Publication Cover
Historical Biology
An International Journal of Paleobiology
Volume 35, 2023 - Issue 11
1,125
Views
1
CrossRef citations to date
0
Altmetric
Research Articles

Actinopterygian and chondrichthyan ichthyoliths reveal enhanced cosmopolitanism in Late Triassic marine ecosystems

ORCID Icon, &
Pages 2022-2035 | Received 13 Jul 2022, Accepted 28 Sep 2022, Published online: 15 Nov 2022

References

  • Agassiz L. 1832. Untersuchungen über die fossilen Fische aus der Lias-Formation. Jahrbuch für Mineralogie, Geognosie, Geologie und Petrefaktenkunde. 3:139–149.
  • Agassiz L. 1835. Recherches Sur Les Poissons Fossiles. Neuchatel: Imprimerie de Petitpierre; p. 33–52.
  • Agassiz L. 1843. Research On Fossil Fish. Imprimerie de Petitpierre. Volume I (book 18). Neuchatel.
  • Allard H, Carpenter SC, Duffin CJ, Benton MJ. 2015. Microvertebrates from the classic Rhaetian bone beds of Manor Farm Quarry, near Aust (Bristol, UK). Proceedings of the Geologists’ Association. 126:762–776. doi:10.1016/j.pgeola.2015.09.002.
  • Andreev PS, Cuny G. 2012. New Triassic stem selachimorphs (Chondrichthyes, Elasmobranchii) and their bearing on the evolution of dental enameloid in Neoselachii. Journal of Vertebrate Paleontology. 32(2):255–266. doi:10.1080/02724634.2012.644646.
  • Balini M, Jenks JF, Martin R, McRoberts CA, Orchard MJ, Silberling NJ. 2015. The Carnian/Norian boundary succession at Berlin-Ichthyosaur State Park (Upper Triassic, central Nevada, USA). Paläontol Z. 89(3):399–433. doi:10.1007/s12542-014-0244-2.
  • Baumiller TK, Salamon MA, Gorzelak P, Mooi R, Messing CG, Gahn FJ. 2010. Post-Paleozoic crinoid radiation in response to benthic predation preceded the Mesozoic marine revolution. Proc Natl Acad Sci USA. 107(13):5893–5896. doi:10.1073/pnas.0914199107.
  • Beltan L, Freneix S, Janvier P, Lopez-Paulsen O. 1987. The Triassic fauna of the Vitiacua Formation in the Villamontes region (Department of Chuquisaca, Bolivia). Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 1987(2):99–115. doi:10.1127/njgpm/1987/1987/99.
  • Benton MJ, Wu F. 2022. Triassic Revolution. Front Earth Sci. 10:899541. doi:10.3389/feart.2022.899541.
  • Böss H-P. 1982. Locomotion and feeding in Mesozoic durophagous fishes. Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 164(1–2):167–171. doi:10.1127/njgpa/164/1982/167.
  • Brough J 1935. On the Structure and Relationships of the Hybodont Sharks. Memories and Proceedings of the Manchester Literary and Philosophical Society. 79:35–50.
  • Bryant HC. 1914. Teeth of a cestraciont shark from the upper Triassic of Northern California. Berkeley (CA): University of California Press. 27-30.
  • Bucur II, Rigaud S, Del Piero N, Fucelli A, Heerwagen E, Peybernes C, Peyrotty G, Verard C, Chablais J, Martini R. 2020. Upper Triassic calcareous algae from the Panthalassa Ocean. Rivista Italiana di Paleontologia e Stratigrafia126. 499–540. doi:10.13130/2039-4942/13681.
  • Camp CL, Welles SP. 1956. Triassic dicynodont reptiles. Part I. Memoirs of the University of California 13:255–341.
  • Cavicchini I, Heyworth HC, Duffin CJ, Hildebrandt C, Benton MJ. 2018. A Rhaetian microvertebrate fauna from Stowey Quarry, Somerset, U.K. Proceedings of the Geologists’ Association. 129:144–158. doi:10.1016/j.pgeola.2018.02.001.
  • Cawley JJ, Marramà G, Carnevale G, Villafaña JA, López‐Romero FA, Kriwet J. 2021. Rise and fall of †Pycnodontiformes: diversity, competition and extinction of a successful fish clade. Ecol Evol. 11(4):1769–1796. doi:10.1002/ece3.7168.
