The oldest Cenozoic mammal fauna of Europe: implication of the Hainin reference fauna for mammalian evolution and dispersals during the Paleocene

References

• Archibald, J. D. & Rose, K. D. 2005. Womb with a view: the rise of placentals. Pp. 1–8 in J. D. Archibald & K. D. Rose (eds) The rise of placental mammals, origins and relationships of the major extant clades. The Johns Hopkins University Press, Baltimore.
   Google Scholar
• Archibald, J. D., Clemens, W. A., Gingerich, P. D., Krause, D. W., Lindsay, E. H. & Rose, K. D. 1987. First North American land mammal ages of the Cenozoic era. Pp. 24–76 in M. O. Woodburne (ed.) Cenozoic mammals of North America. University of California Press, Berkeley.
   Google Scholar
• Aubry, M.-P., Berggren, W. A., Stott, L. & Sinha, A. 1996. The upper Paleocene–lower Eocene stratigraphic record and the Paleocene–Eocene boundary carbon isotope excursion: implications for geochronology. Geological Society of London Special Publication, 101, 353–380.
   Google Scholar
• Aubry, M.-P., Thiry, M., Dupuis, C. & Berggren, W. A. 2005. The Sparnacian deposits of the Paris Basin: a lithostratigraphic classification. Stratigraphy, 2, 65–100.
   Google Scholar
• Aubry, M.-P., Ouda, K., Dupuis, C., Berggren, W. A. & Van Couvering, J. A. 2007. The Global Standard Stratotype-section and Point (GSSP) for the base of the Eocene Series in the Dababiya section (Egypt). Episodes, 30, 271–286.
   Web of Science ®Google Scholar
• Badgley, C. 1986. Counting individuals in mammalian fossil assemblages from fluvial environments. Palaios, 1, 328–338.
   Google Scholar
• Beard, K. 2002. East of Eden at the Paleocene/Eocene Boundary. Science, 295, 2028–2029.
   PubMed Web of Science ®Google Scholar
• Berggren, W. A. & Aubry, M.-P. 1996. A late Paleocene-early Eocene NW European and North Sea magnetobiochronological correlation work. Geological Society of London Special Publication, 101, 309–352.
   Google Scholar
• Berggren, W. A. & Aubry, M.-P. 1998. The Paleocene/Eocene epoch/series boundary: chronostratigraphic framework and estimated geochronology. Pp. 18–36 in M.-P. Aubry, S. Lucas & W. A. Berggren (eds) Late Paleocene-early Eocene climatic and biotic events in the marine and terrestrial records. Columbia University Press, New York.
   Google Scholar
• Bignot, G. 1993. The position of the Montian stage and related facies within the stratigraphic-palaeogeographic framework of NW Europe during the Danian. Contributions to Tertiary and Quaternary Geology, 29, 47–59.
   Google Scholar
• BiochroM'97 1997. Synthèse et tableaux de corrélations. Pp. 769–805 in J.-P. Aguilar, S. Legendre & J. Michaux (eds), Actes du Congrès BiochroM'97, Montpellier, France, 14-17 April 1997. Mémoires et Travaux de l'E.P.H.E., Institut de Montpellier, 21.
   Google Scholar
• Bloch, J. I., Fischer, D. C., Rose, K. D. & Gingerich, P. D. 2001. Stratocladistic analysis of Paleocene Carpolestidae (Mammalia, Plesiadapiformes) with description of a new late Tiffanian genus. Journal of Vertebrate Paleontology, 21, 119–131.
   Web of Science ®Google Scholar
• Bloch, J. I., Secord, R. & Gingerich, P. D. 2004. Systematics and phylogeny of late Paleocene and early Eocene Palaeoryctinae (Mammalia, Insectivora) from the Clarks Fork and Bighorn Basins, Wyoming. Contributions from the Museum of Paleontology, University of Michigan, 31, 119–154.
   Google Scholar
• Bloch, J. I., Silcox, M. T., Boyer, D. M. & Sargis, E. J. 2007. New Paleocene skeletons and the relationship of plesiadapiforms to crown-clade primates. Proceedings of the National Academy of Sciences, 104, 1159–1164.
   PubMed Web of Science ®Google Scholar
• Boyer, D. M., Prasad, G. V. R., Krause, D. W., Godinot, M., Goswami, A., Verma, O. & Flynn, J. 2010. New postcrania of Deccanolestes from the Late Cretaceous of India and their bearing on the evolutionary and biogeographic history of euarchontan mammals. Naturwissenschaften, 97, 365–377.
   PubMed Web of Science ®Google Scholar
• Boyer, D. M., Costeur, L. & Lipman, Y. 2012. Earliest Record of Platychoerops (Primates, Plesiadapidae), A New Species From Mouras Quarry, Mont de Berru, France. American Journal of Physiological Anthropology, 149, 329–346.
   PubMed Web of Science ®Google Scholar
• Brikiatis, L. 2014. The De Geer, Thulean and Beringia routes: key concepts for understanding early Cenozoic biogeography. Journal of Biogeography, 41, 1036–1054.
   Web of Science ®Google Scholar
• Brinkhuis, H., Schouten, S., Collinson, M. E., Sluijs, A. et al. 2006 Episodic fresh surface waters in the Eocene Arctic Ocean. Nature, 441, 606–609
   PubMed Web of Science ®Google Scholar
• Codrea, V., Solomon, A. A., Venczel, M. & Smith, T. 2014. A new kogaionid multituberculate mammal from the Maastrichtian of the Transylvanian Basin, Romania. Comptes Rendus Palevol, 13, 489–499.
