Redescription of Antarctic eutardigrade Dastychius improvisus (Dastych, 1984) and some remarks on phylogenetic relationships within Isohypsibioidea

References

• Altschul SF, Gish W, Miller W, Myers EW, Lipman DJ. 1990. Basic local alignment search tool. Journal of Molecular Biology 215(3):403–410. DOI:10.1016/S0022-2836(05)80360-2.
   PubMed Web of Science ®Google Scholar
• Beasley CW. 1995. The phylum Tardigrada. Abilene: USA. p. 1–1014. Third Edition by G. Ramazzotti and W. Maucci, English Translation. Published by the translator Clark Beasley.
   Google Scholar
• Beasley CW, Kaczmarek Ł, Michalczyk Ł. 2008. Doryphoribius mexicanus, a new species of Tardigrada (Eutardigrada: Hypsibiidae) from Mexico (North America). Proceedings of the Biological Society of Washington 121(1):34–40. DOI:10.2988/07-30.1.
   Web of Science ®Google Scholar
• Bertolani R. 1982. 15. Tardigradi. Guide per il riconoscimento delle specie animali delle acque interne Italiane. Verona, Italy: Consiglio Nazionale Delle Ricerche. pp. 104.
   Google Scholar
• Bertolani R. 2003. Thulinius, new generic name substituting for Thulinia Bertolani, 1981 (Tardigrada, Eutardigrada). Zootaxa 314(1):1–4. DOI:10.11646/zootraxa.314.1.1.
   Google Scholar
• Bertolani R, Bartels PJ, Guidetti R, Cesari M, Nelson DR. 2014b. Aquatic tardigrades in the Great Smoky Mountains National Park, North Carolina and Tennessee, U.S.A., with the description of a new species of Thulinius (Tardigrada, Isohypsibiidae). Zootaxa 3764(5):524–536. DOI:10.11646/zootaxa.3764.5.2.
   Web of Science ®Google Scholar
• Bertolani R, Guidetti R, Marchioro T, Altiero T, Rebecchi L, Cesari M. 2014a. Phylogeny of Eutardigrada: New molecular data and their morphological support lead to the identification of new evolutionary lineages. Molecular Phylogenetics and Evolution 76:110–126. DOI: 10.1016/j.ympev.2014.03.006.
   PubMed Web of Science ®Google Scholar
• Binda MG. 1969. Nuovi dati su tardigradi di Sicilia con descrizione di due nuove specie. Bollettino delle Sedute della Accademia Gioenia di Scienze Naturali in Catania 9:623–633.
   Google Scholar
• Binda MG. 1980. Tardigradi di Lucania. Animalia 7:79–91.
   Google Scholar
• Binda MG, Pilato G. 1986. Ramazzottius, nuovo genere di Eutardigrado (Hypsibiidae). Animalia 13(1/3):159–166.
   Google Scholar
• Binda MG, Pilato G, Lisi O. 2005. Remarks on Macrobiotus furciger Murray, 1906 and description of three new species of the furciger group (Eutardigrada, Macrobiotidae). Zootaxa 1075(1):55–68. DOI:10.11646/zootaxa.1075.1.3.
   Google Scholar
• Casquet J, Thebaud C, Gillespie RG. 2012. Chelex without boiling, a rapid and easy technique to obtain stable amplifiable DNA from small amounts of ethanol-stored spiders. Molecular Ecology Resources 12(1):136–141. DOI:10.1111/j.1755-0998.2011.03073.x.
   PubMed Web of Science ®Google Scholar
• Cesari M, McInnes S, Bertolani R, Rebecchi L, Guidetti R. 2016a. Genetic diversity and biogeography of the south polar water bear Acutuncus antarcticus (Eutardigrada: Hypsibiidae) – Evidence that it is a truly pan-Antarctic species. Invertebrate Systematics 30(6):635–649. DOI:10.1071/IS15045.
   Web of Science ®Google Scholar
• Cesari M, Vecchi M, Palmer A, Bertolani R, Pilato G, Rebecchi L, Guidetti R. 2016b. What if the claws are reduced? Morphological and molecular phylogenetic relationships of the genus Haplomacrobiotus May, 1948 (Eutardigrada, Parachela). Zoological Journal of the Linnean Society 178(4):819–827. DOI:10.1111/zoj.12424.
