Web of Science ®Google Scholar
• Jensen JA. 1966. Dinosaur eggs from the Upper Cretaceous North Horn Formation of central Utah. Brigham Young Univ. Geol.Stud. 13:55–67.
   Google Scholar
• Jin XS, Varricchio DJ, Poust AW, He T. 2020. An oviraptorosaur adult-egg association from the Cretaceous of Jiangxi Province, China. J Vertebr Paleontol. e1739060.
   Web of Science ®Google Scholar
• Karlsson O, Lilja C. 2008. Eggshell structure, mode of development and growth rate in birds. Zoology. 111(6):494–502. doi:https://doi.org/10.1016/j.zool.2007.11.005.
   PubMed Web of Science ®Google Scholar
• Kundrát M, Cruickshank ARI, Manning TW, Nudds J. 2008. Embryos of therizinosauroid theropods from the Upper Cretaceous of China: diagnosis and analysis of ossification patterns. Acta Zool (Stockholm). 89(3):231–251. doi:https://doi.org/10.1111/j.1463-6395.2007.00311.x.
   Google Scholar
• Kurzanov SM, Mikhailov KE. 1989. Dinosaur eggshells from the Lower Cretaceous of Mongolia. In: Gillette DD, Lockley MG, editors. Dinosaur Tracks and Traces. Cambridge: Cambridge University Press; p. 109–113.
   Google Scholar
• Lucas SG, Bray ES, Emry RJ, Hirch KF. 2012. Dinosaur eggshell and the Cretaceous-Paleogene boundary in the Zaysan Basin, eastern Kazakstan. J Stratigr. 36(2):417–435.
   Google Scholar
• Mikhailov KE. 1991. Classification of fossil eggshells of amniotic vertebrates. Palaeontol. 36(2):193–238.
   Google Scholar
• Mikhailov KE. 1994. Eggs of sauropod and ornithopod dinosaurs from the Cretaceous deposits of Mongolia. Paleontol J. 28(3):141–159.
   Google Scholar
• Mikhailov KE. 1997. Fossil and recent eggshell in amniotic vertebrates: fine structure, comparative morphology and classification. Spec Pap Palaeontol. 56:1–76.
   Google Scholar
• Moreno-Azanza M, Canudo JI, Gasca JM. 2014. Spheroolithid eggshells in the Lower Cretaceous of Europe. Implications for eggshell evolution in ornithischian dinosaurs. Cretac Res. 51:75–87. doi:https://doi.org/10.1016/j.cretres.2014.05.017.
   Web of Science ®Google Scholar
• Norell MA, Clark JM, Chiappe LM, Demberelyin D. 1995. A nesting dinosaur. Nature. 378(6559):774. doi:https://doi.org/10.1038/378774a0.
   Web of Science ®Google Scholar
• Norell MA, Clark JM, Demberelyin D, Rhinchen B, Chiappe LM, Davidson AR, Mckenna MC, Altangerel P, Novacek MJ. 1994. A theropod dinosaur embryo and the affinities of the flaming cliffs dinosaur eggs. Science. 266(5186):779–782. doi:https://doi.org/10.1126/science.266.5186.779.
   PubMed Web of Science ®Google Scholar
• Prieto-Marquez A, Erickson GM, Ebersole JA. 2016. A primitive hadrosaurid from southeastern North America and the origin and early evolution of ‘duck-billed’ dinosaurs. J Vertebr Paleontol. 36(2):e1054495. doi:https://doi.org/10.1080/02724634.2015.1054495.
   Web of Science ®Google Scholar
• Pu HY, Zelenitsky DK, Lü JC, Currie PJ, Carpenter K, Xu L, Koppelhus EB, Jia SH, Xiao L, Chuang HL, et al. 2017. Perinate and eggs of a giant caenagnathid dinosaur from the Late Cretaceous of central China. Nat Commun. 8(1):14952. doi:https://doi.org/10.1038/ncomms14952.
   PubMedGoogle Scholar
• Sato T, Cheng YN, Wu XC, Zelenitsky DK, Hsiao YF. 2005. A pair of shelled eggs inside a female dinosaur. Science. 308(5720):375. doi:https://doi.org/10.1126/science.1110578.
   PubMed Web of Science ®Google Scholar
• Sellés AG, Galobart À. 2016. Reassessing the endemic European Upper Cretaceous dinosaur egg Cairanoolithus. Hist Biol. 28(5):583–596. doi:https://doi.org/10.1080/08912963.2014.998666.
   Web of Science ®Google Scholar
• Sellés AG, Via B, Galobart À. 2014. Spheroolithus europaeus, oosp. nov. (late Maastrichtian, Catalonia), the youngest oological record of hadrosauroids in Eurasia. J Vertebr Paleontol. 34(2):725–729. doi:https://doi.org/10.1080/02724634.2013.819360.
   Google Scholar
• Shao ZF, Fan SH, Jia SH, Tanaka K, Lü JC. 2014. Intact theropod dinosaur eggs with embryonic remains from the late Cretaceous of southern China. Geol. Bull. Of China. 33(7):941–948.
