New camerate crinoid genera from the Upper Ordovician (Katian) of Estonia: evolutionary origin of family Opsiocrinidae and a phylogenetic assessment of Ordovician Monobathrida

References

• Ainsaar, L. & Meidla, T. 2001. Facies and stratigraphy of the middle Caradoc mixed siliciclastic-carbonate sediments in eastern Baltoscandia. Proceedings of the Estonian Academy of Sciences, Geology, 50, 5–23.
   Google Scholar
• Ausich, W. I. 1985. New crinoids and revision of Superfamily Glyptocrinacea (early Silurian, Ohio). Journal of Paleontology, 59, 793–808.
   Web of Science ®Google Scholar
• Ausich, W. I. 1986. Early Silurian rhodocrinitacean crinoids (Brassfield Formation, Ohio). Journal of Paleontology, 60, 84–106.
   Web of Science ®Google Scholar
• Ausich, W. I. 1996. Crinoid plate circlet homologies. Journal of Paleontology, 70, 955–964.
   Web of Science ®Google Scholar
• Ausich, W. I. 1998a. Early phylogeny and subclass division of the Crinoidea (Phylum Echinodermata). Journal of Paleontology, 72, 499–510.
   Web of Science ®Google Scholar
• Ausich, W. I. 1998b. Phylogeny of Arenig to Caradoc crinoids (Phylum Echinodermata) and suprageneric classification of the Crinoidea. The University of Kansas Paleontological Contributions, 9, 1–36.
   Google Scholar
• Ausich, W. I. & Wilson, M. A. 2016. Llandovery (early Silurian) crinoids from Hiiumaa Island, western Estonia. Journal of Paleontology, 90, 1138–1147.
   Web of Science ®Google Scholar
• Ausich, W. I., Wilson, M. A. & Vinn, O. 2012. Crinoids from the Silurian of western Estonia. Acta Palaeontologica Polonica, 57, 613–631.
   Web of Science ®Google Scholar
• Ausich, W. I., Kammer, T. W., Rhenberg, E. C. & Wright, D. F. 2015. Early phylogeny of crinoids within the pelmatozoan clade. Palaeontology, 58, 937–952.
   Web of Science ®Google Scholar
• Ausich, W. I., Wilson, M. A. & Vinn, O. 2015. Wenlock and Pridoli (Silurian) crinoids from Saaremaa, western Estonia (Phylum Echinodermata). Journal of Paleontology, 89, 72–81.
   Web of Science ®Google Scholar
• Bergström, S. M., Chen, X., Gutiérrez-Marco, J. C. & Dronov, A. 2009. The new chronostratigraphic classification of the Ordovician System and its relations to major regional series and stages and to δ13C chemostratigraphy. Lethaia, 42, 97–107.
   Web of Science ®Google Scholar
• Bergström, S. M., Schmitz, B., Saltzman, M. R. & Huff, W. D. 2010. The Upper Ordovician Guttenberg δ13C excursion (GICE) in North America and Baltoscandia: occurrence, chronostratigraphic significance, and paleoenvironmental relationships. Geological Society of America, Special Papers, 466, 37–67.
   Google Scholar
• Cocks, L. R. M. & Torsvik, T. H. 2005. Baltica from the late Precambrian to mid-Palaeozoic times: the gain and loss of a terrane's identity. Earth-Science Reviews, 72, 39–66.
   Web of Science ®Google Scholar
• Cole, S. R. 2017. Phylogeny and morphologic evolution of the Ordovician Camerata (Class Crinoidea, Phylum Echinodermata). Journal of Paleontology, 91, 815–828.
   Web of Science ®Google Scholar
• Cole, S. R. & Ausich, W. I. 2015. Phylogenetic analysis of the Ordovician Diplobathrida (Subclass Camerata, Class Crinoidea): implications for early camerate evolution. Cuadernos del museo Geominero, 19, 41–44.
   Google Scholar
• Cole, S. R., Ausich, W. I., Colmenar, J. & Zamora, S. 2017. Filling the Gondwanan gap: diverse crinoids from the Castillejo and Fombuena formations (Middle and Upper Ordovician, Iberian Chains, Spain). Journal of Paleontology, 91, 715–734.