   Web of Science ®Google Scholar
• Ciobanu DA, Zawierucha K, Moglan I, Kaczmarek Ł. 2014. Milnesium berladnicorum sp. n. (Eutardigrada, Apochela, Milnesiidae), a new species of water bear from Romania. ZooKeys 429:1–11. DOI: 10.3897/zookeys.429.7755.
   Web of Science ®Google Scholar
• Convey P, McInnes SJ. 2005. Exceptional tardigrade-dominated ecosystems in Ellsworth Land. Antarctica. Ecology 86(2):519–527. DOI:10.1890/04-0684.
   Web of Science ®Google Scholar
• Dabert M, Dastych H, Dabert J. 2015. Molecular data support the dispersal ability of the glacier tardigrade Hypsibius klebelsbergi Mihelčič, 1959 across the environmental barrier (Tardigrada). Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg 17:233–240.
   Google Scholar
• Dabert M, Dastych H, Hohberg K, Dabert J. 2014. Phylogenetic position of the enigmatic clawless eutardigrade genus Apodibius Dastych, 1983 (Tardigrada), based on 18S and 28S rRNA sequence data from its type species A. confusus. Molecular Phylogenetics and Evolution 70:70–75. DOI: 10.1016/j.ympev.2013.09.012.
   Web of Science ®Google Scholar
• Darriba D, Taboada GL, Doallo R, Posada D. 2012. jModelTest 2: More models, new heuristics and parallel computing. Nature Methods 9:772. DOI: 10.1038/nmeth.2109.
   PubMed Web of Science ®Google Scholar
• Dastych H. 1983. Apodibius confusus gen. n. sp. n. a new water-bear from Poland (Tardigrada). Bulletin of the Polish Academy of Sciences. Biological Sciences 31:41–46.
   Google Scholar
• Dastych H. 1984. The Tardigrada from Antarctica with description of several new species. Acta Zoologica Cracoviensia 27:377–436.
   Google Scholar
• Dastych H. 2018. Redescription and revalidation of the sub-Antarctic tardigrade Hypsibius murrayi (Richters, 1907) based on the rediscovered type material (Tardigrada, Panarthropoda). Entomologie heute 30:95–115.
   Google Scholar
• Degma P, Bertolani R, Guidetti. R 2020. Actual checklist of Tardigrada species. https://iris.unimore.it/retrieve/270444/Actual%20checklist%20of%20Tardigrada%2037th%20Edition%2008_07_20.pdf
   Google Scholar
• Degma P, Guidetti R. 2007. Notes to the current checklist of Tardigrada. Zootaxa 1579:41–53. DOI: 10.11646/zootaxa.1579.1.2.
   Web of Science ®Google Scholar
• Doyère LMF. 1840. Memoire sur les Tardigrades. I. Annales Des Sciences Naturelles Paris 14:269–362.
   Google Scholar
• Folmer O, Black M, Hoeh W, Lutz R, Vrijenhoek R. 1994. DNA primers for amplification of mitochondrial cytochrome c oxidase subunit I from diverse metazoan invertebrates. Molecular Marine Biology and Biotechnology 3:294–299.
   PubMedGoogle Scholar
• Gąsiorek P, Michalczyk Ł. 2020. Revised Cornechiniscus (Heterotardigrada) and new phylogenetic analyses negate echiniscid subfamilies and tribes. Royal Society Open Science 7:200581. DOI: 10.1098/rsos.200581.
   Web of Science ®Google Scholar
• Gąsiorek P, Morek W, Stec D, Blagden B, Michalczyk Ł. 2019a. Revisiting Calohypsibiidae and Microhypsibiidae: Fractonotus Pilato, 1998 and its phylogenetic position within Isohypsibiidae (Eutardigrada: Parachela). Zoosystema 41(6):71–89. DOI:10.5252/zoosystema2019v41a6.
   Web of Science ®Google Scholar
• Gąsiorek P, Stec D, Morek W, Michalczyk Ł. 2018. An integrative redescription of Hypsibius dujardini (Doyère, 1840), the nominal taxon for Hypsibioidea (Tardigrada: Eutardigrada). Zootaxa 4415:45–75. DOI: 10.11646/zootaxa.4415.1.2.
