A new species of Numidotherium (Mammalia: Proboscidea) from the Eocene of Libya and the early phylogeny of the Proboscidea

LITERATURE CITED

• Andrews , C. W. 1901 . Preliminary note on some recently discovered extinct vertebrates from Egypt . Geological Magazine , 8 : 400 – 436 .
   Google Scholar
• Andrews , C. W. 1904 . Note on the Barypoda, a new order of ungulate mammals . Geological Magazine , 1 : 481 – 482 .
   Google Scholar
• Andrews , C. W. 1906 . A Descriptive Catalogue of the Tertiary Vertebrata of the Fayum, Egypt 324 London : British Museum (Natural History) .
   Google Scholar
• Arambourg , C. and Magnier , P. 1961 . Gisements de vertébrés dans le bassin tertiaire de Syrte (Libye). Comptes Rendus de l'Académie des Sciences . Série II , 252 : 1181 – 1183 .
   Google Scholar
• Bertrand , P. 1988 . “ Evolution de la structure de l'émail chez les Proboscidea primitif: aspect phylogénétique et fonctionell; ” . In Teeth Revisited. Proceedings of the VIIth International Symposium on Dental Morphology Edited by: Russell , D. E. , Santora , J. P. and Sigogneau-Russell , D. 109 – 124 . Paris : Mémoires de Muséum National d'Histoire Naturelle .
   Google Scholar
• Bown , T. M. and Kraus , M. J. 1988 . Geology and paleoenvironment of the Oligocene Jebel Qatrani Formation and adjacent rocks, Fayum Depression, Egypt. United States Geological Survey . Professional Paper , 1452 : 1 – 64 .
   Google Scholar
• Cifelli , R. L. 1982 . The petrosal structure of Hyopsodus with respect to that of some other ungulates, and its phylogenetic implications . Journal of Paleontology , 56 : 795 – 805 .
   Web of Science ®Google Scholar
• Coiffait , P. E. , Coiffait , B. , Jaeger , J.-J. and Mahboubi , M. 1984 . Un noveau gisement à mammifères fossiles d'âge Eocène supérieur sur le versant sud des Namencha (Algérie orientale): découverte des plus anciens ronguers d'Afrique. Comptes Rendus de l'Académie des Sciences . Série II , 299 : 893 – 898 .
   Google Scholar
• Court , N. 1994a . Periotic anatomy of Moeritherium: homology or homoplasy in the auditory region of Tethytheria? . Zoological Journal of the Linnean Society , 112 : 13 – 28 .
   Web of Science ®Google Scholar
• Court , N. 1994b . Stance and gait in Numidotherium koholense, a primitive proboscidean from the Eocene of Algeria . Zoological Journal of the Linnean Society , 111 : 297 – 338 .
   Google Scholar
• Court , N. and Jaeger , J.-J. 1991 . Anatomy of the periotic bone in Numidotherium koholense: an example of parallel evolution in the inner ear of tethytheres. Comptes Rendus de l'Académie des Sciences . Série II , 312 : 559 – 565 .
   Google Scholar
• Court , N. and Mahboubi , M. 1993 . Reassessment of lower Eocene Seggeurius amourensis: aspects of primitive dental morphology in the mammalian order Hyracoidea . Journal of Paleontology , 67 : 889 – 893 .
   Web of Science ®Google Scholar
• Domning , D. P. , Ray , C. E. and McKenna , M. C. 1986 . Two new Oligocene desmostylians and a discussion of tethytherian systematics . Smithsonian Contributions to Paleobiology , 59 : 1 – 56 .
   Google Scholar
• Fischer , M. S. 1990 . Un trait unique de l'oreille des éléphants et des siréniens (Mammalia): un paradox phylogénétique. Comptes Rendus de l'Académie des Sciences . Série II , 311 : 157 – 162 .
   Google Scholar
• Fischer , M. S. and Tassy , P. 1993 . “ The interrelations between Proboscidea, Sirenia, Hyracoidea and Mesaxonia: the morphological evidence; ” . In Mammal Phylogeny: Placentals Edited by: Szalay , F. S. , Novacek , M. J. and McKenna , M. C. 217 – 243 . New York : Springer-Verlag .
   Google Scholar
• Gazin , C. L. 1965 . A study of the early Tertiary condylarthran mammal Meniscotherium . Smithsonian Miscellaneous Collections , 149 : 1 – 98 .
