References
- American Public Health Association (APHA). 2000. Standard methods for the examination of Water and Wastewater, 20th ed. Washington, D.C: American Public Health Association, American Water Works Association, Water Environment Federation.
- Anderson, N. J., R. Harriman, D. B. Ryves, and S. T. Patrick. 2001. Dominant factors controlling variability in the ionic composition of West Greenland lakes. Arctic, Antarctic, and Alpine Research 33 (4):1–15. doi:https://doi.org/10.2307/1552551.
- Anderson, N. J., J. E. Saros, J. E. Bullard, S. M. Cahoon, S. McGowan, E. A. Bagshaw, C. D. Barry, R. Bindler, B. T. Burpee, J. L. Carrivick, et al. 2017. The Arctic in the twenty-first century: Changing biogeochemical linkages across a paraglacial landscape of Greenland. Bioscience 67 (2):118–33.
- Battarbee, R. W. 1986. Diatom analysis. In Handbook of Holocene palaeoecology and palaeohydrology, ed. B. E. Berglund, 527–70. New York: Wiley-Interscience.
- Bergström, A. K. 2010. The use of TN:TP and DIN:TP ratios as indicators for phytoplankton nutrient limitation in oligotrophic lakes affected by N deposition. Aquatic Sciences 72:277–81. doi:https://doi.org/10.1007/s00027-010-0132-0.
- Bhatia, M. P., E. B. Kujawinski, S. B. Das, C. F. Breier, P. B. Henderson, and M. A. Charette. 2013. Greenland meltwater as a significant and potentially bioavailable source of iron to the ocean. Nature Geoscience 6 (4):274–78. doi:https://doi.org/10.1038/ngeo1746.
- Burpee, B., J. E. Saros, R. M. Northington, and K. S. Simon. 2016. Microbial nutrient limitation in Arctic lakes in a permafrost landscape of southwest Greenland. Biogeosciences 13 (2):365–74. doi:https://doi.org/10.5194/bg-13-365-2016.
- Carrivick, J. L., and F. S. Tweed. 2013. Proglacial lakes: Character, behavior and geological importance. Quaternary Science Reviews 78:34–52. doi:https://doi.org/10.1016/j.quascirev.2013.07.028.
- Cole, J. J., M. L. Pace, S. R. Carpenter, and J. F. Kitchell. 2000. Persistence of net heterotrophy in lakes during nutrient addition and food web manipulations. Limnology and Oceanography 45 (8):1718–30. doi:https://doi.org/10.4319/lo.2000.45.8.1718.
- Cremer, H., and B. Wagner. 2004. Planktonic diatom communities in High Arctic lakes (Store Koldewey, Northeast Greenland). Canadian Journal of Botany 82 (12):1744–57. doi:https://doi.org/10.1139/b04-127.
- Daly, G. L., and F. Wania. 2005. Organic contaminants in mountains. Environmental Science & Technology 39 (2):385–98. doi:https://doi.org/10.1021/es048859u.
- Das, B., R. D. Vinebrooke, A. Sanchez-Azofeifa, B. Rivard, and A. P. Wolfe. 2005. Inferring sedimentary chlorophyll concentrations with reflectance spectroscopy: A novel approach to reconstructing historical changes in the trophic status of mountain lakes. Canadian Journal of Fisheries and Aquatic Sciences 62 (5):1067–78. doi:https://doi.org/10.1139/f05-016.
- Fegel, T. S., J. S. Baron, A. G. Fountain, G. F. Johnson, and E. K. Hall. 2016. The differing biogeochemical and microbial signatures of glaciers and rock glaciers. Journal of Geophysical Research: Biogeosciences 121:919–32.
- Findlay, S. E., R. L. Sinsabaugh, W. V. Sobczak, and M. Hoostal. 2003. Metabolic and structural response of hyporheic microbial communities to variations in supply of dissolved organic matter. Limnology and Oceanography 48 (4):1608–17. doi:https://doi.org/10.4319/lo.2003.48.4.1608.
- Fox, J., and S. Weisberg. 2011. An {R} companion to applied regression, 2nd ed. Thousand Oaks, CA: Sage.