   PubMed Web of Science ®Google Scholar
• Gąsiorek P, Stec D, Morek W, Michalczyk Ł. 2019b. Deceptive conservatism of claws: Distinct phyletic lineages concealed within Isohypsibioidea (Eutardigrada) revealed by molecular and morphological evidence. Contributions to Zoology 88(1):78–132. DOI:10.1163/18759866-20191350.
   Web of Science ®Google Scholar
• Gąsiorek P, Stec D, Morek W, Zawierucha K, Kaczmarek Ł, Lachowska-Cierlik D, Michalczyk Ł. 2016. An integrative revision of Mesocrista Pilato, 1987 (Tardigrada: Eutardigrada: Hypsibiidae). Journal of Natural History 50:2803–2828. DOI: 10.1080/00222933.2016.1234654.
   Web of Science ®Google Scholar
• Gąsiorek P, Zawierucha K, Stec D, Michalczyk Ł. 2017. Integrative redescription of a common Arctic water bear Pilatobius recamieri (Richters, 1911). Polar Biology 40:2239–2252. DOI: 10.1007/s00300-017-2137-9.
   Web of Science ®Google Scholar
• Goujon M, Mcwilliam H, Li W, Valentin F, Squizzato S, Paern J, Lopez R. 2010. A new bioinformatics analysis tools framework at EMBL–EBI. Nucleic Acids Resources 38:695–699. DOI: 10.1093/nar/gkq313.
   PubMed Web of Science ®Google Scholar
• Guidetti R, Bertolani R. 2005. Tardigrade taxonomy: An updated check list of the taxa and a list of characters for their identification. Zootaxa 845:1–46. DOI: 10.11646/zootaxa.845.1.1.
   Web of Science ®Google Scholar
• Guidetti R, McInnes SJ, Cesari M, Rebecchi L, Rota-Stabelli O. 2017. Evolutionary scenarios for the origin of an Antarctic tardigrade species based on molecular clock analyses and biogeographic data. Contributions to Zoology 86(2):97–110. DOI:10.1163/18759866-08602001.
   Web of Science ®Google Scholar
• Guidetti R, Rebecchi L, Bertolani R, Jönsson KI, Kristensen RM, Cesari M. 2016. Morphological and molecular analyses on Richtersius (Eutardigrada) diversity reveal its new systematic position and lead to the establishment of a new genus and a new family within Macrobiotoidea. Zoological Journal of the Linnean Society 178:834–845. DOI: 10.1111/zoj.12428.
   Web of Science ®Google Scholar
• Guidetti R, Rebecchi L, Cesari M, McInnes SJ. 2014. Mopsechiniscus franciscae, a new species of a rare genus of Tardigrada from continental Antarctica. Polar Biology 37(9):1221–1233. DOI:10.1007/s00300-014-1514-x.
   Web of Science ®Google Scholar
• Guidetti R, Schill RO, Bertolani R, Dandekar T, Wolf M. 2009. New molecular data for tardigrade phylogeny, with the erection of Paramacrobiotus gen. n.. Journal of Zoological Systematics and Evolutionary Research 47:315–321. DOI: 10.1111/j.1439-0469.2009.00526.x.
   Web of Science ®Google Scholar
• Guil N, Giribet G. 2012. A comprehensive molecular phylogeny of tardigrades – Adding genes and taxa to a poorly resolved phylum-level phylogeny. Cladistics 28:21–49. DOI: 10.1111/j.1096-0031.2011.00364.x.
   PubMed Web of Science ®Google Scholar
• Guil N, Jørgensen A, Kristensen R. 2019. An upgraded comprehensive multilocus phylogeny of the Tardigrada tree of life. Zoologica Scripta 48:120–137. DOI: 10.1111/zsc.12321.
   Web of Science ®Google Scholar
• Hall TA. 1999. BioEdit: A user-friendly biological sequence alignment editor and analysis program for Windows 95/98/NT. Nucleic Acids Symposium Series 41:95–98. DOI: 10.14601/Phytopathol_Mediterr-14998u1.29.
