Testing the monophyly of the ground-dweller spider genus Harpactea Bristowe, 1939 (Araneae, Dysderidae) with the description of three new species

References

• Alicata, P. (1966). Le Harpactea (Araneae, Dysderidae) della fauna italiana e considerazioni sulla loro origine. Atti Dell’ Accademia Gioenia Di Scienze Naturali in Catania, 6(18), 190–221.
   Google Scholar
• Arnedo, M. A., Oromi, P., Murria, C., Macias-Hernandez, N., & Ribera, C. (2007). The dark side of an island radiation: Systematics and evolution of troglobitic spiders of the genus Dysdera Latreille (Araneae:Dysderidae) in the Canary Islands. Invertebrate Systematics, 21(6), 623–660. https://doi.org/10.1071/IS07015
   Web of Science ®Google Scholar
• Arnedo, M., Gasparo, F., & Opatova, V. (2009). Systematics and phylogeography of the Dysdera erythrina species complex (Araneae, Dysderidae) in Sardinia. ZooKeys, 16, 319–345. https://doi.org/10.3897/zookeys.16.128
   Web of Science ®Google Scholar
• Arnedo, M., Oromi, P., & Ribera, C. (2001). Radiation of the Spider Genus Dysdera (Araneae, Dysderidae) in the Canary Islands: Cladistic Assessment Based on Multiple Data Sets. Cladistics, 17(4), 313–353. https://doi.org/10.1006/clad.2001.0168
   Web of Science ®Google Scholar
• Astrin, J. J., Höfer, H., Spelda, J., Holstein, J., Bayer, S., Hendrich, L., Huber, B. A., Kielhorn, K.-H., Krammer, H.-J., Lemke, M., Monje, J. C., Morinière, J., Rulik, B., Petersen, M., Janssen, H., & Muster, C. (2016). Towards a DNA barcode reference database for spiders and harvestmen of Germany. PLoS One, 11(9), e0162624–24. https://doi.org/10.1371/journal.pone.0162624
   PubMed Web of Science ®Google Scholar
• Ballarin, F., Pantini, P., & Hansen, H. (2011). Catalogo ragionato dei ragni (Arachnida, Araneae) del Veneto. Memorie Del Museo Civico Di Storia Naturale Di Verona, 2, 1–151.
   Google Scholar
• Belloni, S., Martinis, B., & Orombelli, G. (1971). Karst of Italy. Karst Important Region of the Northern Emisphere. M. Herak and V. T. Stringfield (Eds). Elsavier Publishing Company Amsterdam.
   Google Scholar
• Bidegaray-Batista, L., & Arnedo, M. a. (2011). Gone with the plate: the opening of the Western Mediterranean basin drove the diversification of ground-dweller spiders. BMC Evolutionary Biology, 11(1), 317https://doi.org/10.1186/1471-2148-11-317
   PubMedGoogle Scholar
• Bidegaray-Batista, L., Ferrández, M. Á., & Arnedo, M. A. (2014). Winter is coming: Miocene and Quaternary climatic shifts shaped the diversification of Western-Mediterranean Harpactocrates (Araneae, Dysderidae) spiders. Cladistics, 30(4), 428–446. https://doi.org/10.1111/cla.12054
   Web of Science ®Google Scholar
• Bidegaray-Batista, L., Macías-Hernández, N., Oromí, P., & Arnedo, M. A. (2007). Living on the edge: demographic and phylogeographical patterns in the woodlouse-hunter spider Dysdera lancerotensis Simon, 1907 on the eastern volcanic ridge of the Canary Islands. Molecular Ecology, 16(15), 3198–3214. https://doi.org/10.1111/j.1365-294X.2007.03351.x
   PubMed Web of Science ®Google Scholar
• Bosselaers, J., & Van Keer, J. (2016). A redescription of Harpactea dufouri (Thorell, 1873) (Araneae, Dysderidae), its occurrence outside the Balearic Islands, and some notes on the corticalis group of the genus. European Journal of Taxonomy, 222(222), 1–13. https://doi.org/10.5852/ejt.2016.222
   Google Scholar
• Brignoli, P. M. (1978). Ragni di Turchia V. Specie nuove o interessanti, cavernicole ed epigee, di varie famiglie (Araneae). Revue Suisse de Zoologie., 85, 461–541. https://doi.org/10.5962/bhl.part.82243
   Google Scholar
• Burger, M., & Kropf, C. (2007). Genital morphology of the haplogyne spider Harpactea lepida (Arachnida, Araneae, Dysderidae). Zoomorphology, 126(1), 45–52. https://doi.org/10.1007/s00435-007-0029-1
   Web of Science ®Google Scholar
• Chatzaki, M., & Arnedo, M. A. (2006). Taxonomic revision of the epigean representatives of the spider subfamily Harpacteinae (Araneae: Dysderidae) on the island of Crete. Zootaxa, 1169(1), 1–32. https://doi.org/10.11646/zootaxa.1169.1.1
   Google Scholar
• Corrado, C. (1953). Le Grotte della bassa Valcellina. Riv,. Mens. C.A.I, 72(11/12), 367–372.