   Web of Science ®Google Scholar
• Deline, B. & Ausich, W. I. 2011. Testing the plateau: a reexamination of disparity and morphologic constraints in early Paleozoic crinoids. Paleobiology, 37, 214–236.
   Web of Science ®Google Scholar
• Donovan, S. K. & Cope, J. C. W. 1989. A new camerate crinoid from the Arenig of south Wales. Palaeontology, 32, 101–107.
   Web of Science ®Google Scholar
• Dronov, A. & Holmer, L. E. 1999. Depositional sequences in the Ordovician of Baltoscandia. Acta Universitatis Carolinae Geologica, 43, 133–136.
   Google Scholar
• Dronov, A. V., Ainsaar, L., Kaljo, D., Meidla, T., Saadre, T. & Einasto, R. 2011. Ordovician of Baltoscandia: facies, sequences and sea-level changes. Cuadernos del Museo Geominero, 14, 143–150.
   Google Scholar
• Foote, M. 1992. Paleozoic record of morphological diversity in blastozoan echinoderms. Proceedings of the National Academy of Sciences, 89, 7325–7329.
   PubMed Web of Science ®Google Scholar
• Foote, M. 1994. Morphological disparity in Ordovician–Devonian crinoids and the early saturation of morphological space. Paleobiology, 20, 320–344.
   Web of Science ®Google Scholar
• Foote, M. 1997. The evolution of morphological disparity. Annual Review of Ecology and Systematics, 28, 129–152.
   Google Scholar
• Foote, M. 1999. Morphological diversity in the evolutionary radiation of Paleozoic and post-Paleozoic crinoids. Paleobiology, 25, 1–115.
   Web of Science ®Google Scholar
• Gould, S. J. 1989. Wonderful Life. Norton, New York, 347 pp.
   Google Scholar
• Hatch, J. R., Jacobson, S. R., Witzke, B. J., Risatti, J. B., Anders, D. E., Wattney, W. L., Newell, K. D. & Vuletich, A. K. 1987. Possible late Middle Ordovician organic carbon isotope excursion: evidence from Ordovician oils and hydrocarbon source rocks, mid-continent and east-central United States. American Association of Petroleum Geologists, Bulletin, 71, 1342–1354.
   Web of Science ®Google Scholar
• Hecker, R. F. 1958. New data on the genus Achradocystites (Echinodermata, Paracrinoidea). Trudy Instituta geologii AN ESSR. 3, 145–162. [ In Russian, with Germany summary].
   Google Scholar
• Hints, L. 1998. Oandu Stage (Caradoc) in central North Estonia. Proceedings of the Estonian Academy of Sciences, Geology, 47, 158–172.
   Google Scholar
• Hints, L. & Meidla, T. 1997. Pirgu Stage. Pp. 234–241 in A. Raukas & A. Teedumäe (eds) Geology and mineral resources of Estonia. Estonian Academy Publishers, Tallinn.
   Google Scholar
• Jaekel, O. 1899. Stammesgeschichte der Pelmatozoen. J. Springer, Berlin, 441 pp., 18 pls.
   Google Scholar
• Jaekel, O. 1918. Phylogenie und System der Pelmatozoen. Paläeontologische Zeitschrift, 3, 1–128.
   Google Scholar
• Jell, P. A. 1999. Silurian and Devonian crinoids from central Victoria. Memoirs of the Queensland Museum, 43, 1–114.
   Google Scholar
• Kaljo, D. 2004. Diversity of late Ordovician rugose corals in Baltoscandia: role of environmental changes and comparison with other areas. Proceedings of the Estonian Academy of Sciences, Geology, 53, 233–245.
   Google Scholar
• Kammer, T. W., Sumrall, C. D., Zamora, S., Ausich, W. I. & Deline, B. 2013. Oral region homologies in Paleozoic crinoids and other plesiomorphic pentaradial echinoderms. PLoS ONE, 8, 1–16.
   Web of Science ®Google Scholar
• Kesling, R. V. 1967. Cystoids. Pp. S85–S267 in R. C. Moore (ed.) Treatise on invertebrate paleontology, Part S. Echinodermata 1. Geological Society of America and University of Kansas Press, Lawrence.