- German, D. P., M. N. Weintraub, A. S. Grandy, C. L. Lauber, Z. L. Rinkes, and S. D. Allison. 2011. Optimization of hydrolytic and oxidative enzyme methods for ecosystem studies. Soil Biology and Biochemistry 43 (7):1387–97. doi:https://doi.org/10.1016/j.soilbio.2011.03.017.
- Gervais, F. 1997. Light-dependent growth, dark survival, and glucose uptake by cryptophytes isolated from a freshwater chemocline. Journal of Phycology 33 (1):18–25. doi:https://doi.org/10.1111/j.0022-3646.1997.00018.x.
- Granshaw, F. D., and A. G. Fountain. 2006. Glacier change (1958–1998) in the north Cascades national park complex, Washington, USA. Journal of Glaciology 52 (177):251–56. doi:https://doi.org/10.3189/172756506781828782.
- Hanna, E., P. Huybrechts, K. Steffen, J. Cappelen, R. Huff, C. Shuman, T. Irvine-Fynn, S. Wise, and M. Griffiths. 2008. Increased runoff from melt from the Greenland Ice Sheet: A response to global warming. Journal of Climate 21 (2):331–41. doi:https://doi.org/10.1175/2007JCLI1964.1.
- Hanna, E., S. H. Mernild, J. Cappelen, and K. Steffen. 2012. Recent warming in Greenland in a long-term instrumental (1881–2012) climatic context: I. Evaluation of surface air temperature records. Environmental Research Letters 7 (4):045404. doi:https://doi.org/10.1088/1748-9326/7/4/045404.
- Hastings, M. G., J. C. Jarvis, and E. J. Steig. 2009. Anthropogenic impacts on nitrogen isotopes of ice-core nitrate. Science 324 (5932):1288. doi:https://doi.org/10.1126/science.1170510.
- Hawkings, J., J. Wadham, M. Tranter, J. Telling, E. Bagshaw, A. Beaton, S. L. Simmons, D. Chandler, A. Tedstone, and P. Nienow. 2016. The Greenland Ice Sheet as a hot spot of phosphorus weathering and export in the Arctic. Global Biogeochemical Cycles 30:191–210. doi:https://doi.org/10.1002/2015GB005237.
- Hawkings, J. R., J. L. Wadham, M. Tranter, E. Lawson, A. Sole, T. Cowton, A. J. Tedstone, I. Bartholomew, P. Nienow, D. Chandler, et al. 2015. The effect of warming climate on nutrient and solute export from the Greenland Ice Sheet. Geochemical Perspectives Letters 1:94–104. doi:https://doi.org/10.7185/geochemlet.1510.
- Heiri, O., A. F. Lotter, and G. Lemcke. 2001. Loss on ignition as a method for estimating organic and carbonate content in sediments: Reproducibility and comparability of results. Journal of Paleolimnology 25 (1):101–10. doi:https://doi.org/10.1023/A:1008119611481.
- Hodson, A., P. Mumford, and D. Lister. 2004. Suspended sediment and phosphorus in proglacial rivers: Bioavailability and potential impacts upon the P status of ice: Marginal receiving waters. Hydrological Processes 18 (13):2409–22. doi:https://doi.org/10.1002/(ISSN)1099-1085.
- Holloway, J. M., and R. A. Dahlgren. 2002. Nitrogen in rock: Occurrences and biogeochemical implications. Global Biogeochemical Cycles 16 (4):1118. doi:https://doi.org/10.1029/2002GB001862.
- Interlandi, S. J., and S. S. Kilham. 2001. Limiting resources and the regulation of diversity in phytoplankton communities. Ecology 82 (5):1270–82. doi:https://doi.org/10.1890/0012-9658(2001)082[1270:LRATRO]2.0.CO;2.
- Jøsrgensen, E. G. 1969. The adaptation of plankton algae IV. Light adaptation in different algal species. Physiologia Plantarum 22 (6):1307–15. doi:https://doi.org/10.1111/j.1399-3054.1969.tb09121.x.