   Google Scholar
• Iharos G. 1966. Neue Tardigraden-arten aus Ungarn. Acta Zoologica Academiae Scientiarum 12:111–122.
   Web of Science ®Google Scholar
• Jørgensen A, Faurby S, Hansen JG, Mobjerg N, Kristensen RM. 2010. Molecular phylogeny of Arthrotardigrada (Tardigrada). Molecular Phylogenetics and Evolution 54(3):1006–1015. DOI:10.1016/j.ympev.2009.10.006.
   Web of Science ®Google Scholar
• Jørgensen A, Kristensen RM. 2004. Molecular phylogeny of Tardigrada – Investigation of the monophyly of Heterotardigrada. Molecular Phylogenetics and Evolution 32:666–670. DOI: 10.1016/j.ympev.2004.04.017.
   Web of Science ®Google Scholar
• Kaczmarek Ł, Bartylak T, Roszkowska M. 2020b. Two new genera of long clawed Isohypsibioidea Guil, Jørgensen & Kristensen, 2019. Zootaxa 4729(2):293–299. DOI:10.11646/zootaxa.4729.2.10.
   Web of Science ®Google Scholar
• Kaczmarek Ł, Cytan J, Zawierucha K, Diduszko D, Michalczyk Ł. 2014. Tardigrades from Peru (South America), with descriptions of three new species of Parachela. Zootaxa 3790(2):357–379. DOI:10.11646/zootaxa.3790.2.5.
   PubMed Web of Science ®Google Scholar
• Kaczmarek Ł, Grobys D, Kulpa A, Bartylak T, Kmita H, Kepel M, Kepel A, Roszkowska M. 2019. Two new species of the genus Milnesium Doyère, 1840 (Tardigrada, Apochela, Milnesiidae) from Madagascar. ZooKeys 884:1–22. DOI: 10.3897/zookeys.884.29469.
   PubMed Web of Science ®Google Scholar
• Kaczmarek Ł, Michalczyk Ł. 2017. The Macrobiotus hufelandi group (Tardigrada) revisited. Zootaxa 4363(1):101–123. DOI:10.11646/zootaxa.4363.1.4.
   PubMed Web of Science ®Google Scholar
• Kaczmarek Ł, Mioduchowska M, Kačarević U, Kubska K, Parnikoza I, Gołdyn B, Roszkowska M. 2020a. New records of Antarctic Tardigrada with comments on interpopulation variability of the Paramacrobiotus fairbanksi Schill, Förster, Dandekar and Wolf, 2010. Diversity 12:108. DOI: 10.3390/d12030108.
   Web of Science ®Google Scholar
• Kaczmarek Ł, Parnikoza I, Gawlak M, Esefeld J, Peter H-U, Kozeretska I, Roszkowska M. 2018. Tardigrades from Larus dominicanus Lichtenstein, 1823 nests on the Argentine Islands (maritime Antarctic). Polar Biology 41(2):283–301. DOI:10.1007/s00300-017-2190-4.
   Web of Science ®Google Scholar
• Kiehl E, Dastych H, D’Haese J, Greven H. 2007. The 18S rDNA sequences support polyphyly of Hypsibiidae (Eutardigrada). Journal of Limnology 66:21–25. DOI: 10.4081/jlimnol.2007.s1.21.
   Web of Science ®Google Scholar
• Kihm JH, Kim S, McInnes SJ, Zawierucha K, Rho HS, Kang P, Park TYS. 2020. Integrative description of a new Dactylobiotus (Eutardigrada: Parachela) from Antarctica that reveals an intraspecific variation in tardigrade egg morphology. Scientific Reports 10:9122. DOI: 10.1038/s41598-020-65573-1.
   Web of Science ®Google Scholar
• Kimble JM. 2004. Cryosols. Permafrost-affected soils. Berlin: Springer.
   Google Scholar
• Kumar S, Stecher G, Li M, Knyaz C, Tamura K. 2018. MEGA X: Molecular Evolutionary Genetics Analysis across Computing Platforms. Molecular Biology and Evolution 35(6):1547–1549. DOI:10.1093/molbev/msy096.