   Web of Science ®Google Scholar
• Cope, E. D. 1882. Notes on Eocene Mammalia. American Naturalist, 16, 522.
   Google Scholar
• Cope, E. D. 1884. The Tertiary Marsupialia. American Naturalist, 18, 686–697.
   Google Scholar
• Crochet, J.-Y. & Sigé B. 1983. Les Mammifères Montiens de Hainin (Paléocène de Belgique). Part III : Marsupiaux. Palaeovertebrata, 13, 51–64.
   Google Scholar
• De Bast, E. & Smith, T. 2013. Reassessment of the small ‘arctocyonid’ Prolatidens waudruae from the early Paleocene of Belgium, and its phylogenetic relationships with ungulate-like mammals. Journal of Vertebrate Paleontology, 33, 964–976.
   Web of Science ®Google Scholar
• De Bast, E. & Smith, T. 2014. Intercontinental dispersal of mammals during the Paleocene: new data from Europe. Journal of Vertebrate Paleontology, Program and Abstracts, 118.
   Google Scholar
• De Bast, E., Sigé, B. & Smith, T. 2012. Diversity of the adapisoriculid mammals from the early Palaeocene of Hainin, Belgium. Acta Palaeontologica Polonica, 57, 35–52.
   Web of Science ®Google Scholar
• De Bast, E., Steurbaut, E. & Smith, T. 2013. New mammals from the marine Selandian of Maret, Belgium, and their implications for the age of the Paleocene continental deposits of Walbeck, Germany. Geologica Belgica, 16, 236–244.
   Web of Science ®Google Scholar
• De Geyter, G., De Man, E., Herman, J., Jacobs, P., Moorkens, T., Steurbaut, E. & Vandenberghe, N. 2006. Disused Paleogene regional stages from Belgium: Montian, Heersian, Landenian, Paniselian, Bruxellian, Laekenian, Ledian, Wemmelian and Tongrian. Geologica Belgica, 9, 203–213.
   Google Scholar
• Douglass, E. 1908. Vertebrate fossils from the Fort Union beds. Annals of Carnegie Museum, 5(2), 11–26.
   Google Scholar
• Flynn, J. J. & Wesley-Hunt, G. D. 2005. Carnivora. Pp. 175–195 in K. D. Rose & J. D. Archibald (eds) The rise of placental mammals, origins and relationships of the major extant clades. The Johns Hopkins University Press, Baltimore.
   Google Scholar
• Folie, A. 2007. Evolution of the amphibians and squamates of the continental Cretaceous-Paleogene transition in Europe. Journal of Vertebrate Paleontology, 27(3), 75A.
   Web of Science ®Google Scholar
• Folie, A., Sigé, B. & Smith, T. 2005. A new scincomorph lizard from the Palaeocene of Belgium and the origin of Scincoidea in Europe. Naturwissenschaften, 92, 542–546.
   PubMed Web of Science ®Google Scholar
• Folie, A. & Smith, T. 2012.The oldest scolecophidian snake. Journal of Vertebrate Paleontology, Program and abstracts, 95.
   Web of Science ®Google Scholar
• Folie, A., Smith, R. & Smith, T. 2013. New amphisbaenian lizards from the early Paleogene of Europe and their implications for the early evolution of modern amphisbaenians. Geologica Belgica, 16, 227–235.
   Web of Science ®Google Scholar
• Fox, R. C. 2004. A new palaeoryctid (Insectivora: Mammalia) from the late Paleocene of Alberta, Canada. Journal of Paleontology, 78, 612–616.
   Web of Science ®Google Scholar
• Fox, R. C., Scott, C. S. & Buckley, G. A. 2015. A ‘giant’ purgatoriid (plesiadapiformes) from the Paleocene of Montana, USA: mosaic evolution in the earliest primates. Palaeontology, 58, 277–291.
   Web of Science ®Google Scholar
• Gazin, C. L. 1956. Paleocene mammalian faunas of the Bison Basin in south-central Wyoming. Smithsonian Miscellaneous Collections 131(6), 1–57.
   Google Scholar
• Gheerbrant, E. 1988. Afrodon chleuhi nov. gen., nov. sp., insectivore (Mammalia, Eutheria) lipotyphlé (?) du Paléocène marocain : données préliminaires. Comptes Rendus de l'Académie des Sciences de Paris, 307, 1303–1309.
   Google Scholar
• Gheerbrant, E. 1991. Bustylus (Eutheria, Adapisoriculidae) and the absence of ascertained marsupials in the Palaeocene of Europe. Terra Nova, 3, 586–592.
   Web of Science ®Google Scholar
• Gheerbrant, E. 1992. Les mammifères paléocènes du Bassin d'Ouarzazate (Maroc). I. Introduction générale et Palaeoryctidae. Palaeontographica, Abteilung A, 224, 67–132.
   Google Scholar
• Gheerbrant, E. 1995. Les Mammifères paléocènes du Bassin d'Ouarzazate (Maroc) III. Adapisoriculidae et autres mammifères (Carnivora, ?Creodonta, Condylarthra, ?Ungulata et incertae sedis). Palaeontographica, Abteilung A, 237, 39–132.
   Google Scholar
• Gheerbrant, E. & Astibia, H. 2012. Addition to the Late Cretaceous Laño mammal faunule (Spain) and to the knowledge of European “Zhelestidae” (Lainodontinae nov.). Bulletin de la Société Géologique de France, 183, 537–546.