- Kammerlander, B., K. A. Koinig, E. Rott, R. Sommaruga, B. Tartarotti, F. Trattner, and B. Sonntag. 2016. Ciliate community structure and interactions within the planktonic food web in two alpine lakes of contrasting transparency. Freshwater Biology 61 (11):1950–65. doi:https://doi.org/10.1111/fwb.2016.61.issue-11.
- Knight, P. G., R. I. Waller, C. J. Patterson, A. P. Jones, and Z. P. Robinson. 2000. Glacier advance, ice-marginal lakes and routing of meltwater and sediment: Russell Glacier, Greenland. Journal of Glaciology 46 (154):423–26. doi:https://doi.org/10.3189/172756500781833160.
- Kopáček, J., J. Borovec, J. Hejzlar, K. U. Ulrich, S. A. Norton, and A. Amirbahman. 2005. Aluminum control of phosphorus sorption by lake sediments. Environmental Science & Technology 39 (22):8784–89. doi:https://doi.org/10.1021/es050916b.
- Kopáček, J., J. Hejzlar, J. Borovec, P. Porcal, and I. Kotorova. 2000. Phosphorus availability in an acidified watershed-lake ecosystem. Limnology and Oceanography 45 (1):212–25. doi:https://doi.org/10.4319/lo.2000.45.1.0212.
- Köster, D., and R. Pienitz. 2006. Seasonal diatom variability and paleolimnological inferences: A case study. Journal of Paleolimnology 35 (2):395–416. doi:https://doi.org/10.1007/s10933-005-1334-7.
- Krammer, K., and H. Lange-Bertalot. 1986–1991. Bacillariophyceae. In Sußwasserflora von Mitteleuropa, eds. H. Ettl, G. Gartner, J. Gerloff, H. Heynig, and D. Mollenhauer. Vols. 2 (1–4). Stuttgart/Jena: Gustav Fischer Verlag. (Freshwater flora of central Europe.).
- Lawson, E. C., J. L. Wadham, M. Tranter, M. Stibal, G. P. Lis, C. E. Butler, J. Laybourn-Parry, P. Nienow, D. Chandler, and P. Dewsbury. 2014. Greenland Ice Sheet exports labile organic carbon to the Arctic oceans. Biogeosciences 11 (14):4015–28. doi:https://doi.org/10.5194/bg-11-4015-2014.
- Lewis, S. M., and L. C. Smith. 2009. Hydrologic drainage of the Greenland Ice Sheet. Hydrological Processes 23 (14):2004–11. doi:https://doi.org/10.1002/hyp.v23:14.
- McClain, M. E., E. W. Boyer, C. L. Dent, S. E. Gergel, N. B. Grimm, P. M. Groffman, S. C. Hart, J. W. Harvey, C. A. Johnston, E. Mayorga, et al. 2003. Biogeochemical hot spots and hot moments at the interface of terrestrial and aquatic ecosystems. Ecosystems 6 (4):301–12. doi:https://doi.org/10.1007/s10021-003-0161-9.
- Nielsen, A. B., 2010. Present conditions in Greenland and the Kangerlussuaq area. Posiva Oy. Working Report no. 2010-07.
- Norton, S. A., R. H. Perry, J. E. Saros, G. L. Jacobson, I. J. Fernandez, J. Kopáček, T. A. Wilson, and M. D. SanClements. 2011. The controls on phosphorus availability in a Boreal lake ecosystem since deglaciation. Journal of Paleolimnology 46 (1):107–22. doi:https://doi.org/10.1007/s10933-011-9526-9.
- Oksanen, J., F. G. Blanchet, M. Friendly, R. Kindt, P. Legendre, D. McGlinn, P. R. Minchin, R. B. O’Hara, G. L. Simpson, P. Solymos, et al. 2017. vegan: Community ecology package. R Package Version 2.4-2.
- Pace, M. L., and J. J. Cole. 2000. Effects of whole-lake manipulations of nutrient loading and food web structure on planktonic respiration. Canadian Journal of Fisheries and Aquatic Sciences 57 (2):487–96. doi:https://doi.org/10.1139/f99-279.
- Perren, B. B., Y. Axford, and D. S. Kaufman. 2017. Alder, nitrogen, and lake ecology: Terrestrial-aquatic linkages in the postglacial history of lone spruce Pond, southwestern Alaska. PloS ONE 12 (1):0169106. doi:https://doi.org/10.1371/journal.pone.0169106.