   Google Scholar
• Kesling, R. V. 1968. Ameliacrinus benderi, a new dicyclic camerate crinoid from the Middle Devonian Silica Formation in northwestern Ohio. University of Michigan Museum of Paleontology Contribution, 22, 155–162.
   Google Scholar
• Kesling, R. V. & Meyer, D. L. 1963. The crinoid Opsiocrinus mariae Kier in the Bell Shale of Michigan. University of Michigan Museum of Paleontology Contribution, 18, 177–184.
   Google Scholar
• Kier, P. M. 1952. Echinoderms of the middle Devonian Silica Formation of Ohio. University of Michigan Museum of Paleontology Contribution, 10, 59–81.
   Google Scholar
• Kier, P. M. 1958. Infrabasals in the crinoid Opsiocrinus Kier. University of Michigan Museum of Paleontology Contribution, 14, 201–206.
   Google Scholar
• Kröger, B., Hints, L. & Lehnert, O. 2014a. Age, facies, and geometry of the Sandbian/Katian (Upper Ordovician) pelmatozoan-bryozoan-receptaculitid reefs of the Vasalemma Formation, northern Estonia. Facies, 60, 963–986.
   Web of Science ®Google Scholar
• Kröger, B., Hints, L. & Lehnert, O. 2017. Ordovician reef and mound evolution: the Baltoscandian picture. Geological Magazine, 154, 683–706.
   Web of Science ®Google Scholar
• Kröger, B., Hints, L., Lehnert, O., Männik, P. & Joachimski, M. 2014b. The early Katian (Late Ordovician) reefs near Saku, northern Estonia and the age of the Saku Member, Vasalemma Formation. Estonian Journal of Earth Sciences, 63, 271–276.
   Web of Science ®Google Scholar
• Männil, R. 1960. Stratigrafia oanduskogo (‘Vasalemmaskogo’) gorisonta. Trudy Instituta Geologii Akademii Nauk Estonskoi SSR, 5, 89–122. [In Russian, with English summary].
   Google Scholar
• Meidla, T., Ainsaar, L., Hints, L., Hints, O., Martma, T. & Nõlvak, J. 1999. The mid-Caradocian biotic and isotopic event in the Ordovician of the East Baltic. Acta Universitatis Carolinae, Geologica, 43, 503–506.
   Google Scholar
• Miller, J. S. 1821. A natural history of the Crinoidea, or lily-shaped animals; with observations on the genera, Asteria, Euryale, Comatula and Marsupites. Bryan & Co., Bristol, 150 pp.
   Google Scholar
• Moore, R. C. 1952. Crinoids. Pp. 604–652 in R. C. Moore, C. G. Lalicker & A. G. Fischer (eds) Invertebrate fossils. McGraw-Hill, New York.
   Google Scholar
• Moore, R. C. & Laudon, L. R. 1943. Evolution and classification of Paleozoic crinoids. Geological Society of America Special Paper, 46, 1–154.
   Google Scholar
• Moore, R. C. & Teichert, C. 1978. Treatise on invertebrate paleontology, Part T. Echinodermata 2. Geological Society of America and University of Kansas Press, Lawrence, 1027 pp.
   Google Scholar
• Nestor, H. & Einasto, R. 1997. Ordovician and Silurian carbonate sedimentation basin. Pp. 192–204 in A. Raukas & A. Teedumäe (eds) Geology and mineral resources of Estonia. Estonian Academy Publishers, Tallinn, Estonia.
   Google Scholar
• Põlma, L., Sarv, L. & Hints, L. 1988. Lithology and fauna of the Caradoc Series type sections in North Estonia. Valgus, Tallinn, 101 pp. [In Russian, with English summary].
   Google Scholar
• Rõõmusoks, A. 1970. Stratigrafia Viruskoi i Harioskoi Serii (Ordovik) Severnoi Estonii. Valgus, Tallinn, 343 pp. [In Russian, with English summary].
   Google Scholar
• Schluter, D. 2000. The ecology of adaptive radiation. Oxford University Press, New York, 296 pp.
   Google Scholar
• Shavit, L., Penny, D., Hendy, M. D. & Holland, B. R. 2007. The problem of rooting rapid radiations. Molecular Biology and Evolution, 24, 2400–2411.