   Google Scholar
• Deeleman-Reinhold, C. L. (1993). The genus Rhode and the harpacteine genera Stalagtia, Folkia, Minotauria, and Kaemis (Araneae, Dysderidae) of Yugoslavia and Crete, with remarks on the genus Harpactea. Revue Arachnologique, 10(6), 105–135.
   Google Scholar
• Deltshev, C., & Lazarov, S. (2018). Two new spider species, Harpactea simovi sp. n. and H. stoevi sp. n. (Araneae: Dysderidae), from the Balkan Peninsula. Acta Zoologica Bulgarica, 70, 3–7.
   Web of Science ®Google Scholar
• Dimitrov, D., Deltshev, C., & Lazarov, S. (2019). Description of Harpactea popovi sp. n. from Bulgaria with further taxonomic notes on related species (Araneae, Dysderidae). Zootaxa, 4568(3), 593–600. https://doi.org/10.11646/zootaxa.4568.3.13
   Web of Science ®Google Scholar
• Dimitrov, D., & Lazarov, S. (1999). Two new species of Harpactea from Bulgaria (Araneae: Dysderidae). Berichte des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck, 86, 127–129.
   Google Scholar
• Forster, R. R., & Platnick, N. I. (1985). A review of the austral spider family Orsolobidae (Arachnida, Araneae), with notes on the superfamily Dysderoidea. Bulletin of the American Museum of Natural History, 181, 1–230.
   Google Scholar
• Gasparo, F. (1997). Miscellanea biospelologica. Parte I: Friuli. In E. Boegan, (Ed) “Atti e Memorie della Commissione Grotte, (Vol. 34, pp. 17–48). Trieste.
   Google Scholar
• Gasparo, F. (2013). Descrizione di una nuova specie del genere Harpactea Bristowe, 1939 del Carso Triestino (Araneae, Dysderidae). Atti Del Museo Civico Di Storia Naturale - Trieste, 56, 207–217.
   Google Scholar
• Gasparo, F., & Thaler, K. (2000). I ragni cavernicoli della Venezia Giulia (Italia nord-orientale) (Arachnida, Araneae). Atti e Memorie, Commissione Grotte “Eugenio Boegan” 37, 17–55.
   Google Scholar
• Glavaš, I. (2007). Nova istraživanja jame Biokovke u 2007. godini. Subterranea Croatica, 9, 13–16.
   Google Scholar
• Goloboff, P. A., & Catalano, S. A. (2016). TNT version 1.5, including a full implementation of phylogenetic morphometrics. Cladistics, 32(3), 221–238. Retrieved from http://www.lillo.org.ar/phylogeny/tnt/ https://doi.org/10.1111/cla.12160
   PubMed Web of Science ®Google Scholar
• Grottolo, M. (1994). Note complementari su Orotrechus schwienbacheri Gottolo e Martinelli, 1991 (Coleoptera Carabidae Trechinae). «Natura Bresciana» Ann. Mus. Civ. Se. Nat., Brescia, 29(1993), 185–192.