  • Chen L, Cuny G, Wang X. 2007. The chondrichthyan fauna from the Middle-Late Triassic of Guanling (Guizhou province, SW China). Historical Biology. 19(4):291–300. doi:10.1080/08912960701248234.
  • Cope ED, Cope ED. 1870. Synopsis of the extinct batrachia, reptilia and aves of North America, Part 1. Transactions of the American Philosophical Society, New Series. 14(1):1–104. doi:10.2307/1005355.
  • Core Team R. 2020. A language and environment for statistical computing. Vienna (Austria): R Foundation for Statistical Computing. [accessed 2021 Apr 10]. https://www.R-project.org/
  • Crofts SB, Neenan JM, Scheyer TM, Summers AP. 2017. Tooth occlusal morphology in the durophagous marine reptiles, Placodontia (Reptilia: Sauropterygia). Paleobiology. 43(1):114–128. doi:10.1017/pab.2016.27.
  • Cross SRR, Ivanovski N, Duffin CJ, Hildebrandt C, Parker A, Benton MJ. 2018. Microvertebrates from the basal Rhaetian Bone Bed (latest Triassic) at Aust Cliff, S.W. Proceedings of the Geologists’ Association. 129:635–653. doi:10.1016/j.pgeola.2018.06.002.
  • Cuny G, Benton MJ. 1999. Early radiation of the Neoselachian sharks in Western Europe. Geobios. 32(2):193–204. doi:10.1016/S0016-6995(99)80032-1.
  • Cuny G, Rieppel O, Sander PM. 2001. The shark fauna from the Middle Triassic (Anisian) of North-Western Nevada. Zoological Journal of the Linnean Society. 133(3):285–301. doi:10.1111/j.1096-3642.2001.tb00627.x.
  • Cuny G, Risnes S. 2005. The enameloid microstructure of the teeth of synechodontiform sharks (Chondrichthyes: Neoselachii). PalArch’s J Vertebr Palaeontol. 3:8–19.
  • Dillon E, Norris R, O’Dea A. 2017. Dermal denticles as a tool to reconstruct shark communities. Mar Ecol Prog Ser. 566:117–134. doi:10.3354/meps12018.
  • Duffin CJ. 1993. Late Triassic shark teeth (Chondrichthyes, Elasmobranchii) from Saint-Nicolas-de-Port (north-east France). Service géologique de Belgique. 264:7–32.
  • Duffin CJ, Delsate D. 1993. The age of the Upper Triassic vertebrate fauna from Attert (Province of Luxembourg, Belgium). Belgian Geological Survey Professional Paper. 264:33–44.
  • Ferguson HG, Muller SW. 1949. Structural Geology of the Hawthorne and Tonopah Quadrangles. Washington (D.C.): USGS.
  • Fischer V, Cappetta H, Vincent P, Garcia G, Goolaerts S, Martin JE, Roggero D, Valentin X. 2014. Ichthyosaurs from the French Rhaetian indicate a severe turnover across the Triassic–Jurassic boundary. Naturwissenschaften. 101(12):1027–1040. doi:10.1007/s00114-014-1242-7.
  • Gozzi E 2004. Analisi tassonomica e morfofunzionale di Saurichthys e Birgeria (Osteichthyes, Actinopterygii) [Unpublished PhD]: Università degli Studi di Milano.
  • Guex J, Bartolini A, Atudorei V, Taylor D. 2004. High-resolution ammonite and carbon isotope stratigraphy across the Triassic–Jurassic boundary at New York Canyon (Nevada). Earth and Planetary Science Letters. 225(1–2):29–41. doi:10.1016/j.epsl.2004.06.006.
  • Hautmann M. 2004. Early Mesozoic evolution of alivincular bivalve ligaments and its implications for the timing of the ‘Mesozoic marine revolution. 37(2):165–172. doi:10.1080/00241160410005835.
  • Hogler JA. 1992. Taphonomy and Paleoecology of Shonisaurus popularis (Reptilia: Ichthyosauria). PALAIOS. 7(1):108–117. doi:10.2307/3514800.
  • Huxley TH. 1880. On the application of the laws of evolution to the arrangement of the vertebrata, and more particularly of the Mammalia. Proceedings of the Zoological Society of London. 43:649–662.
  • Jeppsson L, Anehus R, Fredholm D. 1999. The Optimal Acetate Buffered Acetic Acid Technique for Extracting Phosphatic Fossils. Journal of Paleontology. 73(5):964–972. http://www.jstor.org/stable/1306854.