- Peter, H., and R. Sommaruga. 2017. Alpine glacier-fed turbid lakes are discontinuous cold polymictic rather than dimictic. Inland Waters 7 (1):45–54. doi:https://doi.org/10.1080/20442041.2017.1294346.
- Porder, S., and S. Ramachandran. 2013. The phosphorus concentration of common rocks: A potential driver of ecosystem P status. Plant and Soil 367 (1–2):41–55. doi:https://doi.org/10.1007/s11104-012-1490-2.
- Psenner, R., R. Pucsko, and M. Sager. 1984. Die Fraktionierung organischer und anorganischer phosphorverbindungen von sedimenten: Versuch einer definition ökologisch wichtiger Frakionen. Archive Fur Hydrobiologie Supplement 70 (1):111–55.
- Saros, J. E., S. J. Interlandi, S. Doyle, T. J. Michel, and C. E. Williamson. 2005. Are the deep chlorophyll maxima in alpine lakes primarily induced by nutrient availability, not UV avoidance? Arctic, Antarctic, and Alpine Research 37 (4):557–63. doi:https://doi.org/10.1657/1523-0430(2005)037[0557:ATDCMI]2.0.CO;2.
- Saros, J. E., R. M. Northington, C. L. Osburn, B. T. Burpee, and N. J. Anderson. 2016. Thermal stratification in small arctic lakes of southwest Greenland affected by water transparency and epilimnetic temperatures. Limnology & Oceanography 61 (4):1530–42. doi:https://doi.org/10.1002/lno.10314.
- Saros, J. E., K. C. Rose, D. W. Clow, V. C. Stephens, A. B. Nurse, H. A. Arnett, J. R. Stone, C. E. Williamson, and A. P. Wolfe. 2010. Melting alpine glaciers enrich high-elevation lakes with reactive nitrogen. Environmental Science & Technology 44 (13):4891–96. doi:https://doi.org/10.1021/es100147j.
- Sheibley, R. W., M. Enache, P. W. Swarzenski, P. W. Moran, and J. R. Foreman. 2014. Nitrogen deposition effects on diatom communities in lakes from three national parks in Washington State. Water, Air, & Soil Pollution 225 (2):1857. doi:https://doi.org/10.1007/s11270-013-1857-x.
- Sinsabaugh, R. L., and C. M. Foreman. 2001. Activity profiles of bacterioplankton in a eutrophic river. Freshwater Biology (46):1239–49. doi:https://doi.org/10.1046/j.1365-2427.2001.00748.x.
- Sinsabaugh, R. L., B. H. Hill, and J. J. F. Shah. 2009. Ecoenzymatic stoichiometry of microbial organic nutrient acquisition in soil and sediment. Nature 462 (7274):795–98. doi:https://doi.org/10.1038/nature08632.
- Sinsabaugh, R. L., C. L. Lauber, M. N. Weintraub, B. Ahmed, S. D. Allison, C. Crenshaw, A. R. Contosta, D. Cusack, S. Frey, M. E. Gallo, et al. 2008. Stoichiometry of soil enzyme activity at global scale. Ecology Letters 11 (11):1252–64.
- Slemmons, K. E., A. Medford, B. L. Hall, J. R. Stone, S. McGowan, T. Lowell, M. Kelly, and J. E. Saros. 2016. Changes in glacial meltwater alter algal communities in lakes of Scoresby Sund, Renland, East Greenland throughout the holocene: Abrupt reorganizations began 1000 years before present. The Holocene 27 (7):929–40. doi:https://doi.org/10.1177/0959683616678468.
- Slemmons, K. E., and J. E. Saros. 2012. Implications of nitrogen-rich glacial meltwater for phytoplankton diversity and productivity in alpine lakes. Limnology and Oceanography 57 (6):1651–63. doi:https://doi.org/10.4319/lo.2012.57.6.1651.