   Web of Science ®Google Scholar
• Gheerbrant, E. & Rage, J.-C. 2006. Paleobiogeography of Africa: How distinct from Gondwana and Laurasia? Palaeogeography, Palaeoclimatology, Palaeoecology, 241, 224–246.
   Web of Science ®Google Scholar
• Gheerbrant, E. & Russell, D. E. 1989. Presence of the genus Afrodon [Mammalia, Lipotyphla, Adapisoriculidae] in Europe; new data for the problem of Trans-Tethyan relations between Africa and Europe around the K/T boundary. Palaeogeography, Palaeoclimatology, Palaeoecology, 76, 1–15.
   Web of Science ®Google Scholar
• Gheerbrant, E., Abrial, C. & Cuenca, G. 1997. Nouveau mammifère condylarthre du Paléocène supérieur de Campo (Pyrénées aragonaises, Espagne). Comptes rendus de l'académie des sciences de Paris, Série IIa, 324, 599–606.
   Google Scholar
• Gheerbrant, E., Amaghzaz, M., Bouya, B., Goussard, F. & Letenneur, C. 2014. Ocepeia (Middle Paleocene of Morocco): The oldest skull of an Afrotherian mammal. PLoS ONE 9(2), e89739.
   PubMed Web of Science ®Google Scholar
• Gheerbrant, E., Codrea, V., Hosu, A., Sen, S., Guernet, C., de Lapparent de Broin, F. & Riveline, J. 1999. Découverte dans les Calcaires de Rona (Thanétien ou Sparnacien), Transylvanie, Roumanie: les plus anciens mammifères cénozoïques d'Europe orientale. Eclogae Geologicae Helvetiae, 92, 517–535.
   Google Scholar
• Gingerich, P. D. 1976. Cranial anatomy and Evolution of Early Tertiary Plesiadapidae (Mammalia, Primates). University of Michigan Papers on Paleontology, 15, 1–141.
   Google Scholar
• Gingerich, P. D. 1989. New earliest Wasatchian mammalian fauna from the Eocene of northwestern Wyoming: composition and diversity in a rarely sampled high- floodplain assemblage. University of Michigan Papers on Paleontology, 28, 1–97.
   Google Scholar
• Gingerich, P. D. 1998. Paleobiological perspectives on Mesonychia, Archaeoceti and the origin of whales. Pp. 423–449 in J. G. M. Thewissen (ed.) The emergence of whales: evolutionary patterns in the origin of Cetacea. Plenum Press, New York.
   Google Scholar
• Gingerich, P. D. 2006. Environment and evolution through the Paleocene–Eocene thermal maximum. Trends in Ecology and Evolution, 21, 246–253.
   PubMed Web of Science ®Google Scholar
• Godfriaux, I. & Robaszinsky, F. 1974. Le Montien continental et le Dano-Montien marin des sondages de Hainin (Hainaut, Belgique). Annales de la Société Géologique de Belgique, 97, 185–200.
   Google Scholar
• Godfriaux, I. & Thaler, L. 1972. Note sur la découverte de dents de mammifères dans le Montien continental du Hainaut (Belgique). Académie Royale de Belgique, Bulletins Classe Sciences, Series 5, 58, 536–541.
   Google Scholar
• Godfriaux, I., Aguilar, J.-P. & Thaler, L. 1975. Exploitation par puits du Montien continental à mammifères de Hainin (Belgique). 3^ème Réunion Annuelle des Sciences de la Terre, Montpellier, 170.
   Google Scholar
• Godinot, M., Smith, T. & Smith, R. 1996. Mode de vie et affinités de Paschatherium (Condylarthra, Hyopsodontidae) d'après ses os du tarse. Palaeovertebrata, 25, 225–242.
   Google Scholar
• Goswami, A., Prasad, G. V., Upchurch, P., Boyer, D., Seiffert, E. R., Verma, O., Gheerbrant, E. & Flynn, J. 2011. A radiation of basal arboreal mammals beginning in the late Cretaceous of India. Proceedings of the National Academy of Sciences, USA, 108, 16333–16338.
   PubMed Web of Science ®Google Scholar
• Gregory, W. K. 1910. The orders of mammals. Bulletin of the American Museum of Natural History, 27, 1–524.
   Google Scholar
• Groessens-Van Dyck, M.-C. 1984. Les tortues du Paléocène continental de Hainin et Vinalmont (Belgique). Pp. 133–139 in F. de Broin & E. Jiménez-Fuentes (eds), Studia Palaeocheloniologica 1. Studia Geologica Salmanticensia, Vol. Esp. 1, Ediciones Universidad de Salamanca, Spain.
   Google Scholar
• Guth, C. 1962. Un insectivore de Menat. Annales de Paléontologie, 48, 1–10.
   Google Scholar
• Haeckel, E. 1866. Generelle Morphologie der Organismen; Allgemeine Grundzüge der organischen Formen-Wissenschaft, mechanisch begründet durch die von Charles Darwin reformirte Deszendenz-Theorie. Reimer, Berlin, 475 pp.
   Google Scholar
• Hahn, G. & Hahn, R. 1983. Multituberculata. Pp. 1–409 in F. Westphal (ed.) Fossilium Catalogus. I: Animalia. Pars 127. Kugler Publications, Amsterdam.
   Google Scholar
• Halliday, T. J. D., Upchurch, P. & Goswami, A. 2015. Resolving the relationships of Paleocene placental mammals. Biological Reviews DOI: 10.1111/brv.12242.
   PubMed Web of Science ®Google Scholar
• Hooker, J. J. 1996a. Mammalian biostratigraphy across the Paleocene–Eocene boundary in the Paris, London and Belgian basins. Geological Society of London Special Publication, 101, 205–218.