   Google Scholar
• Gücel, S., Charalambidou, I., Göçmen, B., & Kunt, K. B. (2019). New data of spiders (Arachnida, Araneae) of Cyprus. 1. Dysderidae found in caves. ZooKeys, 825, 43–53. https://doi.org/10.3897/zookeys.825.29029
   Google Scholar
• Hlaváč, P., Bregović, P., & Jalžić, B. (2019). Endogean and cavernicolous Coleoptera of the Balkans. XVIII. Strong radiation in caves of the Central Dinarides: seven new species of Thaumastocephalus Poggi et al., 2001 (Staphylinidae: Pselaphinae)(. Zootaxa, 4559(1), 90–110. https://doi.org/10.11646/zootaxa.4559.1.3
   PubMed Web of Science ®Google Scholar
• Kalyaanamoorthy, S., Minh, B. Q., Wong, T. K. F., Von Haeseler, A., & Jermiin, L. S. (2017). ModelFinder: Fast model selection for accurate phylogenetic estimates. Nature Methods, 14(6), 587–589. https://doi.org/10.1038/nmeth.4285
   PubMed Web of Science ®Google Scholar
• Katoh, K., Misawa, K., Kuma, K., & Miyata, T. (2002). MAFFT: a novel method for rapid multiple sequence alignment based on fast Fourier transform. Nucleic Acids Research, 30(14), 3059–3066. https://doi.org/10.1093/nar/gkf436
   PubMed Web of Science ®Google Scholar
• Kearse, M., Moir, R., Wilson, A., Stones-Havas, S., Cheung, M., Sturrock, S., Buxton, S., Cooper, A., Markowitz, S., Duran, C., Thierer, T., Ashton, B., Meintjes, P., & Drummond, A. (2012). Geneious Basic: An integrated and extendable desktop software platform for the organization and analysis of sequence data. Bioinformatics (Oxford, England)), 28(12), 1647–1649. https://doi.org/10.1093/bioinformatics/bts199
   PubMed Web of Science ®Google Scholar
• Lanfear, R., Frandsen, P. B., Wright, A. M., Senfeld, T., & Calcott, B. (2017). Partitionfinder 2: New methods for selecting partitioned models of evolution for molecular and morphological phylogenetic analyses. Molecular Biology and Evolution, 34(3), 772–773. https://doi.org/10.1093/molbev/msw260
   PubMed Web of Science ®Google Scholar
• Lazarov, S., & Dimitrov, D. (2018). Description of Harpactea petrovi sp. n. (Araneae: Dysderidae) from Bulgaria. Acta Zoologica Bulgarica, 70, 293–296.
   Web of Science ®Google Scholar
• Lazarov, S., & Naumova, M. (2010). Two new Harpactea species from Bulgaria (Araneae: Dysderidae). Revue Suisse de Zoologie, 117, 101–110.
   Google Scholar
• Le Peru, B. (2011). The Spiders of Europe, a synthesis of data: Volume 1 Atypidae to Theridiidae. Mémoires de La Société Linnéenne de Lyon, 2, 1–522.
   Google Scholar
• Lohaj, R., & Delić, T. (2019). Playing hard to get: two new species of subterranean Trechini beetles (Coleoptera, Carabidae, Trechinae) from the Dinaric Karst. Deutsche Entomologische Zeitschrift, 66(1), 1–15. https://doi.org/10.3897/dez.66.31754
   Web of Science ®Google Scholar
• Macías-Hernández, N., Oromí, P., & Arnedo, M. A. (2008). Patterns of diversification on old volcanic islands as revealed by the woodlouse-hunter spider genus Dysdera (Araneae, Dysderidae) in the eastern Canary Islands. Biological Journal of the Linnean Society, 94(3), 589–615. https://doi.org/10.1111/j.1095-8312.2008.01007.x
   Web of Science ®Google Scholar
• Macías-Hernández, N., Oromí, P., & Arnedo, M. a. (2010). Integrative taxonomy uncovers hidden species diversity in woodlouse hunter spiders (Araneae, Dysderidae) endemic to the Macaronesian archipelagos. Systematics and Biodiversity, 8(4), 531–553. https://doi.org/10.1080/14772000.2010.535865
   Web of Science ®Google Scholar
• Mammola, S., Cardoso, P., Angyal, D., Balázs, G., Blick, T., Brustel, H., Carter, J., Ćurčić, S., Danflous, S., Dányi, L., Déjean, S., Deltshev, C., Elverici, M., Fernández, J., Gasparo, F., Komnenov, M., Komposch, C., Kováč, L., Kunt, K. B., … Isaia, M. (2019). Continental data on cave-dwelling spider communities across Europe (Arachnida: Araneae). Biodiversity Data Journal, 7, e38492https://doi.org/10.3897/BDJ.7.e38492