  • Johns MJ. 1996. Diagnostic Pedicle Features of Middle and Late Triassic Elasmobranch Scales from Northeastern British Columbia, Canada. Micropaleontology. 42(4):335. doi:10.2307/1485956.
  • Johns MJ, Barnes CR, Orchard MJ. 1997. Taxonomy and biostratigraphy of Middle and Late Triassic elasmobranch ichthyoliths from northeastern British Columbia. Geological Survey of Canada Bulletin. doi:10.4095/209171.
  • Jordan DS. 1907. The fossil fishes of California, with supplementary notes on other species of extinct fishes. University of California Publications, Bulletin of the Department of Geology. 42:335–350.
  • Kelley NP, Motani R, Jiang D, Rieppel O, Schmitz L. 2014. Selective extinction of Triassic marine reptiles during long-term sea-level changes illuminated by seawater strontium isotopes. Palaeogeography, Palaeoclimatology, Palaeoecology. 400:9–16. doi:10.1016/j.palaeo.2012.07.026.
  • Lakin RJ, Duffin CJ, Hildebrandt C, Benton MJ. 2016. The Rhaetian vertebrates of Chipping Sodbury, South Gloucestershire, UK, a comparative study. Proceedings of the Geologists’ Association. 127:40–52. doi:10.1016/j.pgeola.2016.02.010.
  • Landon ENU, Duffin CJ, Hildebrandt C, Davies TG, Simms MJ, Benton MJ. 2017. The first discovery of crinoids and cephalopod hooklets in the British Triassic. Proceedings of the Geologists’ Association. 128:360–373. doi:10.1016/j.pgeola.2017.03.005.
  • Larina E, Bottjer DJ, Corsetti FA, Thibodeau AM, Berelson WM, West AJ, Yager JA. 2021. Ecosystem change and carbon cycle perturbation preceded the end-Triassic mass extinction. Earth and Planetary Science Letters. 576:117180. doi:10.1016/j.epsl.2021.117180.
  • Laws RA. 1982. Late Triassic depositional environments and molluscan associations from west-central Nevada. Palaeogeography, Palaeoclimatology, Palaeoecology. 37(2):131–148. doi:10.1016/0031-0182(82)90036-0.
  • Lazăr I, Panaiotu CE, Grigore D, Sandy MR, Peckmann J. 2011. An unusual brachiopod assemblage in a Late Jurassic (Kimmeridgian) stromatactis mud-mound of the Eastern Carpathians (Hăghimaş Mountains), Romania. Facies. 57(4):627–647. doi:10.1007/s10347-010-0261-x.
  • Leach W. 1822. Dapedium politum. Devon Transactions of the Geological Society of London.
  • Lombardo C, Tintori A. 2005. Feeding specializations in Late Triassic fishes. In: Annali dell’Università degli Studi di Ferrara Museologia Scientifica e Naturalistica. volume speciale. p. 11.
  • López-Arbarello A, Rauhut OWM, Cerdeño E. 2010. The Triassic fish faunas of the Cuyana Basin, Western Argentina. Palaeontology. 53(2):249–276. doi:10.1111/j.1475-4983.2010.00931.x.
  • Lucas SG, Taylor DG, Guex J, Krainer K. 2007. Updated proposal for Global Stratotype Section and Point for the base of the Jurassic System in the New York Canyon area, Nevada, USA. ISJS Newsletter. 34(1):34–42.
  • Maisey JG. 1989. Hamiltonichthys mapesi, g. & sp. nov. (Chondrichthyes, Elasmobranchii), from the Upper Pennsylvanian of Kansas. American Museum of Natural History Novitates. 2931:1–42.
  • Maisey JG, Denton JSS. 2016. Dermal denticle patterning in the Cretaceous hybodont shark Tribodus limae (Euselachii, Hybodontiformes), and its implications for the evolution of patterning in the chondrichthyan dermal skeleton. Journal of Vertebrate Paleontology. 36(5):e1179200. doi:10.1080/02724634.2016.1179200.
  • Manzanares E, Pla C, Ferrón HG, Botella H. 2018. Middle-Late Triassic chondrichthyans remains from the Betic Range (Spain). J Iber Geol. 44(1):129–138. doi:10.1007/s41513-017-0027-1.