   Google Scholar
• Hooker, J. J. 1996b. A primitive emballonurid bat (Chiroptera, Mammalia) from the earliest Eocene of England. Palaeovertebrata, 25, 287–300.
   Google Scholar
• Hooker, J. J. 2010. The mammal fauna of the early Eocene Blackheath Formation of Abbey Wood, London. Monographs of the Palaeontographical Society, 165, 1–162.
   Google Scholar
• Hooker, J. J. 2013. Origin and evolution of the Pseudorhyncocyonidae, a European Paleogene family of insectivorous placental mammals. Palaeontology, 56, 807–835.
   Web of Science ®Google Scholar
• Hooker, J. J. 2015. A two-phase Mammalian Dispersal Event across the Paleocene–Eocene transition. Newsletters on Stratigraphy, 48, 201–220.
   Web of Science ®Google Scholar
• Hooker, J. J. & Millbank, C. 2001. A Cernaysian mammal from the Upnor Formation (Late Palaeocene, Herne Bay, UK) and its implications for correlation. Proceedings of the Geologists' Association, 112, 331–338.
   Web of Science ®Google Scholar
• Hooker, J. J. & Russell, D. E. 2012. Early Palaeogene Louisinidae (Macroscelidea, Mammalia), their relationships and north European diversity. Zoological Journal of the Linnean Society, 164, 856–936.
   Web of Science ®Google Scholar
• Hooker, J. J., Russell, D. E. & Phélizon, A. 1999. A new family of Plesiadapiformes (Mammalia) from the old world Lower Paleogene. Palaeontology, 42, 377–407.
   Web of Science ®Google Scholar
• Huxley, T. H. 1880. On the application of the laws of evolution to the arrangement of the Vertebrata and more particularly of the Mammalia. Proceedings of the Zoological Society of London, 43, 649–662.
   Google Scholar
• Janis, C. M., Archibald, J. D., Cifelli, R. L., Lucas, S. G., Schaff, C. R., Schoch, R. M. & Williamson, T. E. 1998a. Archaic ungulates and ungulatelike mammals. Pp. 247–259 in C. M. Janis, K. M. Scott & L. L. Jacobs (eds) Evolution of Tertiary Mammals of North America. Volume 1: Terrestrial carnivores, ungulates and ungulate like mammals. Cambridge University Press, Cambridge.
   Google Scholar
• Janis, C. M., Scott, K. M. & Jacobs, L. L. (eds) 1998b. Evolution of Tertiary Mammals of North America. Volume 1: Terrestrial carnivores, ungulates and ungulate like mammals. Cambridge University Press, Cambridge, 703 pp.
   Google Scholar
• Janis, C. M., Gunnell, G. F. & Uhen, M. D. (eds) 2008. Evolution of Tertiary Mammals of North America. Volume 2: Small mammals, xenarthrans and marine mammals. Cambridge University Press, New York, 795 pp.
   Google Scholar
• Jehle, M., Godinot, M., Delsate, D., Phélizon, A. & Pellouin, J.-L. 2012. A new late Paleocene micromammal fauna from Montchenot (Paris Basin). Preliminary results. Palaeobiodiversity and Palaeoenvironments, 92, 487–496.
   Google Scholar
• Kielan-Jaworowska, Z. & Hurum, J. H. 2001. Phylogeny and systematics of multituberculate mammals. Palaeontology, 44, 389–429.
   Web of Science ®Google Scholar
• Kielan-Jaworowska, Z., Cifelli, R. L. & Luo, Z.-X. 2004. Mammals from the age of dinosaurs. Origins, evolution, and structure. Columbia University Press, New York, 630 pp.
   Google Scholar
• King, S. 2016. A revised correlation of Tertiary rocks in the British Isles and adjacent areas of NW Europe. Chapter 14, East Anglia and London Basin (Paleocene and Eocene). Geological Society of London Special Report, 27, 329–375.
   Google Scholar
• Krause, D. W. 1982. Jaw movement, dental function, and diet in the Paleocene multituberculate Ptilodus. Paleobiology, 8, 265–281.
   Web of Science ®Google Scholar
• Kretzoi, M. & Kretzoi, M. 2000. Index generum et subgenerum mammalium. Fossilium Catalogus Animalia, 137, 1–433.
   Google Scholar
• Lemoine, V. 1883. Sur l’Adapisorex, un nouveau genre de mammifère de la faune cernaysienne des environs de Reims. Comptes Rendus de l'Académie des Sciences de Paris, 97, 1325–1327.
   Google Scholar
• Lemoine, V. 1885. Etude sur quelques petits mammifères de la faune cernaysienne des environs de Reims. Bulletins de la Société Géologique de France, 13, 203–217.
   Google Scholar
• Lillegraven, J. A. 1969. Latest Cretaceous mammals of upper part of Edmonton Formation of Alberta, Canada, and review of Marsupial-Placental dichotomy in mammalian evolution. The University of Kansas Paleontological Contributions, 50 (Vertebrata 12), 1–122.
   Google Scholar
• Linnaeus, C. 1758. Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata, 824 pp.
   Google Scholar
• Linnaeus, C. 1766. Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio duodecima, reformata, 532 pp.
   Google Scholar
• Lofgren, D. L., Lillegraven, J. A., Clemens, W. A., Gingerich, P. D. & Williamson, T. E. 2004. Paleocene biochronology: the Puercan through Clarkforkian Land Mammal Ages. Pp. 43–105 in M. O. Woodburne (ed.) Late Cretaceous and Cenozoic mammals of North America, biostratigraphy and geochronology. Columbia University Press, New York.
