References
- Adams, W. P. 1966. Studies of ablation and run-off on an Arctic glacier. Doctoral diss., McGill University, Montreal, Canada.
- Bagshaw, E. A., M. Tranter, J. L. Wadham, A. G. Fountain, and M. Mowlem. 2011. High-resolution monitoring reveals dissolved oxygen dynamics in an Antarctic cryoconite hole. Hydrological Processes 25 (18):2868–77. doi:https://doi.org/10.1002/hyp.8049.
- Baker, B. B., and R. K. Moseley. 2007. Advancing treeline and retreating glaciers: Implications for conservation in Yunnan, P.R. China. Arctic, Antarctic, and Alpine Research 39 (2):200–09. doi:https://doi.org/10.1657/1523-0430(2007)39[200:atargi]2.0.co;2.
- Buda, J., E. Łokas, M. Pietryka, D. Richter, W. Magowski, N. S. Iakovenko, D. L. Porazinska, T. Budzik, M. Grabiec, J. Grzesiak, et al. 2020. Biotope and biocenosis of cryoconite hole ecosystems on ecology glacier in the maritime Antarctic. Science of the Total Environment 274. doi:https://doi.org/10.1016/j.scitotenv.2020.138112.
- Burnham, K. P., and D. R. Anderson. 2002. Model selection and multimodel inference: A practical information-theoretic approach. New York: Springer.
- Caccianiga, C., C. Andreis, G. Diolaiuti, C. D’Agata, C. Mihalcea, and C. Smiraglia. 2011. Alpine debris-covered glaciers as a habitat for plant life. The Holocene 21:1011–20. doi:https://doi.org/10.1177/0959683611400219.
- Cook, J. M., I. Hodson, J. Telling, A. Anesio, T. Irvine-Fynn, and C. Bellas. 2010. The mass–area relationship within cryoconite holes and its implications for primary production. Annals of Glaciology 51:106–10. doi:https://doi.org/10.3189/172756411795932038.
- Cook, J. M., M. Sweet, O. Cavalli, A. Taggart, and A. Edwards. 2018. Topographic shading influences cryoconite morphodynamics and carbon exchange. Arctic, Antarctic, and Alpine Research 50:1. doi:https://doi.org/10.1080/15230430.2017.1414463.
- Darcy, J. L., E. M. S. Gendron, P. Sommers, D. L. Porazinska, and S. K. Schmidt. 2018. Island biogeography of cryoconite hole bacteria in Antarctica’s Taylor Valley and around the world. Frontiers in Ecology and Evolution 6:180. doi:https://doi.org/10.3389/fevo.2018.00180.
- De Smet, W. H., and E. A. Van Rompu. 1994. Rotifera and Tardigrada from some cryoconite holes on a Spitsbergen Svalbard glacier. Belgian Journal of Zoology 124:27–37.
- Deharveng, L., C. A. D’Haese, and A. Bedos. 2008. Global diversity of springtails (Collembola; Hexapoda) in freshwater. Hydrobiologia 595:329–38. doi:https://doi.org/10.1007/s10750-007-9116-z.
- Edwards, A., A. Anesio, S. M. Rassner, B. Sattler, B. Hubbard, W. T. Perkins, M. Young, and G. W. Griffith. 2011. Possible interactions between bacterial diversity, microbial activity and supraglacial hydrology of cryoconite holes in Svalbard. The ISME Journal 5:150–60. doi:https://doi.org/10.1038/ismej.2010.100.
- Eisenbeis, G. 1982. Physiological absorption of liquid water by Collembola: Absorption by the ventral tube at different salinities. Journal of Insect Physiology 28:11–20. doi:https://doi.org/10.1016/0022-1910(82)90017-8.
- ESRI. 2013. Arc map version 10.2. Redlands, CA: ESRI Inc.