  • Martindale RC, Corsetti FA, James NP, Bottjer DJ. 2015. Paleogeographic trends in Late Triassic reef ecology from northeastern Panthalassa. Earth Science Reviews. 142:18–37. doi:10.1016/j.earscirev.2014.12.004.
  • Mears EM, Rossi V, MacDonald E, Coleman G, Davies TG, Arias-Riesgo C, Hildebrandt C, Thiel H, Duffin CJ, Whiteside DI, et al. 2016. The Rhaetian (Late Triassic) vertebrates of Hampstead Farm Quarry, Gloucestershire, UK. Proceedings of the Geologists’ Association. 127:478–505. doi:10.1016/j.pgeola.2016.05.003.
  • Metz ET. 2019. Description, phylogenetic analysis and taphonomy of a new Thalattosaur from the Brisbois member of the Vester Formation (Carnian/Norian) of central Oregon. University of Alaska, Fairbanks.
  • Montague-Judd D. 1999. Paleo-upwelling and the distribution of Mesozoic marine reptiles. University of Arizona.
  • Moreau M, Duffin CJ, Hildebrandt C, Hutchinson D, Parker A, Carpenter S, Benton MJ. 2021. Microvertebrates from the Rhaetian basal bone bed of Saltford, near Bath, SW England. Proceedings of the Geologists’ Association.132:174–187. 10.1016/j.pgeola.2020.11.003.
  • Müller MK. 2011. The fish fauna of the Late Jurassic Solothurn Turtle Limestone (NW Switzerland). Swiss J Geosci. 104(S1):(S1):133–146. doi:10.1007/s00015-011-0061-5.
  • Muller SWM, Ferguson HG. 1939. Mesozoic stratigraphy of the Hawthorne and Tonopah quadrangles, Nevada. GSA Bulletin. 50(10):1573–1624. doi:10.1130/GSAB-50-1573.
  • Nordén KK, Duffin CJ, Benton MJ. 2015. A marine vertebrate fauna from the Late Triassic of Somerset, and a review of British placodonts. Proceedings of the Geologists' Association. 126:564–581. doi:10.1016/j.pgeola.2015.07.001.
  • Oldow JS. 1984. Evolution of a Late Mesozoic back-arc fold and thrust belt, Northwestern Great Basin, U.S.A. Tectonophysics. 102(1–4):245–274. doi:10.1016/0040-1951(84)90016-7.
  • Orchard MJ, McRoberts CA, Tozer ET, Johns MJ, Sandy MR, Shaner JS. 2001a. An intercalibrated biostratigraphy of the Upper Triassic of Black Bear Ridge, Williston Lake, northeast British Columbia. Geological Survey of Canada. doi:10.4095/211991.
  • Orchard MJ, Zonneveld JP, McRoberts CA, Sandy MR, Tozer ET, Carrelli GG. 2001b. Fossil succession and sequence stratigraphy of the Upper Triassic of Black Bear Ridge, northeast British Columbia, a GSSP prospect for the Carnian-Norian boundary. Albertiana. 25:10–22.
  • Ősi A, Botfalvai G, Prondvai E, Hajdu Z, Czirják G, Szentesi Z, Pozsgai E, Götz AE, Makádi L, Csengődi D, et al. 2013. First report of Triassic vertebrate assemblages from the Villány Hills (Southern Hungary). Central European Geology. 56:297–335. doi:10.1556/CEuGeol.56.2013.4.2.
  • Patterson C. 1966. British Wealden sharks. Bulletin of the British Museum of Natural History (Geology). 11: 283–350. 31 fig., 5 pl.
  • Plieninger W. 1847. Zahne aus der oberen Grenzbreccie des Keupers bei Degerloch und Steinenbronn. Jahreshefte des Vereins für Vaterländische Naturkunde in Württemberg. 164–167:164–167.
  • Reif W-E. 1978. Types of morphogenesis of the dermal skeleton in fossil sharks. Paläontol Z. 52(1–2):110–128. doi:10.1007/BF03006733.
  • Reilly M, Breyer J, Oldow J. 1980. Petrographic provinces and provenance of the Upper Triassic Luning Formation, west-central Nevada: summary. Geological Society of America Bulletin. 91(10):573–575. doi:10.1130/0016-7606(1980)91<573:PPAPOT>2.0.CO;2.