   PubMed Web of Science ®Google Scholar
• Simms, M. J. 1993. Reinterpretation of thecal plate homology and phylogeny in the Class Crinoidea. Lethaia, 26, 303–312.
   Web of Science ®Google Scholar
• Sprinkle, J. 1982. Acolocrinus. Pp. 111–118 in J. Sprinkle (ed.) Echinoderm faunas from the Bromide Formation (Middle Ordovician) of Oklahoma. The University of Kansas Paleontological Contributions, 1.
   Google Scholar
• Sprinkle, J. & Moore, R. C. 1978. Hybocrinida. Pp. T564–T574 in R. C. Moore & C. Teichert (eds) Treatise on invertebrate paleontology, Part T. Echinodermata 2. Geological Society of America and University of Kansas Press, Lawrence.
   Google Scholar
• Stukalina, G. A. 2000. Paleozoic crinoids. VSEGEI Press, St. Petersburg, 283 pp. [In Russian].
   Google Scholar
• Stukalina, G. A. & Hints, L. 1989. On the morphology and systematics of Achradocystites (Paracrinoidea). Pp. 58–72 in D. Kaljo (ed.) Fossil and recent echinoderm researches. Academy of Sciences of the USSR, Tallinn. [In Russian, with English summary].
   Google Scholar
• Swofford, D. L. 2003. PAUP* version 4.0.b10 Phylogenetic analysis using parsimony and other methods. Sinauer Associates, Sunderland.
   Google Scholar
• Torsvik, T. T. & Cocks, L. R. M. 2013. New global palaeogeographical reconstructions for the Early Palaeozoic and their generation. Pp. 5–24 in D. A. T. Harper & T. Servais (eds) Early Palaeozoic biogeography and palaeogeography. Geological Society of London, Memoir, 38.
   Google Scholar
• Ubaghs, G. 1978. General morphology. Pp. T58–T216 in R. C. Moore & C. Teichert (eds) Treatise on invertebrate paleontology, Part T. Echinodermata 2. Geological Society of America and University of Kansas Press, Lawrence.
   Google Scholar
• Volborth, A. 1870. Über Achradocystites und Cystoblastus, zwei neue Crinoideen – Gattungen, eingeleitet durch kritische Betrauchtungen uber die Organe der Cystideen. Mémoires de l'Académie Impériale des Sciences de St.-Pétersbourg, Serie 7, 16, 1–15.
   Google Scholar
• Wachsmuth, C. & Springer, F. 1885. Discussion of the classification and relations of the brachiate crinoids, and conclusions of the generic descriptions. Pp. 225–364 in Revision of the Palaeocrinoidea. Part 3, Section 1. Proceedings of the Academy of Natural Sciences of Philadelphia, Philadelphia.
   Google Scholar
• Webster, G. D., Waters, J. & Chen, X. 2009. Revision of the Chen and Yao Devonian to Permian crinoids from Western Yunnan. Palaeobiodiversity and Palaeoenvironments, 89, 119–160.
   Google Scholar
• Wright, D. F. 2015. Fossils, homology, and ‘phylogenetic paleo-ontogeny’: a reassessment of primary posterior plate homologies among fossil and living crinoids with insight from developmental biology. Paleobiology, 41, 570–591.
   Web of Science ®Google Scholar
• Wright, D. F. 2017. Bayesian estimation of fossil phylogenies and the evolution of early to middle Paleozoic crinoids (Echinodermata). Journal of Paleontology, 91, 799–814.
   Web of Science ®Google Scholar
• Wright, D. F. & Toom, U. 2017. New crinoids from the Baltic region (Estonia): fossil tip-dating phylogenetics constrains the origin and Ordovician–Silurian diversification of the Flexibilia (Echinodermata). Palaeontology, 60, 893–910.
   Web of Science ®Google Scholar
• Wright, D. F., Ausich, W. I., Cole, S. R., Rhenberg, E. C. & Peter, M. E. 2017. Phylogenetic taxonomy and classification of the Crinoidea (Echinodermata). Journal of Paleontology, 91, 829–846.
   Web of Science ®Google Scholar
• Zittel, K. A. von. 1879. Echinoderms. Pp. 308–560 in Handbuch der Palaeontologie, Volume 1, Palaeozoologie. R. Oldenbourg, Leipzig.
   Google Scholar