References

• Ballarín I. 1985. Árboles y arbustos del Moncayo. Zaragoza: Diputación General de Aragón.
   Google Scholar
• Barr TC. 1968. Cave ecology and the evolution of troglobites. In: Dobzhansky T, Hecht MK, Steere WC, editors. Evolutionary Biology. New York: Appleton-Century-Crofts. p. 35–102.
   Google Scholar
• Carceller F, Arribas O. 1993. Directrices de gestión de las masas forestales para el mantenimiento de la biodiversidad faunística, el caso del Moncayo, Zaragoza. Congreso Florestal Español. 4:27–33.
   Google Scholar
• Cate PC [Web Service]. 2004. Fauna Europaea: Elateridae, Dima. In: Alonso-Zarazaga MA, de Jong YSDM, editors. Fauna Europaea: Coleoptera. Fauna Europaea version 2.6; [cited 2014 June 5]. Available from http://www.faunaeur.org
   Google Scholar
• Cobos A. 1961. Sobre las Dima Charpentier (Coleoptera, Elateridae) ibéricas del supuesto subgénero Celox Schauf., y rehabilitación de D. assoi Pérez Arcas. Miscelánea Zoológica. 1(4):109–113.
   Google Scholar
• Darlington Jr PJ. 1943. Carabidae of mountains and islands: data on the evolution of isolated faunas, and on atrophy of wings. Ecological Monographs. 13(1):37–61.
   Google Scholar
• Desutter-Grandcolas L. 1997. Are troglobitic taxa troglobiomorphic? A test using phylogenetic inference. International Journal of Speleology. 26:1–19.
   Google Scholar
• Dethier M, Hubart JM. 2005. La “troglobitude”: adaptations à la vie souterraine. Notes fauniques de Glembloux. 57:29–48.
   Google Scholar
• Fuente JM. 1929. Catálogo sistemático-geográfico de los Coleópteros observados en la Península Ibérica, Pirineos propiamente dichos y Baleares. Boletín de la Sociedad Entomológica de España. 12:108–121.
   Google Scholar
• Gers C. 1992. Ecologie et biologie des populations d’arthropodes, terrestres du milieu souterrain superficiel: Fonctionnement et Ecologie évolutive [PhD thesis]. Toulouse: Université Paul Sabatier.
   Google Scholar
• Gers C. 1998. Diversity of energy fluxes and interactions between arthropod communities: from soil to cave. Acta Ecologica. 19:205–213.
   Google Scholar
• Giachino PM, Vailati D. 2010. The subterranean environment. Hypogean life, concepts and collecting techniques. Verona: WBA Handbooks.
   Google Scholar
• Gould SJ, Vrba ES. 1982. Exaptation – a missing term in the science of form. Paleobiology. 8(1):4–15.
   Web of Science ®Google Scholar
• Heads SW. 2010. The first fossil spider cricket (Orthoptera: Gryllidae: Phalangopsinae): 20 million years of troglobiomorphosis or exaptation in the dark? Zoological Journal of the Linnean Society. 158:56–65.
   Web of Science ®Google Scholar
• Hernando C, Ribera I, Vogler AP. 1999. Alpine and cave or endogean habitats as postglacial refugia: Examples from paleartic ground beetles, with comments on their possible origins (Coleoptera: Carabidae). The Coleopterist Bulletin. 53:31–39.
   Web of Science ®Google Scholar
• Jeannel R. 1943. Les fossiles vivants des cavernes. Paris: Gallimard.
   Google Scholar
• Juberthie C, Delay D, Bouillon M. 1980. Extension du milieu souterrain en zone non calcaire: description d’un nouveau milieu et de son peuplement par les Coléoptères troglobies. Mémoires de Biospéologie. 7:19–52.
   Google Scholar
• Kühnelt W. 1957. Biología del suelo. Madrid: Consejo Superior de Investigaciones Científicas.
   Google Scholar
• Leseigneur L. 1972. Coléoptères Elatérides de la faune de France continentale et de Corse. Supplément au Bulletin mensuel de la Société Linnéenne de Lyon. 41:1–381.
   Google Scholar
• Löbl I, Smetana A. 2014. Errata for volume 4 [Elateridae]. Catalogue of Palaearctic Coleoptera; [cited 2014June 5]. Available from http://www.ville-ge.ch/mhng/pdf/palcoleo_errata.pdf
   Google Scholar
• López H, Oromí P. 2010. A pitfall trap for sampling the mesovoid shallow substratum (MSS) fauna. Speleobiology Notes. 2:7–11.
   Google Scholar
• McClure EH, Lim BL, Winn SE. 1967. Fauna of Dark Cave, Batu Caves, Kuala Lumpur, Malaysia. Pacific Insects. 9(3):399–428.
   Web of Science ®Google Scholar
• Molero-Baltanás R, Gaju Ricart M, Bach C. 2013: Los Tisanuros y Lepismas (Microcoryphia y Zygentoma). In: Ruano F, Tierno de Figueroa M, Tinaut A, editors. Los Insectos de Sierra Nevada: 200 años de historia. Santa Fe, Granada: GRX Servicios Gráficos; p. 88–98.