   Google Scholar
• Gazin , C. L. 1968 . A study of the condylarthran mammal Hyopsodus . Smithsonian Miscellaneous Collections , 153 : 1 – 90 .
   Google Scholar
• Gingerich , P. D. , Russell , D. E. and Wells , N. A. 1990 . Astragalus of Anthracobune (Mammalia, Proboscidea) from the early middle Eocene of Kashmir. Contributions from the Museum of Paleontology . University of Michigan , 28 : 71 – 77 .
   Google Scholar
• Harris , J. M. 1973 . Prodeinotherium from Gebel Zelten, Libya. Bulletin of the British Museum (Natural History) . Geology , 23 : 211 – 274 .
   Google Scholar
• Harris , J. M. 1975 . Evolution of the feeding mechanism in the family Deinotheriidae (Mammalia: Proboscidea) . Zoological Journal of the Linnean Society , 56 : 331 – 362 .
   Web of Science ®Google Scholar
• Harris , J. M. 1978 . “ Deinotherioidea and Barytherioidea; ” . In Evolution of African Mammals Edited by: Maglio , V. J. and Cooke , H. B. S. 315 – 332 . Cambridge : Harvard University Press .
   Google Scholar
• Lundberg , J. G. 1972 . Wagner networks and ancestors . Systematic Zoology , 21 : 398 – 413 .
   Google Scholar
• MacInnes , D. G. 1942 . Miocene and post-Miocene Proboscidea from East Africa. Transactions of the Zoological Society . London , 24 : 33 – 106 .
   Google Scholar
• Mahboubi , M. , Ameur , R. , Crochet , J. Y. and Jaeger , J.-J. 1984 . Earliest known proboscidean from the early Eocene of north-west Africa . Nature , 308 : 543 – 544 .
   PubMedGoogle Scholar
• Mahboubi , M. , Ameur , R. , Crochet , J. Y. and Jaeger , J.-J. 1986 . El Kohol (Saharan Atlas, Algeria): a new Eocene mammal locality in North-West Africa. Palaeontographica, Abt . A. , 192 : 15 – 49 .
   Google Scholar
• Matsumoto , H. 1923 . A contribution to the knowledge of Moeritherium . Bulletin of the American Museum of Natural History , 48 : 97 – 139 .
   Google Scholar
• McKenna , M. C. 1975 . “ Towards a phylogenetic classification of the Mammalia; ” . In Phylogeny of the Primates: A Multidisciplinary Approach Edited by: Luckett , W. P. and Szalay , F. S. 21 – 46 . New York : Plenum Press .
   Google Scholar
• Norell , M. A. and Novacek , M. J. 1992 . The fossil record and evolution: comparing cladistic and paleontological evidence for vertebrate history . Science , 255 : 1690 – 1693 .
   PubMed Web of Science ®Google Scholar
• Novacek , M. J. 1986 . The skull of the leptictid insectivorans and the higher level classification of the eutherian mammals . Bulletin of the American Museum of Natural History , 183 : 1 – 111 .
   Web of Science ®Google Scholar
• Novacek , M. J. 1992a . Fossils, topologies, missing data, and the higher level phylogeny of eutherian mammals . Systematic Biology , 41 : 58 – 73 .
   Web of Science ®Google Scholar
• Novacek , M. J. 1992b . Mammalian phylogeny: shaking the tree . Nature , 356 : 121 – 125 .
   PubMed Web of Science ®Google Scholar
• Novacek , M. J. and Wyss , A. R. 1986 . Higher level relationships of the recent eutherian orders: morphological evidence . Cladistics , 2 : 257 – 287 .
   PubMedGoogle Scholar
• Novacek , M. J. , Wyss , A. R. and McKenna , M. C. 1988 . “ The major groups of eutherian mammals; ” . In Phylogeny and Classification of the Tetrapods Edited by: Benton , M. J. 31 – 71 . Oxford, , England : Clarendon Press .
   Google Scholar
• Prothero , D. R. , Manning , E. M. and Fischer , M. 1988 . “ The phylogeny of ungulates; ” . In The Phylogeny and Classification of the Tetrapods Edited by: Benton , M. J. 201 – 234 . Oxford : Clarendon Press .
   Google Scholar
• Radinsky , L. B. 1966 . The adaptive radiation of the phenacodontid condylarths and the origin of the Perissodactyla . Evolution , 20 : 408 – 417 .