   Google Scholar
• López-Martínez, N. & Peláez-Campomanes, P. 1999. New mammals from the south-central Pyrenees (Tremp Formation, Spain) and their bearing on late Paleocene marine-continental correlations. Bulletins de la Société Géologique de France, 170, 681–696.
   Web of Science ®Google Scholar
• López-Martínez, N., Smith, R., Peláez-Campomanes, P. & Smith, T. 2006. The acme of the micromammal Paschatherium across the Paleocene-Eocene boundary in continental Europe. Micropaleontology, 3, 267–280.
   Web of Science ®Google Scholar
• Marsh, O. C. 1889. Discovery of cretaceous mammalia. American Journal of Science, 38, 81–92.
   Google Scholar
• Marshall, L. G., De Muizon, C. & Sigé, B. 1983. Late Cretaceous mammals (Marsupialia) from Bolivia. Géobios, 16, 739–745.
   Web of Science ®Google Scholar
• Matthew, W. D. 1918. A revision of the Lower Eocene Wasatch and Wind River faunas. Part V. Insectivora (Continued), Glires, Edentata. Bulletin of the American Museum of Natural History, 38, 429–483.
   Google Scholar
• Matthew, W. D. 1937. Paleocene faunas of the San Juan Basin, New Mexico. Transactions of the American Philosophical Society, 30, 1–510.
   Google Scholar
• Matthew, W. D. & Granger, W. 1921. New genera of Paleocene mammals. American Museum Novitates, 13, 1–7.
   Google Scholar
• McKenna, M. C. 1975. Toward a phylogenetic classification of the Mammalia. Pp. 21–46 in W. P. Luckett & F. S. Szalay (eds.) Phylogeny of the Primates. Plenum Press, New York.
   Google Scholar
• Matthews, S. C. 1973. Notes on open nomenclature and on synonymy lists. Palaeontology, 16, 713–719.
   Google Scholar
• McKenna, M. C. & Bell, S. K. 1997. Classification of mammals above the species level. Columbia University Press, New York. 631 pp.
   Google Scholar
• Meng J., Hu, Y.-M. & Li, C.-K. 2003. The osteology of Rhombomylus (Mammalia, Glires): Implications for phylogeny and evolution of Glires. Bulletin of the American Museum of National History, 275, 1–247.
   Google Scholar
• Middleton, M. D. & Dewar, E. W. 2004. New mammals from the Early Paleocene Littleton Fauna (Denver Formation, Colorado). New Mexico Museum of Natural History and Science Bulletin, 26, 21–31.
   Google Scholar
• Missiaen, P. & Smith, T. 2008. The Gashatan (late Paleocene) mammal fauna from Subeng, Inner Mongolia, China. Acta Palaeontologica Polonica, 53, 357–378.
   Web of Science ®Google Scholar
• Missiaen, P. 2011. An updated mammalian biochronology and biogeography for the early Paleogene of Asia. Vertebrata Palasiatica, 49, 29–52.
   Google Scholar
• Muizon, C. de & Cifelli, R. L. 2000. The ‘condylarths’ (archaic Ungulata, Mammalia) from the early Palaeocene of Tiupampa (Bolivia): implications on the origin of the South American ungulates. Geodiversitas, 22, 47–150.
   Google Scholar
• Peláez-Campomanes, P., López-Martínez, N., Álvarez-Sierra, M. A. & Daams, R. 2000. The earliest mammal of the European Paleocene: the multituberculate Hainina. Journal of Paleontology, 74, 701–711.
   Web of Science ®Google Scholar
• Piton, L.-E. 1940. Paléontologie du gisement éocène de Menat (Puy-de-Dôme). Vallier, Clermont-Ferrand, 303 pp.
   Google Scholar
• Pomerol, C. 1989. Stratigraphy of the Palaeogene: hiatuses and transitions. Proceedings of the Geologists' Association, 100, 313–324.
   Google Scholar
• Prasad, G. V. R., Jaeger, J. J., Sahni, A., Gheerbrant, E. & Khajuria, C. 1994. Eutherian mammals from the Upper Cretaceous (Maastrichtian) Intertrappean Beds of Naskal, Andra Pradesh, India. Journal of Vertebrate Paleontology, 14, 260–277.
   Web of Science ®Google Scholar
• Prasad, G. V. R., Verma, O., Gheerbrant, E., Goswami, A., Khosla, A., Parmar, V. & Sahni, A. 2010. First mammal evidence from Late Cretaceous of India for biotic dispersal between India and Africa at the KT transition. Comptes Rendus Palevol, 9, 63–71.
   Web of Science ®Google Scholar
• Prothero, D. R., Manning, E. M. & Fischer, M. 1988. The phylogeny of the Ungulates. Pp. 173–181 in M. J. Benton (ed.) The phylogeny and classification of the tetrapods, Volume 2: Mammals. Springer-Verlag, New York.
   Google Scholar
• Rădulescu, C. & Samson, P.-M. 1996. The first multituberculate skull from the Late Cretaceous (Maastrichtian) of Europe (Hateg basin, Rumania). Anuarul Institutului Geologic al Romaniei, 69, 177–178.
   Google Scholar
• Rose, K. D. 1981a. The Clarkforkian land-mammal age and mammalian faunal composition across the Paleocene–Eocene boundary. University of Michigan Papers on Paleontology, 26, 1–197.