   PubMed Web of Science ®Google Scholar
• Mammola, S., Cardoso, P., Ribera, C., Pavlek, M., & Isaia, M. (2018). A synthesis on cave-dwelling spiders in Europe. Journal of Zoological Systematics and Evolutionary Research, 56(3), 301–316. https://doi.org/10.1111/jzs.12201
   Web of Science ®Google Scholar
• Miller, M. A., Pfeiffer, W., & Schwartz, T. (2010, April 2016). Creating the CIPRES Science Gateway for inference of large phylogenetic trees [Paper presentation]. 2010 Gateway Computing Environments Workshop, GCE 2010, https://doi.org/10.1109/GCE.2010.5676129
   Google Scholar
• Mosetti, S. (1954a). Gruppo Triestino Speleologi, Atti del VI Congr. Naz. di Spel., Trieste 1954: LXV-LXVI
   Google Scholar
• Mosetti, S. (1954b). Le Grotte della Valcellina, Atti del VI Congr. Naz. di Spel., Trieste 1954: 310-32
   Google Scholar
• Nentwig, W., Blick, T., Bosmans, R., Gloor, D., Hänggi, A., Kropf, C. (2020). Araneae. Retrieved from https://www.araneae.nmbe.ch
   Google Scholar
• Nguyen, L. T., Schmidt, H. A., Von Haeseler, A., & Minh, B. Q. (2015). IQ-TREE: A fast and effective stochastic algorithm for estimating maximum-likelihood phylogenies. Mol. Biol. Evol, 32(1), 268–274. https://doi.org/10.1093/molbev/msu300
   PubMed Web of Science ®Google Scholar
• Pantini, P., & Isaia, M. (2019). Araneae.it: the online Catalog of Italian spiders, with addenda on other Arachnid Orders occurring in Italy (Arachnida: Araneae, Opiliones, Palpigradi, Pseudoscorpionida, Scorpiones, Solifugae). Fragmenta Entomologica, 51(2), 127–152. https://doi.org/10.4081/fe.2019.374
   Web of Science ®Google Scholar
• Piva, E. (2005). Nuove specie di Orostygia e Oryotus, con note sinonimiche (Coleoptera Cholevidae). Memorie Della Società Entomologica Italiana, 84(1), 3–44. https://doi.org/10.4081/memorieSEI.2005.3
   Google Scholar
• Rambaut, A., Drummond, A. J., Xie, D., Baele, G., & Suchard, M. A. (2018). Posterior summarization in Bayesian phylogenetics using Tracer 1.7. Systematic Biology, 67(5), 901–904. https://doi.org/10.1093/sysbio/syy032
   PubMed Web of Science ®Google Scholar
• Ramírez, M. J., & Michalik, P. (2019). The Spider Anatomy Ontology (SPD)—A Versatile Tool to Link Anatomy with Cross-Disciplinary Data. Diversity, 11(10), 202. https://doi.org/10.3390/d11100202
   Google Scholar
• Roewer, C.-F. (1931). Arachnoideen aus südostalpinen Höhlen gesammelt von Herrn Karl Strasser in den Jahren 1929 und 1930. Mitteilungen Über Höhlen- Und Karstforschung, 1931, 1–17.
   Google Scholar
• Ronquist, F., & Huelsenbeck, J. P. (2003). MrBayes 3: Bayesian phylogenetic inference under mixed models. Bioinformatics (Oxford, England), 19(12), 1572–1574. https://doi.org/10.1093/bioinformatics/btg180
   PubMed Web of Science ®Google Scholar
• Rucner, D., & Rucner, R. (1995). Beitrag zur Kenntnis einiger Arthropoden (Scorpiones, Pseudoscorpiones, Araneae, Acari, Diplopoda und Chilopoda) in den Waldassoziationen Kroatiens. Natura Croatica, 4, 185–225.
   Google Scholar
• Sudar, V., Kuharić, N., Bregović, P., & Kirin, A. (2017). Prva biospeleološka ekspedicija - Biokovo 2017. Subterranea Croatica, 23, 2–20.
   Google Scholar
• Wheeler, W. C., Coddington, J. A., Crowley, L. M., Dimitrov, D., Goloboff, P. A., Griswold, C. E., Hormiga, G., Prendini, L., Ramírez, M. J., Sierwald, P., Almeida-Silva, L., Alvarez-Padilla, F., Arnedo, M. A., Benavides Silva, L. R., Benjamin, S. P., Bond, J. E., Grismado, C. J., Hasan, E., Hedin, M., … Zhang, J. (2017). The spider tree of life: Phylogeny of Araneae based on target-gene analyses from an extensive taxon sampling. Cladistics, 33(6), 574–543. https://doi.org/10.1111/cla.12182
   PubMed Web of Science ®Google Scholar
• World Spider Catalog. (2020). World Spider Catalog. https://doi.org/10.24436/2
   Google Scholar