- Fair, H. 2017. Ecology of aquatic insects in monsoonal temperate glacier streams of Southeast Tibet: A departure from the conceptual model. Doctoral diss., The Ohio State University, Columbus.
- Fjellberg, A. 2010. Cryophilic Isotomidae (Collembola) of the Northwestern Rocky Mountains, U. S. A. Zootaxa 2513:27–79. doi:https://doi.org/10.5281/zenodo.196078.
- Fujii, Y., and K. Higuchi. 1977. Statistical analyses of the forms of the glaciers in the Khumbu Himal. Journal of Japanese Society of Snow and Ice 39:7–14. doi:https://doi.org/10.5331/seppyo.39.special_7.
- Fyffe, C. L., B. W. Brocka, M. P. Kirkbrideb, D. W. F. Mairc, N. S. Arnold, C. Smiragliae, G. Diolaiutie, and F. Diotrif. 2019. Do debris-covered glaciers demonstrate distinctive hydrological behavior compared to clean glaciers? Journal of Hydrology 570:584–97. doi:https://doi.org/10.1016/j.jhydrol.2018.12.069.
- Gerdel, R. W., and F. Drouet. 1960. The cryoconite of the Thule Area, Greenland. Transactions of the American Microscopical Society 79:256–72. doi:https://doi.org/10.2307/3223732.
- Guisan, A., and N. E. Zimmermann. 2000. Predictive habitat distribution models in ecology. Ecological Modelling 135:147–86. doi:https://doi.org/10.1016/S0304-3800(00)00354-9.
- He, Y. Q., Z. X. Li, X. M. Yang, W. X. Jia, X. Z. He, B. Song, N. N. Zhang, and Q. Liu. 2008. Changes of the Hailuogou Glacier, Mt. Gongga, China, against the background of global warming in the last several decades. Journal of China University of Geosciences 19:271–81. doi:https://doi.org/10.1016/S1002-0705(08)60045-X.
- Hodson, A., A. M. Anesio, M. Tranter, A. Fountain, M. Osborn, J. Priscu, J. Laybourn-Parry, and B. Sattler. 2008. Glacial ecosystems. Ecological Monographs 78:41–67. doi:https://doi.org/10.1890/07-0187.1.
- Johnson, J. B., and K. S. Omland. 2004. Model selection in ecology and evolution. Trends in Ecology and Evolution 19:101–08. doi:https://doi.org/10.1016/j.tree.2003.10.013.
- Konopová, B., D. Kolosov, and M. J. O’Donnell. 2019. Water and ion transport across the eversible vesicles in the collophore of the springtail Orchesella cincta. Journal of Experimental Biology 34:261–66. doi:https://doi.org/10.1242/jeb.200691.
- Liu, Q., and S. Liu. 2010. Seasonal evolution of the englacial and subglacial drainage systems of a temperate glacier revealed by hydrological analysis. Sciences in Cold and Arid Regions 21:51–58.
- Liu, Q., S. Y. Liu, Y. Zhang, X. Wang, Y. Zhang, W. Guo, and J. Xu. 2010. Recent shrinkage and hydrological response of Hailuogou glacier, a monsoon temperate glacier on the east slope of Mount Gongga, China. Journal of Glaciology 56:215–24. doi:https://doi.org/10.3189/002214310791968520.
- Makowska, N., K. Zawierucha, J. Mokracka, and R. Koczura. 2016. First report of microorganisms of Caucasus glaciers (Georgia). Biologia 71:620–25. doi:https://doi.org/10.1515/biolog-2016-0086.
- Margesin, R., and V. Miteva. 2011. Diversity and ecology of psychrophilic microorganisms. Research in Microbiology 162:346–61. doi:https://doi.org/10.1016/j.resmic.2010.12.004.
- Mazerolle, M. J. 2020. AICcmodavg: Model selection and multimodel inference based on (Q)AIC(c). R package version 2.3-1. https://cran.r-project.org/package=AICcmodavg.
