A reappraisal of Boluochia zhengi (Aves: Enantiornithes) and a discussion of intraclade diversity in the Jehol avifauna, China

References

• Amadon , D. 1963 . Comparison of fossil and recent species: some difficulties . The Condor , 65 : 407 – 409 .
   Google Scholar
• Cau , A. and Arduini , P. 2008 . Enantiophoenix electrophyla gen. et sp. nov. (Aves, Enantiornithes) from the Upper Cretaceous (Cenomanian) of Lebanon and its phylogenetic relationships . Atti della Società Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano , 149 : 293 – 324 .
   Google Scholar
• Chiappe , L. M. and Walker , C. A. 2002 . “ Skeletal morphology and systematics of the Cretaceous Euenantiornithes (Ornithothoraces: Enantiornithes) ” . In Mesozoic Birds: Above the Heads of Dinosaurs , Edited by: Chiappe , L. M. and Witmer , L. M. 240 – 267 . Berkeley : University of California Press .
   Google Scholar
• Chiappe , L. M. , Lamb , J. P., Jr. and Ericson , P. G. P. 2002 . New enantiornithine bird from the marine Upper Cretaceous of Alabama . Journal of Vertebrate Paleontology , 22 : 170 – 174 .
   Web of Science ®Google Scholar
• Chiappe , L. M. , Suzuki , S. , Dyke , G. J. , Watabe , M. , Tsogtbaatar , K. and Barsbold , R. 2006 . A new enantiornithine bird from the Late Cretaceous of the Gobi Desert . Journal of Systematic Palaeontology , 5 : 193 – 208 .
   Web of Science ®Google Scholar
• Chiappe , L. M. 2007 . Glorified Dinosaurs: the Origin and Early Evolution of Birds , 263 Hoboken : John Wiley & Sons, Inc. .
   Google Scholar
• Chiappe , L. M. , Marugán-Lobón , J. , Ji , S. and Zhou , Z. 2008 . Life history of a basal bird: morphometrics of the Early Cretaceous Confuciusornis . Biology Letters , 4 : 719 – 723 .
   PubMedGoogle Scholar
• Clarke , J. A. , Zhou , Z. and Zhang , F. 2006 . Insight into the evolution of avian flight from a new clade of Early Cretaceous ornithurines from China and the morphology of Yixianornis grabaui . Journal of Anatomy , 208 : 287 – 308 .
   PubMed Web of Science ®Google Scholar
• He , H. Y. , Wang , X. L. , Zhou , Z. H. , Wang , F. , Jin , F. , Boven , A. , Shi , G. H. and Zhu , R. X. 2004 . Timing of the Jiufotang Formation (Jehol Group) in Liaoning, northeastern China, and its implications . Geophysical Research Letters , 31 : L12605
   Web of Science ®Google Scholar
• He , H.-Y. , Wang , X.-L. , Jin , F. , Zhou , Z.-H. , Wang , F. , Yang , L.-K. , Ding , X. , Boven , A. and Zhu , R.-X. 2006 . The 40Ar/39Ar dating of the early Jehol Biota from Fengning, Hebei Province, northern China . Geochemistry, Geophysics, Geosystems , 7 : 1 – 8 .
   Web of Science ®Google Scholar
• Hopson , J. A. 2001 . “ Ecomorphology of avian and nonavian theropod phalangeal proportions: implications for the arboreal versus terrestrial origin of bird flight ” . In New Perspectives on the Origin and Early Evolution of Birds , Edited by: Gauthier , J. and Gall , L. F. 211 – 235 . New Haven : Peabody Museum of Natural History .
   Google Scholar
• Hou , L. and Zhang , J. 1993 . A new fossil bird from Lower Cretaceous of Liaoning . Vertebrata PalAsiatica , 31 : 217 – 224 .
   Google Scholar
• Hou , L. 1997 . A carinate bird from the Upper Jurassic of western Liaoning, China . Chinese Science Bulletin , 42 : 413 – 416 .
   Web of Science ®Google Scholar
• Hou , L. , Chiappe , L. M. , Zhang , F. and Chuong , C.-M. 2004 . New Early Cretaceous fossil from China documents a novel trophic specialization for Mesozoic birds . Naturwissenschaften , 91 : 22 – 25 .
   PubMed Web of Science ®Google Scholar
• Isler , M. L. , Isler , P. R. and Whitney , B. M. 1999 . Species limits in Antbirds (Passeriformes: Thamnophilidae): the Myrmotherula surinamensis complex . The Auk , 116 : 83 – 96 .
   Google Scholar
• Li , L. , Duan , Y. , Hu , D. , Wang , L. , Cheng , S. and Hou , L. 2006 . New eoentantiornithid bird from the Early Cretaceous Jiufotang Formation of western Liaoning, China . Acta Geologica Sinica (English edition) , 80 : 38 – 41 .