   PubMed Web of Science ®Google Scholar
• Rasmussen , D. T. , Bown , T. M. and Simons , E. L. 1992 . “ The Eocene-Oligocene transition in continental Africa; ” . In Eocene-Oligocene Climatic and Biotic Evolution Edited by: Prothero , D. R. and Berggren , W. A. 548 – 566 . Princeton University Press .
   Google Scholar
• Russell , D. E. 1964 . Les mamminfères Paléocène d'Europe. Mémoires du Muséum National d'Histoire Naturelle . Paris , 13 : 1 – 324 .
   Google Scholar
• Savage , R. J. G. 1969 . Early Tertiary mammal localities in southern Libya. Proceedings of the Geological Society . London , 1623 : 89 – 91 .
   Google Scholar
• Savage , R. J. G. 1971 . “ Review of the fossil mammals of Libya; ” . In Symposium on the Geology of Libya Edited by: Gray , C. 215 – 225 . Tripoli : Faculty of Sciences, University of Libya .
   Google Scholar
• Schlosser , M. 1911 . Beiträge zur Kenntnis der oligozänen Landsäugetiere aus dem Fayum, ägypten . Beiträge zur Paläontologie und geologie Österreich-Ungarns und des Orients , 24 : 51 – 167 .
   Google Scholar
• Shoshani , J. , Savage , R. J. G. and West , R. M. 1989 . A new species of Barytherium (Mammalia, Proboscidea) from Africa, and a discussion of early proboscidean systematics and paleoecology: Abstracts of Papers and Posters Rome : Fifth International Theriological Congress . 160
   Google Scholar
• Sudre , J. 1979 . Nouveaux mammifères éocène du Sahara occidental . Palaeovertebrata , 9 : 83 – 115 .
   Google Scholar
• Swofford , D. L. 1991 . PAUP: Phylogenetic Analysis Using Parsimony, Version 3.1 Champaign, Illinois : Computer program distributed by the Illinois Natural History Survey .
   Google Scholar
• Tassy , P. 1981 . Le crâne de Moeritherium (Proboscidea, Mammalia) de l'Eocène de Dor el Talha (Libye) et le problème de la classification phylogénétique du genre dans les Tethytheria McKenna 1975. Bulletin du Muséum National d'Histoire Naturelle . Paris , 4 : 87 – 147 .
   Google Scholar
• Tassy , P. 1982 . Les principales dichotomies dans l'histoire des Proboscidea (Mammalia): une approche phylogénétique. Géobios . Mémoire Spéciale , 6 : 225 – 245 .
   Google Scholar
• Tassy , P. 1988 . The classification of Proboscidea: how many cladistic classifications? . Cladistics , 4 : 43 – 57 .
   PubMed Web of Science ®Google Scholar
• Tassy , P. 1990 . Phylogénie et classification des Proboscidea (Mammalia): historique et actualité . Annales de Paléontologie , 76 : 159 – 224 .
   Google Scholar
• Tassy , P. and Shoshani , J. 1988 . “ The Tethytheria: elephants and their relatives; ” . In Phylogeny and Classification of the Tetrapods (ed.), Systematics Association Special Volume 35B Edited by: Benton , M. J. 283 – 315 . Oxford : Clarendon Press .
   Google Scholar
• Thewissen , J. G. M. and Domning , D. P. 1992 . The role of phenacodontids in the origin of modem orders of ungulate mammals . Journal of Vertebrate Paleontology , 12 : 494 – 504 .
   Google Scholar
• Wells , N. A. and Gingerich , P. D. 1983 . Review of Eocene Anthracobunidae (Mammalia, Proboscidea) with a new genus and species, Jozaria palustris, from the Kuldana Formation of Kohat (Pakistan). Contributions from the Museum of Paleontology . University of Michigan , 26 : 117 – 139 .
   Google Scholar
• Wight , A. W. R. 1980 . “ Palaeogene vertebrate fauna and regressive sediments of Dur at Talhah, Southern Sirt Basin, Libya; ” . In The Geology of Libya Edited by: Salem , M. J. and Busrewil , M. T. Vol. 1 , 309 – 325 . London : Academic Press .
   Google Scholar
• Williamson , T. E. and Lucas , S. G. 1992 . Meniscotherium (Mammalia, “Condylarthra”) from the Paleocene-Eocene of western North American . Bulletin of the New Mexico Museum of Natural History and Science , 1 : 1 – 75 .
   Google Scholar