   Google Scholar
• Sochava AV. 1969. Dinosaur eggs from the Upper Cretaceous of the Gobi Desert. Paleontol J. 4:517–527.
   Google Scholar
• Tanaka K, Zelenitsky DK, Williamson T, Weil A, Therrien F. 2011. Fossil eggshells from the Upper Cretaceous (Campanian) Fruitland Formation, New Mexico. Hist Biol. 23(1):41–55. doi:https://doi.org/10.1080/08912963.2010.499171.
   Web of Science ®Google Scholar
• Wang Q, Li YG, Zhu XF, Fang KY, Wang XL. 2018. New ootype prismatoolithids from the Late Cretaceous, Laiyang Basin and its significance. Vertebr Palas. 56(3):264–272.
   Google Scholar
• Wang Q, Wang XL, Zhao ZK, Jiang YG. 2010a. A new oogenus of Elongatoolithidae from the Upper Cretaceous Chichengshan Formation of Tiantai basin, Zhejiang province. Vertebr Palas. 48(2):111–118. Chinese with English Summary.
   Google Scholar
• Wang Q, Wang XL, Zhao ZK, Jiang YG. 2012a. A new oofamily of dinosaur egg from the Upper Cretaceous of Tiantai Basin, Zhejiang Province, and its mechanism of eggshell formation. Chinese Sci Bull. doi:https://doi.org/10.1007/s11434-012-5353-2
   Web of Science ®Google Scholar
• Wang Q, Zhao ZK, Wang XL, Jiang YG. 2011. New ootypes of dinosaur eggs from the Late Cretaceous in Tiantai Basin, Zhejiang province, China. Vertebr Palas. 49(4):446–449.
   Google Scholar
• Wang Q, Zhao ZK, Wang XL, Jiang YG, Zhang SK. 2010b. A new oogenus of macroelongatoolithid eggs from the Upper Cretaceous Chichengshan Formation of the Tiantai Basin, Zhejiang Province and a revision of the macroelongatoolithids. Acta Palaeontol. Sinica. 49(1):73–86. Chinese with English Summary.
   Google Scholar
• Wang Q, Zhao ZK, Wang XL, Li N, Zou SL. 2013a. A new form of Elongatoolithidae, Undulatoolithus pengi oogen. Zootaxa. 3746(1):194–200. et oosp. nov. from Pingxiang, Jiangxi, China. doi:https://doi.org/10.11646/zootaxa.3746.1.9.
   PubMed Web of Science ®Google Scholar
• Wang Q, Zhao ZK, Wang XL, Zhang SK, Jiang YG. 2013b. New forms of dictyoolithids from the Tiantai Basin, Zhejiang Province of China and a parataxonomic revision of the dictyoolithids. Vertebr Palas. 51(1):43–54.
   Google Scholar
• Wang XL, Wang Q, Jiang SX, Cheng X, Zhang JL, Zhao ZK, Jiang YG. 2012b. Dinosaur egg faunas of the Upper Cretaceous terrestrial red beds of China and their stratigraphical significance. J Stratigr. 36(2):400–416.
   Google Scholar
• Weishampel DB, Fastovsky DE, Watabe M, Varricchio D, Jackson F, Tsogtbaatar K, Barsbold R. 2008. New oviraptorid embryos from Bugin-Tsav, Nemegt Formation (Upper Cretaceous), Mongolia, with insights into their habitat and growth. J Vertebr Paleontol. 28(4):1110–1119. doi:https://doi.org/10.1671/0272-4634-28.4.1110.
   Web of Science ®Google Scholar
• Wilson JA, Mohabey DM, Peters SE, Head JJ. 2010. Predation upon Hatchling Dinosaurs by a New Snake from the Late Cretaceous of India. PLoS Biol. 8(3):e1000322. doi:https://doi.org/10.1371/journal.pbio.1000322.
   PubMed Web of Science ®Google Scholar
• Wiman C. 1929. Die Kreide-Dinosaurier aus Shantung. Paleontol Sinica Ser C. 6(1):1–67.
   Google Scholar
• Xue XX, Yue LP, Zhang YX. 1994. Comparison of magnetism, biostratigraphy and lithostratigraphic boundary of red strata in Shanyang basin, Shaanxi. Sci China Ser B. 24(4):413–417. Chinese.
   Google Scholar
• Xue XX, Zhang YX, Bi Y, Yue LP, Chen DL. 1996. The development and environmental changes of the intermontane basins in the eastern part of Qinling Mountains. Beijing. Chinese: Geological Publishing House. with English Summary.
   Google Scholar
• Xue XX, Zhang YX, Wang HN. 1987. Subdivision and correlation of the red beds in Shanyang basin. J Northwest U. 17(3):58–63. Chinese with English Abstract.
   Google Scholar
• Yan J, Chen JF, Xie Z, Zhou TX. 2003. Mantle capture in later Cretaceous basalt in eastern Shandong: a new evidence for time range of lithospheric thinning in eastern china. Chinese Sci Bull. 48(14):1570–1574. Chinese.