   Web of Science ®Google Scholar
• Martin , L. D. 1987 . “ The beginning of the modern avian radiation ” . In L’Evolution des Oiseaux d’Apres le Temoignage des Fossiles , Edited by: Mourer-Chauviré , C. 9 – 19 . Lyon : Département des Sciences de la Terre, Université Claude-Bernard .
   Google Scholar
• Morschhauser , E. , Varricchio , D. J. , Gao , C.-H. , Liu , J.-Y. , Wang , X.-R. , Cheng , X.-D. and Meng , Q.-J. 2009 . Anatomy of the Early Cretaceous bird Rapaxavis pani, a new species from Liaoning Province, China . Journal of Vertebrate Paleontology , 29 : 545 – 554 .
   Web of Science ®Google Scholar
• O’Connor , J. K. 2009 . A systematic review of Enantiornithes (Aves: Ornithothoraces) , 600 Los Angeles : University of Southern California . Unpublished doctoral thesis
   Google Scholar
• O’Connor , J. K. , Wang , X.-R. , Chiappe , L. M. , Gao , C.-H. , Meng , Q.-J. , Cheng , X.-D. and Liu , J.-Y. 2009 . Phylogenetic support for a specialized clade of Cretaceous enantiornithine birds with information from a new species . Journal of Vertebrate Paleontology , 29 : 188 – 204 .
   Web of Science ®Google Scholar
• O’Connor , J. K. and Dyke , G. D. 2010 . A reassessment of Sinornis santensis and Cathayornis yandica (Aves: Ornithothoraces) . Proceedings of the VII International Meeting of the Society of Avian Paleontology and Evolution, Records of the Australian Museum , 62 : 7 – 20 .
   Google Scholar
• O’Connor , J. and Chiappe , L. M. forthcoming . A revision of enantiornithine (Aves: Ornithothoraces) skull morphology . Journal of Systematic Palaeontology ,
   Google Scholar
• Sanz , J. L. and Buscalioni , A. D. 1992 . A new bird from the Early Cretaceous of Las Hoyas, Spain, and the early radiation of birds . Palaeontology , 35 : 829 – 845 .
   Web of Science ®Google Scholar
• Sereno , P. C. and Rao , C. 1992 . Early evolution of avian flight and perching: new evidence from the Lower Cretaceous of China . Science , 255 : 845 – 848 .
   PubMed Web of Science ®Google Scholar
• Sereno , P. C. , Rao , C. and Li , J. 2002 . “ Sinornis santensis (Aves: Enantiornithes) from the Early Cretaceous of northeastern China ” . In Mesozoic Birds: Above the Heads of Dinosaurs , Edited by: Chiappe , L. M. and Witmer , L. M. 184 – 208 . Berkeley : University of California Press .
   Google Scholar
• Starck , J. M. and Ricklefs , R. E. 1998 . “ Patterns of development: the altricial–precocial spectrum ” . In Avian Growth and Development , Edited by: Starck , J. M. and Ricklefs , R. E. 3 – 30 . New York : Oxford University Press .
   Google Scholar
• Starck , J. M. and Chinsamy , A. 2002 . Bone microstructure and developmental plasticity in birds and other dinosaurs . Journal of Morphology , 254 : 232 – 246 .
   PubMed Web of Science ®Google Scholar
• Swisher , C. C. III , Wang , X. , Zhou , Z. , Wang , Y. , Jin , F. , Zhang , J. , Xu , X. , Zhang , F. and Wang , Y. 2002 . Further support for a Cretaceous age for the feathered-dinosaur beds of Liaoning, China: new 40Ar/39Ar dating of the Yixian and Tuchengzi formations . Chinese Science Bulletin , 47 : 135 – 138 .
   Web of Science ®Google Scholar
• Wang , X.-R. , O’Connor , J. K. , Zhao , B. , Chiappe , L. M. , Gao , C.-H. and Cheng , X.-D. 2010 . A New Species of Enantiornithes (Aves: Ornithothoraces) based on a Well-Preserved Specimen from the Qiaotou Formation of Northern Hebei, China . Acta Geologica Sinica , 84 : 247 – 256 .
   Web of Science ®Google Scholar
• Yang , W. , Li , S. and Jiang , B. 2007 . New evidence for Cretaceous age of the feathered dinosaurs of Liaoning: zircon U-Pb SHRIMP dating of the Yixian Formation in Sihetun, northeast China . Cretaceous Research , 28 : 177 – 182 .
   Web of Science ®Google Scholar
• Zhang , F. and Zhou , Z. 2000 . A primitive enantiornithine bird and the origin of feathers . Science , 290 : 1955 – 1960 .
   PubMed Web of Science ®Google Scholar
• Zhang , F. , Zhou , Z. , Hou , L. and Gu , G. 2000 . Early diversification of birds: evidence from a new opposite bird . Kexue Tongbao , 45 : 2650 – 2657 .