   Google Scholar
• Navarro G. 1989. Datos sobre la vegetación del moncayo. Turiaso. 9:423–431.
   Google Scholar
• Nitzu, E, Nae A, Băncilă R, Popa I, Giurginca A, Plăiaşu R. 2014. Scree habitats: ecological function, species conservation and spatial-temporal variation in the arthropod community. Systematics and Biodiversity. 12(1):65–75.
   Google Scholar
• Nitzu E, Nae A, Giurginca, A, Popa I. 2010. Invertebrate communities from the Mesovoid Shallow Substratum of the Carpatho-Euxinic Area: Eco-Faunistic and Zoogeographic Analysis. Travaux de l’Institut de Spéologie Émile Racovitza. 49:41–79.
   Google Scholar
• Oromí P, Medina A, Tejedor M. 1986. On the existence of a superficial underground compartment in the Canary Islands. Acta IX Congreso Internacional de Espeleología, Barcelona. 2:147–151.
   Google Scholar
• Ortuño VM, Gilgado JD, Jiménez-Valverde A, Sendra A, Pérez-Suárez G, Herrero-Borgoñón JJ. 2013. The “alluvial mesovoid shallow substratum”, a new subterranean habitat. PLoS ONE 8(10):e76311. doi:10.1371/journal.pone.0076311
   Web of Science ®Google Scholar
• Ortuño VM, Toribio M. 1994. Nuevos datos sobre la distribución y ecología de Nebria vuillefroyi Chaudoir, 1866 (Coleoptera: Nebriidae). Boletín de la Asociación española de Entomología. 18(1–2):204–205.
   Google Scholar
• Peck SB. 1982. A contribution to the knowledge of invertebrate cave faunas of Venezuela. Invertebrate faunas of tropical American caves, part 4. International Journal of Speleology. 12:75–81.
   Google Scholar
• Peck SB, Ruiz-Baliú AE, Garcés González GF. 1998. The cave-inhabiting beetles of Cuba (Insecta: Coleoptera): Diversity, Distribution and Ecology. Journal of Cave and Karst Studies. 60(3):156–166.
   Google Scholar
• Peláez D. 1936–1937. [capture note]. Boletín de la Sociedad Española de Historia Natural. 36:325–326.
   Google Scholar
• Pellicer F. 1980. El periglaciarismo del Moncayo. Geographicalia. 7:3–26.
   Google Scholar
• Pérez Arcas L. 1872. Especies nuevas o críticas de la Fauna Española. Segunda Parte. Anales de la Sociedad Española de Historia Natural. 1:89–137.
   Google Scholar
• Pipan T, López H, Oromí P, Polak S, Culver DC. 2011. Temperature variation and the presence of troglobionts in terrestrial shallow subterranean habitats. Journal of Natural History. 45:253–273.
   Web of Science ®Google Scholar
• Rendoš M, Mock A, Jászay T. 2012. Spatial and temporal dynamics of invertebrates dwelling karstic mesovoid shallow substratum of Sivec National Nature Reserve (Slovakia), with emphasis on Coleoptera. Biologia. 67:1143–1151.
   Web of Science ®Google Scholar
• Růžička V. 1990. The spiders of stony debris. Acta Zoologica Fennica. 190:333–337.
   Google Scholar
• Růžička V. 1999. The first steps in subterranean evolution of spiders (Araneae) in Central Europe. Journal of Natural History. 33:255–265.
   Web of Science ®Google Scholar
• Růžička V, Hajer J, Zacharda M. 1995. Arachnid population patterns in underground cavities of a stony debris field (Araneae, Opiliones, Pseudoscorpionidea, Acari: Prostigmata, Rhagidiidae). Pedobiologia. 39:42–51.
   Web of Science ®Google Scholar
• Růžička V, Zacharda M, Nemcova L, Šmilauer P, Nekola JC. 2012. Periglacial microclimate in low-altitude scree slopes supports relict biodiversity. Journal of Natural History. 46:2145–2157.
   Web of Science ®Google Scholar
• Schimmel R. 1996. Das Monophylum Diminae Candèze, 1863 (Insecta: Coleoptera: Elateridae). Bad Dürkheim: Pollichia, Pfalzmuseum für Naturkunde, 367 p.
   Google Scholar
• Schimmel R, Platia G. 2008. The European species of genus Dima Charpentier 1825: (Coleoptera, Elateridae). Bollettino del Museo Regionale di Scienze Naturali di Torino. 25(2):559–594.
   Google Scholar
• Sket B. 2008. Can we agree on an ecological classification of subterranean animals? Journal of Natural History. 42(21):1549–1563.
   Web of Science ®Google Scholar
• Vandel A. 1964. Biospéologie: la biologie des animaux cavernicoles. Paris: Gauthier-Villars.
   Google Scholar
• Wunderlich J. 1993. The Macaronesian cave-dwelling spider fauna (Arachnida: Araneae). Memoirs of the Queensland Museum. 33(2):681–686.
   Google Scholar
• Zapata JL, Sánchez-Ruiz A. 2012. Catálogo actualizado de los Elatéridos de la Península Ibérica e Islas Baleares (Coleoptera: Elateridae). Arquivos entomolóxicos. 6:115–271.
   Google Scholar