   PubMed Web of Science ®Google Scholar
• Librado P, Rozas J. 2009. DnaSP v5: A software for comprehensive analysis of DNA polymorphism data. Bioinformatics 25:1451–1452. DOI: 10.1093/bioinformatics/btp187.
   PubMed Web of Science ®Google Scholar
• Lindsay DC. 1971. Vegetation of the South Shetland Islands. British Antarctic Survey Bulletin 25:59–83.
   Google Scholar
• McInnes SJ. 2010. Echiniscus corrugicaudatus (Heterotardigrada; Echiniscidae) a new species from Ellsworth Land, Antarctica. Polar Biology 33:59–70. DOI: 10.1007/s00300-009-0684-4.
   Web of Science ®Google Scholar
• Michalczyk Ł, Kaczmarek Ł. 2013. The Tardigrada Register: A comprehensive online data repository for tardigrade taxonomy. Journal of Limnology 72:175–181. DOI: 10.4081/jlimnol.2013.s1.e22.
   Web of Science ®Google Scholar
• Mironov SV, Dabert J, Dabert M. 2012. A new feather mite species of the genus Proctophyllodes Robin, 1877 (Astigmata: Proctophyllodidae) from the long–tailed tit Aegithalos caudatus (Passeriformes: Aegithalidae): Morphological description with DNA barcode data. Zootaxa 3253:54–61. DOI: 10.11646/zootaxa.3253.1.2.
   Google Scholar
• Møbjerg N, Jørgensen A, Eibye-Jacobsen J, Halberg KA, Persson D, Kristensen RM. 2007. New records on cyclomorphosis in the marine eutardigrade Halobiotus crispae (Eutardigrada: Hypsibiidae). Journal of Limnology 66:132–140. DOI: 10.4081/jlimnol.2007.s1.132.
   Web of Science ®Google Scholar
• Morek W, Ciosek J, Michalczyk Ł. 2020. Description of Milnesium pentapapillatum sp. nov., with an amendment of the diagnosis of the order Apochela and abolition of the class Apotardigrada (Tardigrada). Zoologischer Anzeiger 288:107–117. DOI: 10.1016/j.jcz.2020.07.002.
   Web of Science ®Google Scholar
• Morek W, Gąsiorek P, Stec D, Blagden B, Michalczyk Ł. 2016a. Experimental taxonomy exposes ontogenetic variability and elucidates the taxonomic value of claw configuration in Milnesium Doyère, 1840 (Tardigrada: Eutardigrada: Apochela). Contributions to Zoology 85:173–200. DOI: 10.1163/18759866-08502003.
   Web of Science ®Google Scholar
• Morek W, Stec D, Gąsiorek P, Schill RO, Kaczmarek Ł, Michalczyk Ł. 2016b. An experimental test of tardigrade preparation methods for light microscopy. Zoological Journal of the Linnean Society 178:785–793. DOI: 10.1111/zoj.12457.
   Web of Science ®Google Scholar
• Morgan CI. 1976. Studies on the British tardigrade fauna. Some zoogeographical and ecological notes. Journal of Natural History 10:607–632. DOI: 10.1080/00222937600770491.
   Web of Science ®Google Scholar
• Murray J. 1907. Scottish Tardigrada collected by the Lake Survey. Transactions of the Royal Society of Edinburgh 45:641–668. DOI: 10.1017/S0080456800011777.
   Google Scholar
• Nelson DR, Guidetti R, Rebecchi L. 2015. Chapter 17: Phylum Tardigrada. In: Thorp JH, Covich AP, editors. Ecology and general biology: vol. 1: freshwater invertebrates. 4th ed. London: Elsevier. pp. 347–380.
   Google Scholar
• Perry E, Miller WR, Kaczmarek Ł. 2019. Recommended abbreviations for the names of genera of the phylum Tardigrada. Zootaxa 4608:145–154. DOI: 10.11646/zootaxa.4608.1.8.
   Web of Science ®Google Scholar
• Pilato G. 1974. Tardigradi delle acque dolci siciliane - Terza Nota. Animalia 1:235–244.
   Google Scholar
• Pilato G. 1981. Analisi di nuovi caratteri nello studio degli Eutardigradi. Animalia 8:51–57.