   Google Scholar
• Zhang , F. , Zhou , Z. and Hou , L.-H. 2003 . “ Birds ” . In The Jehol Biota: the Emergence of Feathered Dinosaurs, Beaked Birds and Flowering Plant , Edited by: Chang , M. , Chen , P. , Wang , Y. , Wang , Y. and Miao , D. 129 – 150 . Shanghai : Shanghai Scientific and Technical Publishers .
   Google Scholar
• Zhang , F. , Ericson , P. G. and Zhou , Z. 2004 . Description of a new enantiornithine bird from the Early Cretaceous of Hebei, northern China . Canadian Journal of Earth Sciences , 41 : 1097 – 1107 .
   Web of Science ®Google Scholar
• Zhang , F. , Zhou , Z. and Benton , M. J. 2008 . A primitive confuciusornithid bird from China and its implications for early avian flight . Science in China, Series D: Earth Sciences , 51 : 625 – 639 .
   Google Scholar
• Zhang , Y.-g. 2006 . Morphology of the distal tarsometatarsus and perching habits in birds . Sichuan Journal of Zoology , 25 : 21 – 27 .
   Google Scholar
• Zheng , X. , Zhang , Z. and Hou , L. 2007 . A new enantiornitine bird with four long rectrices from the Early Cretaceous of northern Hebei, China . Acta Geologica Sinica (English edition) , 81 : 703 – 708 .
   Web of Science ®Google Scholar
• Zhou , Z. , Jin , F. and Zhang , J. 1992 . Preliminary report on a Mesozoic bird from Liaoning, China . Chinese Science Bulletin , 37 : 1365 – 1368 .
   Web of Science ®Google Scholar
• Zhou , Z. 1995 . Discovery of a new enantiornithine bird from the Early Cretaceous of Liaoning, China . Vertebrata PalAsiatica , 33 : 99 – 113 .
   Google Scholar
• Zhou , Z. and Hou , L. 2002 . “ The discovery and study of Mesozoic birds in China ” . In Mesozoic Birds: Above the Heads of Dinosaurs , Edited by: Chiappe , L. M. and Witmer , L. M. 160 – 183 . Berkeley : University of California Press .
   Google Scholar
• Zhou , Z. and Zhang , F. 2003a . Anatomy of the primitive bird Sapeornis chaoyangensis from the Early Cretaceous of Liaoning, China . Canadian Journal of Earth Sciences , 40 : 731 – 747 .
   Web of Science ®Google Scholar
• Zhou , Z. and Zhang , F. 2003b . Jeholornis compared to Archaeopteryx, with a new understanding of the earliest avian evolution . Naturwissenschaften , 90 : 220 – 225 .
   PubMed Web of Science ®Google Scholar
• Zhou , Z. , Barrett , P. M. and Hilton , J. 2003 . An exceptionally preserved Lower Cretaceous ecosystem . Nature , 421 : 807 – 814 .
   PubMed Web of Science ®Google Scholar
• Zhou , Z. , Chiappe , L. M. and Zhang , F. 2005 . Anatomy of the Early Cretaceous bird Eoenantiornis buhleri (Aves: Enantiornithes) from China . Canadian Journal of Earth Sciences , 42 : 1331 – 1338 .
   Web of Science ®Google Scholar
• Zhou , Z. 2006 . Evolutionary radiation of the Jehol Biota: chronological and ecological perspectives . Geological Journal , 41 : 377 – 393 .
   Web of Science ®Google Scholar
• Zhou , Z.-H. and Zhang , F.-C. 2006 . Mesozoic birds of China—a synoptic review . Vertebrata PalAsiatica , 44 : 74 – 98 .
   Google Scholar
• Zhou , Z. , Clarke , J. and Zhang , F. 2008 . Insight into diversity, body size and morphological evolution from the largest Early Cretaceous enantiornithine bird . Journal of Anatomy , 212 : 565 – 577 .
   PubMed Web of Science ®Google Scholar
• Zhou , Z.-H. , Zhang , F.-C. and Li , Z.-H. 2009 . A new basal ornithurine bird (Jianchangornis microdonta gen. et sp. nov.) from the lower Cretaceous of China . Vertebrata PalAsiatica , 47 : 299 – 310 .
   Google Scholar
• Zhu , R. , Pan , Y. , Shi , R. , Liu , Q. and Li , D . 2007 . Palaeomagnetic and 40Ar/39Ar dating constraints on the age of the Jehol Biota and the duration of depostion of the Sihetun fossil-bearing lake sediments, northeast China . Cretaceous Research , 28 : 171 – 176 .
   Web of Science ®Google Scholar
• Zimmer , K. and Whittaker , A. 2000 . The Rufous Cacholote is two species (Furnariidae: Pseudoseisura) . The Condor , 102 : 409 – 422 .
   Google Scholar