  • Renesto S, Dalla Vecchia FM. 2018. The Late Triassic World: earth in a Time of Transition. Cham: Springer International Publishing; p. 263–313. doi:10.1007/978-3-319-68009-5_8.
  • Rieppel O. 2002. Feeding mechanics in Triassic stem-group sauropterygians: the anatomy of a successful invasion of Mesozoic seas. Zoological Journal of the Linnean Society. 135(1):33–63. doi:10.1046/j.1096-3642.2002.00019.x.
  • Rieppel OC, Kindlimann R, Bucher H. 1996. Anew fossil fish fauna from the Middle Triassic (Anisian) of north-western Nevada. In: Mesozoic Fishes - Systematics and Paleoecology Pfeil, Friedrich. Mesozoic Fishes - Systematics and Paleoecology. Verlag: Germany: Verlag Dr. Friedrich Pfeil; p. 501–512.
  • Rieppel O, Wild R. 1996. A revision of the genus Nothosaurus (Reptilia: Sauropterygia)from the Germanic Triassic, with comments on the status of Conchiosaurus clavatus. Chicago (Ill.): Field Museum of Natural History.
  • Romano C, Jenks JF, Jattiot R, Scheyer TM, Bylund KG, Bucher H. 2017. Marine Early Triassic Actinopterygii from Elko County (Nevada, USA): implications for the Smithian equatorial vertebrate eclipse. J Paleontol. 91(5):1025–1046. doi:10.1017/jpa.2017.36.
  • Sander PM. 2000. Ichthyosauria: their diversity, distribution, and phylogeny. Paläont Z. 74(1):1–35. doi:10.1007/BF02987949.
  • Sander PM, Rieppel OC, Bucher H. 1994. New Marine Vertebrate Fauna from the Middle Triassic of Nevada. Journal of Paleontology. 68(3):676–680. https://www.jstor.org/stable/1306213.
  • Sansom IJ. 2000. Late Triassic (Rhaetian) conodonts and ichthyoliths from Chile. Geol Mag. 137(2):129–135. doi:10.1017/S0016756800003757.
  • Schaeffer B, Dunkle DH. 1950. A semionotid fish from the Chinle Formation, with consideration of its relationships. American Museum Novitates. 1457:1–29. https://digitallibrary.amnh.org/handle/2246/2356.
  • Scheyer TM, Romano C, Jenks J, Bucher H. 2014. Early Triassic Marine Biotic Recovery: the Predators’ Perspective.Farke AA, editor. PLoS ONE. 9(3):e88987. doi:10.1371/journal.pone.0088987.
  • Sibert E, Cramer K, Hastings P, Norris R. 2017. Methods for isolation and quantification of microfossil fish teeth and elasmobranch dermal denticles (ichthyoliths) from marine sediments. Palaeontologia Electronica. 20(1):1–14. doi:10.26879/677.
  • Silberling NJ. 1959. Pre-Tertiary Stratigraphy and Upper Triassic Paleontology of the Union District Shoshone Mountains Nevada. Washington (D.C.): United States Geological Survey.
  • Slater TS, Duffin CJ, Hildebrandt C, Davies TG, Benton MJ. 2016. Microvertebrates from multiple bone beds in the Rhaetian of the M4–M5 motorway junction, South Gloucestershire, U.K. Proceedings of the Geologists' Association. 127:464–477. doi:10.1016/j.pgeola.2016.07.001.
  • Smithwick FM, Johanson Z. 2015. Feeding ecology of the deep-bodied fish Dapedium (Actinopterygii, Neopterygii) from the Sinemurian of Dorset, England. Palaeontology. 58(2):293–311. doi:10.1111/pala.12145.
  • Speed RC. 1978. Paleogeographic and plate tectonic evolution of the early Mesozoic marine province of the western Great Basin Howell, D.G., McDougall, M.K. Mesozoic Paleogeography of the Western United States Pacific Section: SEPM (Society for Sedimentary Geology) . ; p. 253–270.
  • Stanley GD, González-León C, Sandy MR, Daryan BS, Doyle P, Tamura M, Erwin DH. 1994. Upper Triassic Invertebrates from the Antimonio Formation, Sonora, Mexico. Memoir (The Paleontological Society). 36:1–33. http://www.jstor.org/stable/1315620.
  • Storrs G. 1994. Fossil vertebrate faunas of the British Rhaetian (latest Triassic). Zoological Journal of the Linnean Society. 112(1–2):217–259. doi:10.1111/j.1096-3642.1994.tb00319.x.