   Google Scholar
• Pilato G. 2013. The taxonomic value of the structures for the insertion of the stylet muscles in the Eutardigrada, and description of a new genus. Zootaxa 3721(4):365–378. DOI:10.11646/zootaxa.3721.4.4.
   Web of Science ®Google Scholar
• Pilato G, Bertolani R, Binda MG. 1982. Studio degli Isohypsibius del gruppo elegans (Eutardigrada, Hypsibiidae) con descrizione di due nuove specie. Animalia 9:185–198.
   Google Scholar
• Pilato G, Binda MG. 2010. Definition of families, subfamilies, genera and subgenera of the Eutardigrada and keys to their identification. Zootaxa 2404:1–52. DOI: 10.5281/zenodo.194138.
   Google Scholar
• Pilato G, McInnes SJ, Lisi O. 2012. Hebesuncus mollispinus (Eutardigrada, Hypsibiidae), a new species from maritime Antarctica. Zootaxa 3446:60–68. DOI: 10.11646/zootaxa.3446.1.4.
   Google Scholar
• Pilato G, Sabella G, D’Urso V, Lisi O. 2017. Two new species of Eutardigrada from Victoria Land, Antarctica. Zootaxa 4317(3):541–558. DOI:10.11646/zootaxa.4317.3.6.
   Web of Science ®Google Scholar
• Posada D, Buckley TR. 2004. Model selection and model averaging in phylogenetics: Advantages of the AIC and Bayesian approaches over likelihood ratio tests. Systematic Biology 53:793–808. DOI: 10.1080/10635150490522304.
   PubMed Web of Science ®Google Scholar
• Pugh PJA, Convey P. 2008. Surviving out in the cold: Antarctic endemic invertebrates and their refugia. Journal of Biogeography 35:2176–2186. DOI: 10.1111/j.1365-2699.2008.01953.x.
   Web of Science ®Google Scholar
• Ramazzotti G, Maucci W. 1983. Il Phylum Tardigrada. III edizione riveduta e aggiornata. Memorie dell’Istituto Italiano di Idrobiologia 41:1–1012.
   Google Scholar
• Rambaut A, Suchard MA, Xie D, Drummond AJ. 2014. Tracer v1.6. Available from http://beast.bio.ed.ac.uk/Tracer
   Google Scholar
• Rice P, Longden I, Bleasby A. 2000. EMBOSS: The European molecular biology open software suite. Trends in Genetics 16:276–277. DOI: 10.1016/s0168-9525(00)02024-2.
   PubMed Web of Science ®Google Scholar
• Richters F. 1900. Beiträge zur Kenntnis der fauna der Umgebung von Frankfurt a. Frankfurt a. M: M. Ber. Senkenberg. Naturf. Ges. 21–44 pp.
   Google Scholar
• Richters F. 1926. Tardigrada. In: Kükenthal W, Krumbach T, editors. Handbuch der Zoologie. Berlin: Walter de Gruyter & Co. Vol. 3. pp. 58–61.
   Google Scholar
• Riffenburh B. 2007. Encyclopedia of the Antarctic. New York: Taylor Francis Group.
   Google Scholar
• Ronquist F, Huelsenbeck JP. 2003. MrBayes 3: Bayesian phylogenetic inference under mixed models. Bioinformatics 19:1572–1574. DOI: 10.1093/bioinformatics/btg180.
   PubMed Web of Science ®Google Scholar
• Sands CJ, Mcinnes SJ, Marley NJ, Goodall–Copestake WP, Convey P, Linse K. 2008. Phylum Tardigrada: An “individual” approach. Cladistics 24:861–871. DOI: 10.1111/j.1096-0031.2008.00219.x.
   PubMed Web of Science ®Google Scholar
• Schill RO, Steinbrück G. 2007. Identification and differentiation of Heterotardigrada and Eutardigrada species by riboprinting. Journal of Zoological Systematics and Evolutionary Research 45:184–190. DOI: 10.1111/j.1439-0469.2007.00409.x.
   Web of Science ®Google Scholar
• Schultze CAS. 1834. Macrobiotus Hufelandii animal e crustaceorum classe novum, reviviscendi post diuturnam asphixiam et aridiatem potens, etc. 8, 1 tab. C. Berlin: Curths. p. 6. I Table.