- Slemmons, K. E., J. E. Saros, J. R. Stone, S. McGowan, C. T. Hess, and D. Cahl. 2015. Effects of glacier meltwater on the algal sedimentary record of an alpine lake in the central US Rocky Mountains throughout the late Holocene. Journal of Paleolimnology 53 (4):385–99. doi:https://doi.org/10.1007/s10933-015-9829-3.
- Smith, E. M., and Y. T. Prairie. 2004. Bacterial metabolism and growth efficiency in lakes: The importance of phosphorus availability. Limnology and Oceanography 49 (1):137–47. doi:https://doi.org/10.4319/lo.2004.49.1.0137.
- Sommaruga, R. 2015. When glaciers and ice sheets melt: Consequences for planktonic organisms. Journal of Plankton Research 37 (3):509–18. doi:https://doi.org/10.1093/plankt/fbv027.
- Spaulding, S. A., D. J. Lubinski, and M. Potapova, 2010: Diatoms of the United States. Accessed on March 11, 2017. http://westerndiatoms.colorado.edu.
- Stanish, L. F., E. A. Bagshaw, D. M. McKnight, A. G. Fountain, and M. Tranter. 2013. Environmental factors influencing diatom communities in Antarctic cryoconite holes. Environmental Research Letters 8 (4):045006. doi:https://doi.org/10.1088/1748-9326/8/4/045006.
- Tessier, A., P. Campbell, and M. Bisson. 1979. Sequential extraction procedure for the speciation of trace metals. Analytical Chemistry (51):844–51. doi:https://doi.org/10.1021/ac50043a017.
- Tietjen, T., and R. G. Wetzel. 2003. Extracellular enzyme-clay mineral complexes: Enzyme adsorption, alteration of enzyme activity, and protection from photodegradation. Aquatic Ecology 37 (4):331–39. doi:https://doi.org/10.1023/B:AECO.0000007044.52801.6b.
- van den Broeke, M., J. Bamber, J. Ettema, E. Rignot, E. Schrama, W. J. van de Berg, E. van Meijgaard, I. Velicogna, and B. Wouters. 2009. Partitioning recent Greenland mass loss. Science 326 (5955):984–86. doi:https://doi.org/10.1126/science.1178176.
- Vinebrooke, R. D., P. L. Thompson, W. Hobbs, B. H. Luckman, M. D. Graham, and A. P. Wolfe. 2010. Glacially mediated impacts of climate warming on alpine lakes of the Canadian Rocky Mountains. Verhandlungen Des Internationalen Verein Limnologie 30:1449–52.
- Vinšová, P., E. Pinseel, T. J. Kohler, B. van de Vijver, J. D. Žárský, J. Kavan, and K. Kopalová. 2015. Diatoms in cryoconite holes and adjacent proglacial freshwater sediments, Nordenskiöld glacier (Spitsbergen, High Arctic). Czech Polar Reports 5:112–33. doi:https://doi.org/10.5817/CPR2015-2-11.
- Wadham, J. L., M. Tranter, M. Skidmore, A. J. Hodson, J. Priscu, W. B. Lyons, M. Sharp, P. Wynn, and M. Jackson. 2010. Biogeochemical weathering under ice: Size matters. Global Biogeochemical Cycles 24 (3):GB3025. doi:https://doi.org/10.1029/2009GB003688.
- Wickham, H. 2009. ggplot2: Elegant Graphics for Data Analysis. New York: Springer-Verlag.
- Williams, J. J., A. Nurse, J. E. Saros, J. Riedel, and M. Beutel. 2016. Effects of glaciers on nutrient concentrations and phytoplankton in lakes within the Northern Cascades Mountains (USA). Biogeochemistry 131 (3):373–85. doi:https://doi.org/10.1007/s10533-016-0264-y.
- Williams, M. W., M. Knauf, R. Cory, N. Caine, and F. Liu. 2007. Nitrate content and potential microbial signature of rock glacier outflow, Colorado front range. Earth Surface Processes and Landforms 32 (7):1032–47. doi:https://doi.org/10.1002/(ISSN)1096-9837.
- Yallop, M. L., and A. M. Anesio. 2010. Benthic diatom flora in supraglacial habitats: A generic-level comparison. Annals of Glaciology 51 (56):15–22. doi:https://doi.org/10.3189/172756411795932029.