- McIntyre, N. F. 1984. Cryoconite hole thermodynamics. Canadian Journal of Earth Sciences 21:152–56. doi:https://doi.org/10.1139/e84-016.
- Merritt, R. W., K. W. Cummins, and M. B. Berg. 2008. An introduction to the aquatic insects of North America. 4th ed. Dubuque, IA: Kendall Hunt Publishers.
- Milner, A. M. 2016. The Milner and Petts (1994) conceptual model of community structure within glacier-fed rivers: 20 years on. In River science: Research and management for the 21st century, ed. D. J. Gilvear, M. T. Greenwood, M. C. Thoms, and P. J. Wood, 156–70. West Sussex, UK: Wiley and Sons.
- Mueller, D. R., W. F. Vincent, W. H. Pollard, and C. H. Fristen. 2001. Glacial cryoconite ecosystems: A bipolar comparison of algal communities and habitats. Nova Hedwig Beih 123:173–97.
- Pittino, F., M. Maglio, I. Gandolfi, and R. S. Azzoni. 2018. Bacterial communities of cryoconite holes of a temperate alpine glacier show both seasonal trends and year-to-year variability. Annals of Glaciology 59:1–9. doi:https://doi.org/10.1017/aog.2018.16.
- Porazinska, D. L., A. G. Fountain, T. H. Nylen, M. Tranter, R. A. Virginia, and D. H. Wall. 2004. The biodiversity and biogeochemistry of cryoconite holes from McMurdo Dry Valley Glaciers, Antarctica. Arctic, Antarctic, and Alpine Research 36:84–91. doi:https://doi.org/10.1657/1523-0430(2004)036[0084:tbaboc]2.0.co;2.
- R Core Team. 2017. R version 3.3.3: A language and environment for statistical computing. R Vienna, Austria: Foundation for Statistical Computing, Vienna.
- Sommers, P., J. L. Darcy, E. M. S. Gendron, L. F. Stanish, E. A. Bagshaw, D. L. Porazinska, and S. K. Schmidt. 2018. Diversity patterns of microbial eukaryotes mirror those of bacteria in Antarctic cryoconite holes. FEMS Microbial Ecology 94:1–11. doi:https://doi.org/10.1093/femsec/fix167.
- Takeuchi, N., and S. Kohshima. 2004. A snow algal community on Tyndall Glacier in the Southern Patagonia Icefield, Chile. Arctic, Antarctic, and Alpine Research 36:92–99. doi:https://doi.org/10.1657/1523-0430(2004)036[0092:asacot]2.0.co;2.
- Takeuchi, N., S. Kohshima, Y. Yoshimura, K. Setko, and K. Fujita. 2000. Characteristics of cryoconite holes on a Himalayan glacier, Yala Glacier Central Nepal. Bulletin of Glaciological Research 17:51–59.
- Takeuchi, N., R. Sakaki, J. Uetake, N. Nagatsuka, R. Shimada, M. Niwano, and T. Aoki. 2018. Temporal variations of cryoconite holes and cryoconite coverage on the ablation ice surface of Qaanaaq Glacier in northwest Greenland. Annals of Glaciology 59:21–30. doi:https://doi.org/10.1017/aog.2018.19.
- Telling, J., A. M. Anesio, M. Tranter, A. G. Fountain, T. Nylen, J. Hawkings, V. B. Singh, P. Kaur, M. Musilova, and J. L. Wadham. 2014. Spring thaw ionic pulses boost nutrient availability and microbial growth in entombed Antarctic Dry Valley cryoconite holes. Frontiers in Microbiology 5:694. doi:https://doi.org/10.3389/fmicb.2014.00694.
- Tranter, M., A. G. Fountain, C. H. Fritsen, W. B. Lyons, J. C. Priscu, P. J. Statham, and K. A. Welch. 2004. Extreme hydrochemical conditions in natural microcosms entombed within Antarctic ice. Hydrological Processes 18 (2):379–87. doi:https://doi.org/10.1002/hyp.5217.