   Google Scholar
• Young CC. 1954. Fossil reptilian eggs from Laiyang, Shantung, China. Sci Sin. 3:505–522.
   Google Scholar
• Young CC. 1958. The dinosaurian remains of Laiyang, Shantung. Paleontol Sinica New Ser C. 16:1–138.
   Google Scholar
• Zelenitsky DK, Hills LV. 1997. Normal and pathological eggshells of Spheroolithus albertensis, oosp. nov. from the Oldman Formation (Judith River Group, late Campanian), southern Alberta. J Vertebr Paleontol. 17(1):167–171. doi:https://doi.org/10.1080/02724634.1997.10010960.
   Web of Science ®Google Scholar
• Zhang JL, Wang XL, Wang Q, Jiang SX, Cheng X, Li N, Qiu R. 2019. A new saurolophine hadrosaurid (Dinosauria: ornithopoda) from the Upper Cretaceous of Shandong, China. An Acad Bras Cienc. 91(Suppl 2):e20160920. doi:https://doi.org/10.1590/0001-3765201720160920.
   Google Scholar
• Zhang SK. 2010. A parataxonomic revision of the Cretaceous faveoloolithid eggs of China. Vertebr Palas. 48(3):203–219. Chinese with English Summary.
   Google Scholar
• Zhang SK, Yang TR, Li ZQ, Hu YG. 2018. New dinosaur egg material from Yunxian, Hubei Province, China resolves the classification of dendroolithid eggs. Acta Palaeontol Pol. 63(4):671–678. doi:https://doi.org/10.4202/app.00523.2018.
   Web of Science ®Google Scholar
• Zhang YG, Wang KB, Chen SQ, Liu D, Xing H. 2020. Osteological re‐assessment and taxonomic revision of “Tanius laiyangensis” (Ornithischia: hadrosauroidea) from the Upper Cretaceous of Shandong, China. Anat Rec. 303(4):790–800. doi:https://doi.org/10.1002/ar.24097.
   Google Scholar
• Zhao ZK. 1975. The microstructure of dinosaurian eggshells of Nanxiong Basin, Guangdong Province: on the classification of dinosaur eggs. Vertebr Palas. 13(2):105–117. Chinese.
   Google Scholar
• Zhao ZK. 1979. The advancement of research on the dinosaurian eggs in China. In: Mesozoic and Cenozoic Red Beds in South China. Beijing. Chinese: Science Press. Institute of Vertebrate Paleontology and Paleoanthropology, Nanjing Institute of Geology and Paleontology (eds); p. 330–340.
   Google Scholar
• Zhao ZK, Jiang YK. 1974. Microscopic studies on the dinosaurian egg-shells from Laiyang, Shandong Province. Sci Sin. 17:73–83.
   Google Scholar
• Zhao ZK, Mao XY, Chai CF. 1999. Dinosaurs survived the K/T event as evidenced by iridium anomalies in the Nanxiong Basin, South China: evidence from dinosaur eggshells. In: Bravo AM, Reyes T, editors. First International Symposium on Dinosaur Eggs and Babies (Abstracts). Catalonia (Spain): Isona i Conca Dellà; p. 65–67.
   Google Scholar
• Zhao ZK, Wang Q, Zhang SK. 2015. Dinosaur Eggs. Palaeovertebrata Sinica. Vol II, Fascicle 7. Beijing. Chinese: Science Press.
   Google Scholar
• Zhao ZK, Ye J, Li HM, Zhao ZH, Yan Z. 1991. Extinction of the dinosaurs across the Cretaceous-Tertiary boundary in Nanxiong basin, Guangdong province. Vertebr Palas. 29(1):1–20. Chinese with English Summary.
   Google Scholar
• Zhao ZK, Ye J, Wang Q. 2017. Dinosaur extinction and subsequent mammalian recovery during the Cretaceous-Paleogene (K/Pg) transition in the Nanxiong Basin. Chinese Sci Bull. 62(17):1869–1881. Chinese with English Abstract. doi:https://doi.org/10.1360/N972016-00886.
   Google Scholar
• Zhao ZK, Zhang SK, Wang Q, Wang XL. 2013. Dinosaur diversity during the transition between the middle and late parts of the Late Cretaceous in eastern Shandong Province. China: evidence from dinosaur eggshells. Chinese Sci Bull. 58(36):4663–4669. doi:https://doi.org/10.1007/s11434-013-6059-9.
   Google Scholar
• Zhen SN. 1976. A new species of hadrosaur from Shandong. Vertebr Palas. 14: 166–168. Chinese with English Abstract.
   Google Scholar
• Zhu XF, Fang KY, Wang Q, Deng L, Liu YC, Wen J, Wang XL, Wang XR. 2019. The first Similifaveoloolithidae (Wormoolithus luxiensis oogen. et oosp. nov.) from the Upper Cretaceous of Jiangxi Province, China. Hist Biol. 1–10. doi:https://doi.org/10.1080/08912963.2019.1658093
   Google Scholar