   Google Scholar
• Rose, K. D. 1981b. Composition and species diversity in Paleocene and Eocene mammal assemblages: an empirical study. Journal of Vertebrate Paleontology, 1, 367–388.
   Google Scholar
• Rose, K. D. 2006. The beginning of the age of mammals. The Johns Hopkins University Press, Baltimore, 428 pp.
   Google Scholar
• Rose, K. D. & Bown, T. M. 1982. New Plesiadapiform Primates from the Eocene of Wyoming and Montana. Journal of Vertebrate Paleontology, 2, 63–69.
   Google Scholar
• Russell, D. E. 1964. Les mammifères paléocènes d'Europe. Éditions du Muséum, Université de Paris, Paris, 324 pp.
   Google Scholar
• Russell, D. E. 1975. Paleoecology of the Paleocene–Eocene Transition in Europe. Contributions to Primatology, 5, 28–61.
   PubMedGoogle Scholar
• Russell, D. E. 1982. Tetrapods of the Northwest European Tertiary Basin. Geologisches Jahrbuch, 60, 5–74.
   Google Scholar
• Russell, D. E. & Dashzeveg, D. 1986. Early Eocene insectivores (Mammalia) from the People's Republic of Mongolia. Palaeontology, 29, 269–291.
   Web of Science ®Google Scholar
• Savage, D. E. & Russell, D. E. 1983. Mammalian paleofaunas of the world. Addison-Wesley, New York, 427 pp.
   Google Scholar
• Schmidt-Kittler, N. (ed.) 1987. European levels and correlation tables. Münchner Geowissenschaftliche, Abhandlungen A, 10, 15–31.
   Google Scholar
• Scotese, C. R. 2014. The PALEOMAP Project PaleoAtlas for ArcGIS, version 2, Volume 1, Cenozoic Plate Tectonic, Paleogeographic, and Paleoclimatic Reconstructions, Maps 1-15. PALEOMAP Project, Evanston.
   Google Scholar
• Scott, W. B. 1892. A Revision of the North American Creodonta with notes on some genera which have been referred to that group. Proceedings of the Academy of Natural Sciences of Philadelphia, 44, 291–323.
   Google Scholar
• Scott, C. S. & Fox, R. C. 2005. Windows on the evolution of Picrodus (Plesiadapiformes: Primates): Morphology and relationships of a species complex from the Paleocene of Alberta. Journal of Paleontology, 79, 635–657.
   Web of Science ®Google Scholar
• Seiffert, E. R. 2010. The oldest and youngest records of afrosoricid placentals from the Fayum Depression of northern Egypt. Acta Palaeontologica Polonica, 55, 599–616.
   Web of Science ®Google Scholar
• Sigé, B. 1974. Pseudorhyncocyon cayluxi Filhol, 1892. Insectivore géant des Phosphorites du Quercy. Palaeovertebrata, 6, 33–46.
   Google Scholar
• Sigé, B. & Marandat, B. 1997. Apport à la faune du Paléocène inférieur d'Europe: un plésiadapiforme du Montien de Hainin (Belgique). Mémoires et Travaux de l'E.P.H.E., Institut de Montpellier, 21, 679–686.
   Google Scholar
• Silcox, M. T. & Gunnell, G. F. 2007. Plesiadapiformes. Pp. 207–238 in C. M. Janis, G. F. Gunnell & M. D. Uhen (eds) Evolution of Tertiary mammals of North America. Volume 2: Small mammals, xenarthrans and marine mammals. Cambridge University Press, New York.
   Google Scholar
• Silcox M. T., Bloch J. I., Sargis E. J. & Boyer D. M. 2005. Euarchonta (Dermoptera, Scandentia, Primates). Pp 127–144 in K. D. Rose & J. D. Archibald (eds) The rise of placental mammals: origins and relationships of the major extant clades. Johns Hopkins University Press, Baltimore.
   Google Scholar
• Silcox, M. T., Bloch, J. I., Boyer, D. M. & Houde, P. 2010. Cranial anatomy of Paleocene and Eocene Labidolemur kayi (Mammalia: Apatotheria), and the relationships of the Apatemyidae to other mammals. Zoological Journal of the Linnean Society, 160, 773–825.
   Web of Science ®Google Scholar
• Simons, E. L. 1972. Primate Evolution: An Introduction to Man's Place in Nature. The MacMillan Series in Physical Anthropology, MacMillan, New York, 322 pp.
   Google Scholar
• Simpson, G. G. 1935. New Paleocene mammals from the Fort Union of Montana. Proceedings of the United States National Museum, 83, 221–244.
   Google Scholar
• Simpson, G. G. 1937. The Fort Union of the Crazy Mountain Field, Montana and its mammalian faunas. United States National Museum Bulletin, 169, 1–287.
   Google Scholar
• Sloan, R. E. & Van Valen, L. 1965. Cretaceous mammals from Montana. Science, 148, 220–227.
   PubMed Web of Science ®Google Scholar
• Smith, T. 2013. Contribution of Asia to the evolution and paleobiogeography of the earliest modern mammals. Bulletin des Séances de l'Académie Royale des Sciences d'Outre-Mer, 57, 293–305.
   Google Scholar
• Smith T. & Codrea V. 2015. Red Iron-Pigmented Tooth Enamel in a Multituberculate Mammal from the Late Cretaceous Transylvanian “Haţeg Island”. PLoS ONE, 10(7), e0132550.
   PubMed Web of Science ®Google Scholar
• Smith, T. & Smith, R. 2003. Terrestrial mammals as biostratigraphic indicators in upper Paleocene-lower Eocene marine deposits of the southern North Sea Basin. Geological Society of America, Special Paper, 369, 513–520.