- Venables, W. N., and B. D. Ripley. 2002. Modern applied statistics with S. New York: Springer Science+Business Media.
- Vonnahme, T. R., M. Devetter, J. D. Žarský, M. Šabacká, and J. Elster. 2016. Controls on microalgal community structures in cryoconite holes upon High Arctic glaciers, Svalbard. Biogeosciences 12:11751–95. doi:https://doi.org/10.5194/bg-13-659-2016.
- Wharton, R. A., C. P. McKay, G. M. Simmons Jr., and B. C. Parker. 1985. Cryoconite holes on glaciers. BioScience 35:499–503. doi:https://doi.org/10.2307/1309818.
- Yao, T. D., L. Thompson, W. Yang, W. S. Yu, Y. Gao, X. J. Guo, X. X. Yang, K. Q. Duan, H. B. Zhao, B. Q. Xu, et al. 2012. Different glacier status with atmospheric circulations in Tibetan Plateau and surroundings. Nature Climate Change 2:663–67. doi:https://doi.org/10.1038/nclimate1580.
- Zawierucha, K., J. Buda, R. S. Azzoni, M. Niskiewicz, A. Franzetti, and R. Ambrosini. 2019. Water bears dominated cryoconite hole ecosystems: Densities, habitat preferences and physiological adaptations of Tardigrada on an alpine glacier. Aquatic Ecology 53:543–56. doi:https://doi.org/10.1007/s10452-019-09707-2.
- Zawierucha, K., J. Buda, D. Fontaneto, R. Abrosini, A. Franzetti, M. Wierzgon, and M. Bogdziewicz. 2019. Fine-scale spatial heterogeneity of invertebrates within cryoconite holes. Aquatic Ecology 53:179–90. doi:https://doi.org/10.1007/s10452-019-09681-9.
- Zawierucha, K., J. Buda, and A. Nawrot. 2019. Extreme weather events results in the removal of invertebrates from cryoconite holes on an Arctic valley glacier (Longyearbreen, Svalbard). Ecological Research 34:370–79. doi:https://doi.org/10.1111/1440-1703.1276.
- Zawierucha, K., J. Buda, M. Pietryka, M. Pietryka, D. Richter, E. Lokas, S. Lehmann-Konera, N. Makowska, and M. Bogdziewicz. 2018. Snapshot of micro-animals and associated biotic and abiotic environmental variables on the edge of the south-west Greenland ice sheet. Limnology 19:141–50. doi:https://doi.org/10.1007/s10201-017-0528-9.
- Zawierucha, K., M. Kolicka, N. Takeuchi, and L. Kaczmarek. 2015. What animals can live in cryoconite holes? A faunal review. Journal of Zoology 295:159–69. doi:https://doi.org/10.1111/jzo.1219.
- Zawierucha, K., M. Ostrowska, T. R. Vonnahme, M. Devetter, A. P. Nawrot, S. Lehman, and M. Kolicka. 2016. Diversity and distribution of Tardigrada in Arctic cryoconite holes. Journal of Limnology 75:545–59.
- Zawierucha, K., T. R. Vonnahme, M. Devetter, M. Kolicka, M. Ostrowska, S. Chmielewski, and J. Z. Kosicki. 2016. Area, depth and elevation of cryoconite holes in the Arctic do not influence Tardigrada densities. Polish Polar Research 37:325–34. doi:https://doi.org/10.1515/popore-2016-0009.
- Zhang, Y., K. Fujita, S. Y. Liu, Q. Liu, and N. Takayuki. 2011. Distribution of debris thickness and its effect on ice melt at Hailuogou glacier, southeastern Tibetan Plateau, using in situ surveys and ASTER imagery. Journal of Glaciology 571147–57. doi:https://doi.org/10.3189/002214311798843331.