REFERENCES
- Gradstein SR. 1991. Diversity and distribution of Asian Lejeuneaceae subfamily Ptychanthoideae. Tropical Bryology 4: 1–16.
- Hattori S. 1944. Contributio ad Floram Hepaticarum Austro-kiushiuensem. Bulletin of the Tokyo Science Museum 11: 1–203.
- Hattori S. 1952. Hepaticae of Shikoku and Kyushu, southern Japan. 2. Journal of the Hattori Botanical Laboratory 8: 21–46.
- Mizutani M. 1961. A review of Japanese Lejeuneaceae. Journal of the Hattori Botanical Laboratory 24: 115–302.