   Google Scholar
• Smith, T., De Bast, E. & Sigé, B. 2010. Euarchontan affinity for the Paleocene Afro-European adapisoriculid mammals and their origin in the late Cretaceous Deccan Traps of India. Naturwissenschaften, 97, 417–422.
   PubMed Web of Science ®Google Scholar
• Smith, T., Quesnel, F., De Plöeg, G., De Franceschi, D., Métais, G., De Bast, E., Solé, F., Folie, A., Boura, A., Claude, J., Dupuis, C., Gagnaison, C., Iakovleva, A., Martin, J., Maubert, F., Prieur, J., Roche, E., Storme, J.-Y., Thomas, R., Tong, H., Yans, J. & Buffetaut, E. 2014. First Clarkforkian Equivalent Land Mammal Age in the Latest Paleocene Basal Sparnacian Facies of Europe: Fauna, Flora, Paleoenvironment and (Bio)stratigraphy. PLoS One, 9(1), e86229.
   PubMed Web of Science ®Google Scholar
• Smith, T., Rose, K. & Gingerich, P. 2006. Rapid Asia-Europe-North America geographic dispersal of earliest Eocene primate Teilhardina during the Paleocene–Eocene Thermal Maximum. Proceedings of the National Academy of Sciences, 103, 11223–11227.
   PubMed Web of Science ®Google Scholar
• Smith, T., Van Itterbeek, J. & Missiaen, P. 2004. Oldest Plesiadapiform (Mammalia, Proprimates) from Asia and its palaeobiogeographical implications for faunal interchange with North America. Comptes rendus Palevol, 3, 43–52.
   Web of Science ®Google Scholar
• Solé, F. & Smith, T. 2013. Dispersals of placental carnivorous mammals (Carnivoramorpha, Oxyaenodonta & Hyaenodontida) near the Paleocene–Eocene boundary: a climatic and almost worldwide story. Geologica Belgica, 16, 254–261.
   Web of Science ®Google Scholar
• Solé, F., Smith, T., De Bast, E., Codrea, V. & Gheerbrant, E. 2016. New carnivoraforms from the latest Paleocene of Europe and their bearing on the origin and radiation of Carnivoraformes (Carnivoramorpha, Mammalia) Journal of Vertebrate Paleontology. DOI: 10.1080/02724634.2016.1082480
   PubMed Web of Science ®Google Scholar
• Steurbaut, E. 1998. High-resolution holostratigraphy of Middle Paleocene to Early Eocene strata in Belgium and adjacent areas. Palaeontographica, Abteilung A, 247, 91–156.
   Google Scholar
• Steurbaut, E. 2006. Ypresian. Geologica belgica 9, 73–93.
   Google Scholar
• Steurbaut, E. & Sztrákos, K. 2008. Danian/Selandian boundary criteria and North Sea Basin-Tethys correlations based on calcareous nannofossil and foraminiferal trends in SW France. Marine Micropaleontology, 67, 1–29.
   Web of Science ®Google Scholar
• Storch, G. 2008. Skeletal remains of a diminutive primate from the Paleocene of Germany. Naturwissenschaften, 95, 927–930.
   PubMed Web of Science ®Google Scholar
• Storch, G. & Lister, A. M. 1985. Leptictidium nasutum, ein Pseudorhyncocyonide aus dem Eozän der “Grube Messel” bei Darmstadt (Mammalia, Proteutheria). Senckenbergia lethaea, 66, 1–37.
   Google Scholar
• Sudre, J. & Russell, D. E. 1982. Les mammifères montiens de Hainin (Paléocène moyen de Belgique). Part II: Les Condylarthres. Palaeovertebrata, 12, 173–184.
   Google Scholar
• Szalay, F. S. 1968. The Picrodontidae, a Family of Early Primates. American Museum Novitates, 2329, 1–55.
   Google Scholar
• Tabuce, R., Coiffait, B., Coiffait, P.-E., Mahboubi, M. & Jaeger, J.-J. 2001. A new genus of Macroscelidea (Mammalia) from the Eocene of Algeria: a possible origin for elephant shrews. Journal of Vertebrate Paleontology, 21, 535–546.
   Web of Science ®Google Scholar
• Tabuce, R., Telles Antunes, M., Smith, R. & Smith, T. 2006. Dental variability and tarsal morphology of the European Paleocene/Eocene “condylarth” mammal Microhyus. Acta Palaeontologica Polonica, 51, 37–52.
   Web of Science ®Google Scholar
• Taverne, L., Nolf, D. & Folie, A. 2007. On the presence of the osteoglossid fish genus Scleropages (Teleostei, Osteoglossiformes) in the continental Paleocene of Hainin (Mons Basin, Belgium). Belgian Journal of Zoology, 137, 89–97.
   Web of Science ®Google Scholar
• Thaler, L. 1977. État des recherches sur la faune de Mammifères du Montien de Hainin (Belgique). Géobios, Mémoire Spéciale, 1, 57–58.
   Google Scholar
• Ting, S.-Y., Tong, Y.-S., Clyde, W. C., Koch, P. L., Meng, J., Wang, Y.-Q., Bowen, G. J., Li, Q. & Snell, K. E. 2011. Asian Early Paleogene chronology and mammalian faunal turnover events. Vertebrata Palasiatica, 49, 1–28.
   Google Scholar
• Trouessart, E.-L. 1897. Catalogus mammalium tam viventium quam Fossilium, Vol. 1, R. Friedlander und Sohn, Berlin, 664 pp.