   Google Scholar
• Schuster RO, Nelson DR, Grigarick AA, Christenberry D. 1980. Systematic criteria of Eutardigrada. Transactions of the American Microscopical Society 99:284–303. DOI: 10.2307/3226004.
   Google Scholar
• Smith RIL, Corner RWM. 1973. Vegetation of the Arthur Harbour-Argentine Islands region of the Antarctic Peninsula. British Antarctic Survey Bulletin 33(34):89–122.
   Google Scholar
• Stec D, Morek W, Gąsiorek P, Michalczyk Ł. 2018. Unmasking hidden species diversity within the Ramazzottius oberhaeuseri complex, with an integrative redescription of the nominal species for the family Ramazzottiidae (Tardigrada: Eutardigrada: Parachela). Systematics and Biodiversity 16:357–376. DOI: 10.1080/14772000.2018.1424267.
   Web of Science ®Google Scholar
• Stec D, Smolak R, Kaczmarek Ł, Michalczyk Ł. 2015. An integrative description of Macrobiotus paulinae sp. nov. (Tardigrada: Eutardigrada: Macrobiotidae: hufelandi group) from Kenya. Zootaxa 4052(5):501–526. DOI:10.11646/zootaxa.4052.5.1.
   PubMed Web of Science ®Google Scholar
• Stec D, Zawierucha K, Michalczyk Ł. 2017. An integrative description of Ramazzottius subanomalus (Biserov, 1985) (Tardigrada) from Poland. Zootaxa 4300:403–420. DOI: 10.11646/zootaxa.4300.3.4.
   Web of Science ®Google Scholar
• Thulin G. 1928. Über die Phylogenie und das System der Tardigraden. Hereditas Lund 11:207–266.
   Google Scholar
• Tsujimoto M, Kagoshima H, Kanda H, Watanabe K, Imura S. 2020. Reproductive performance of the Antarctic tardigrades, Acutuncus antarcticus (Eutardigrada: Hypsibiidae), revived after being frozen for over 30 years and of their offspring. Zoological Journal of the Linnean Society 188(3):839–847. DOI:10.1093/zoolinnean/zlz137.
   Web of Science ®Google Scholar
• Tsujimoto M, McInnes SJ, Convey P, Imura S. 2014. Preliminary description of tardigrade species diversity and distribution pattern around coastal Syowa Station and inland Sør Rondane Mountains, Dronning Maud Land, East Antarctica. Polar Biology 37(9):1361–1367. DOI:10.1007/s00300-014-1516-8.
   Web of Science ®Google Scholar
• Tumanov DV. 2003. Four new Isohypsibius species from Russian fresh waters (Tardigrada, Hypsibiidae). Bulletin De l’Institut Royal Des Sciences Naturelles De Belgique Biologie 73:183–189.
   Google Scholar
• Vecchi M, Cesari M, Bertolani R, Jönsson KI, Rebecchi L, Guidetti R. 2016. Integrative systematic studies on tardigrades from Antarctica identify new genera and new species within Macrobiotoidea and Echiniscoidea. Invertebrate Systematics 30(4):303–322. DOI:10.1071/IS15033.
   Web of Science ®Google Scholar
• Velasco-Castrillón A, Gibson JAE, Stevens MI. 2014. A review of current Antarctic limno-terrestrial microfauna. Polar Biology 37(10):1517–1531. DOI:10.1007/s00300-014-1544-4.
   Web of Science ®Google Scholar
• Vicente F, Bertolani R. 2013. Considerations on the taxonomy of the Phylum Tardigrada. Zootaxa 3626(2):245–248. DOI:10.11646/zootaxa.3626.2.2.
   PubMed Web of Science ®Google Scholar
• Zawierucha Z, Stec D, Lachowska-Cierlik D, Takeuchi N, Li Z, Michalczyk Ł. 2018. High mitochondrial diversity in a new water bear species (Tardigrada: Eutardigrada) from mountain glaciers in central Asia, with the erection of a new genus Cryoconicus. Annales Zoologici 68(1):179–201. DOI:10.3161/00034541ANZ2018.68.1.007.
   Web of Science ®Google Scholar