  • Stumpf S, Ansorge J, Pfaff C, Kriwet J. 2017. Early Jurassic diversification of pycnodontiform fishes (Actinopterygii, Neopterygii) after the end-Triassic extinction event: evidence from a new genus and species, Grimmenodon aureum. Journal of Vertebrate Paleontology. 37(4):e1344679. doi:10.1080/02724634.2017.1344679.
  • Tackett LS, Bottjer DJ. 2012. Faunal succession of Norian (Late Triassic) level-bottom benthos in the Lombardian Basin: implications for the timing, rate, and nature of the early Mesozoic marine revolution. Palaios. 27(8):585–593. doi:10.2110/palo.2012.p12-028r.
  • Tackett LS, Bottjer DJ. 2016. Paleoecological succession of Norian (Late Triassic) benthic fauna in Eastern Panthalassa (Luning and Gabbs Formations, west-central Nevada). Palaios. 31(4):190–202. doi:10.2110/palo.2015.070.
  • Tackett LS, Kaufman AJ, Corsetti FA, Bottjer DJ. 2014. Strontium isotope stratigraphy of the Gabbs Formation (Nevada): implications for global Norian-Rhaetian correlations and faunal turnover. Lethaia. 47(4):500–511. doi:10.1111/let.12075.
  • Taylor D, Guex J, Lucas SG. 2021. Ammonoids of the latest Triassic Gabbs Formation at New York Canyon, Mineral County, Nevada. New Mexico Museum of Natural History and Science Bulletin. 82:393–425.
  • Thies D, Hauff RB. 2011. A new species of Dapedium LEACH, 1822 (Actinopterygii, Neopterygii, Semionotiformes) from the Early Jurassic of South Germany. Palaeodiversity. 4:185–221.
  • Thies D, Stevens K, Stumpf S. 2021. Stomach contents of the Early Jurassic fish †Lepidotes Agassiz, 1832 (Actinopterygii, Lepisosteiformes) and their palaeoecological implications. Historical Biology. 33(6):868–879. doi:10.1080/08912963.2019.1665040.
  • Tintori A. 1981. Two new pycnodonts (Pisces, Actinopterygii) from the Upper Triassic of Lombardy (N. Italy). Rivista Italiana di Paleontologia e Stratigrafia 86:795–824.
  • Tintori A. 1998. Fish biodiversity in the marine Norian (Late Triassic) of northern Italy: the first Neopterygian radiation. Italian Journal of Zoology 65(sup1):193–198. 65(sup1):193–198. doi:10.1080/11250009809386812.
  • Tintori A, Hitij T, Jiang D, Lombardo C, Sun Z. 2014. Triassic actinopterygian fishes: the recovery after the end-Permian crisis. Integrative Zoology. 9(4):394–411. doi:10.1111/1749-4877.12077.
  • Vermeij GJ. 1993. Evolution and Escalation. Princeton: Princeton University Press.
  • Walker SE, Brett CE. 2002. Post-Paleozoic Patterns in Marine Predation. Was There a Mesozoic and Cenozoic Marine Predatory Revolution? Paleontol Soc Pap. 8:119–194. doi:10.1017/S108933260000108X.
  • Wemple E. 1906. New Cestraciont teeth from the West-American Triassic. University of California Publications, Bulletin of the Department of Geology. 5:71–73.
  • Yonge C. 1979. Cementation in the Bivalvia. Utrecht: Bohn, Scheltema & Holkema; p. 83–106.
  • Zonneveld J-P, Henderson CM, Stanley GD, Orchard MJ, Gingras MK. 2007. Oldest scleractinian coral reefs on the North American craton: upper Triassic (Carnian), northeastern British Columbia, Canada. Palaeogeography, Palaeoclimatology, Palaeoecology. 243(3–4):421–450. doi:10.1016/j.palaeo.2006.08.012.
  • Zuoyu S, Tintori A, Lombardo C, Dayong J, Weicheng H, Yuanlin S, Feixiang W, Rusconi M. 2008. A new species of the genus Colobodus Agassiz, 1844 (Osteichthyes, Actinopterygii) from the Pelsonian (Anisian, Middle Triassic) of Guizhou, South China. Rivista italiana di Paleontologia e Stratigrafia. 114(3):363–376. doi:10.13130/2039-4942/5908.