   Google Scholar
• Vandenberghe, N., Hilgen, F. J. & Speijer, R. P. with contributions by Ogg, J. G., Gradstein, F. M., Hammer, O., Hollis, C. J. & Hooker, J. J. 2012. The Paleogene Period. Pp. 856–907 in F. M. Gradstein, J. G. Ogg, M. Schmitz & G. Ogg (eds) The geologic time scale. Elsevier, Amsterdam.
   Google Scholar
• Van Valen, L. 1966. Deltatheridia, a new order of mammals. Bulletin of the American Museum of Natural History, 132, 1–126.
   Google Scholar
• Van Valen, L. 1967. New Paleocene insectivores and insectivore classification. Bulletin of the American Museum of Natural History, 135, 221–284.
   Google Scholar
• Van Valen, L. 1978. The beginning of the Age of Mammals. Evolutionary Theory, 4, 45–80.
   Google Scholar
• Vianey-Liaud, M. 1979. Les mammifères montiens de Hainin (Paléocène moyen de Belgique). Part I: Multituberculés. Palaeovertebrata, 9, 117–131.
   Google Scholar
• Vianey-Liaud, M. 1986. Les Multituberculés thanétiens de France, et leurs rapports avec les Multituberculés Nord-Américains. Palaeontographica, Abteilung A, 191, 85–171.
   Google Scholar
• Waddell, P. J., Okada, N. & Hasegawa, M. 1999. Towards resolving the interordinal relationships of placental mammals. Systematic Biology, 48, 1–5.
   PubMed Web of Science ®Google Scholar
• Wang, Y., Meng, J., Ni, X. & Li, C. 2007. Major events of Paleogene mammal radiation in China. Geological Journal 42, 415–430.
   Web of Science ®Google Scholar
• Weigelt, J. 1939. Die Aufdeckung der bisher ältesten tertiären Säugtierfauna Deutschlands. Nova Acta Leopoldina, Neue Folge, 7, 1–13.
   Google Scholar
• Wible, J. R., Rougier, G. W., Novacek, M. J. & Asher, R. J. 2009. The eutherian mammal Maelestes gobiensis from the Late Cretaceous of Mongolia and the phylogeny of Cretaceous Eutheria. Bulletin of the American Museum of Natural History, 327, 1–123.
   Web of Science ®Google Scholar
• Williamson, T. E. 1996. The beginning of the age of mammals in the San Juan Basin: Biostratigraphy and evolution of Paleocene mammals of the Nacimiento Formation. New Mexico Museum of Natural History Bulletin, 8, 1–141.
   Google Scholar
• Williamson, T. E., Weil, A. & Standhardt, B. 2011. Cimolestids (Mammalia) from the Early Paleocene (Puercan) of New Mexico. Journal of Vertebrate Paleontology, 31, 162–180.
   Web of Science ®Google Scholar
• Wilson, R. W. 1985. The dentition of the Paleocene “insectivore” genus Acmeodon Matthew & Granger (?Palaeoryctidae, Mammalia). Journal of Paleontology, 59, 713–720.
   Web of Science ®Google Scholar
• Winge, H. 1917. Udsigt over Insektaedernes indbyrdes Slaegtskab. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening Copenhagen, 68, 83–203.
   Google Scholar
• Woodburne, M. O. 2004. Global Events and the North American Mammalian Biochronology. Pp 315–343 in M. O. Woodburne (ed.) Late Cretaceous and Cenozoic mammals of North America. Columbia University Press, New York.
   Google Scholar
• Woodburne, M. O. 2007. Mammal Ages. Stratigraphy, 3, 229–261.
   Google Scholar
• Wu, S., Wu, W., Zhang, F., Ye, J., Ni, X., Sun, J., Edwards, S. V. & Meng, J. 2012. Molecular and Paleontological Evidence for a Post-Cretaceous Origin of Rodents. PLoS One, 7(10), e46445.
   PubMed Web of Science ®Google Scholar
• Yans, J., Amaghzaz, M., Bouya, B., Cappetta, H., Iacumin, P., Kocsis, L., Mouflih, M., Selloum, O., Sen, S., Storme, J.-Y. & Gheerbrant, E. 2014. First carbon isotope chemostratigraphy of the Ouled Abdoun phosphate Basin, Morocco; implications for dating and evolution of earliest African placental mammals. Gondwana Research, 25, 257–269.
   Web of Science ®Google Scholar
• Zachos, J., Pagani, M., Sloan, L., Thomas, E. & Billups, K. 2001. Trends, rhythms, and aberrations in Global Climate 65 Ma to Present. Science, 292, 686–693.
   PubMed Web of Science ®Google Scholar
• Zachos, J., McCarren, H., Murphy, B., Röhl, U. & Westerhold, T. 2010. Tempo and scale of late Paleocene and early Eocene carbon isotope cycles: Implications for the origin of hyperthermals. Earth and Planetary Science Letters, 299, 242–249.
   Web of Science ®Google Scholar
• Zack, S. P., Penkrot, T. A., Bloch, J. I. & Rose, K. D. 2005. Affinities of “hyopsodontids” to elephant shrews and a Holarctic origin of Afrotheria. Nature, 434, 497–501.
   PubMed Web of Science ®Google Scholar
• Zittel, K. A. von. 1892. Handbuch der Paläontologie. 1 Abteilung. Palæozoologie, Band 4, Mammalia. Druck und Verlag von R. Oldenbourg, München and Leipzig, 799 pp.
   Google Scholar