Emended diagnosis and phylogenetic relationships of the Upper Cretaceous fossil snake Najash rionegrina Apesteguía and Zaher, 2006

LITERATURE CITED

• Albino , A. M. 2011 . Evolution of Squamata Reptiles in Patagonia based on the fossil record . Biological Journal of the Linnean Society , 103 : 441 – 457 .
   Web of Science ®Google Scholar
• Albino , A. M. and Caldwell , M. W. 2003 . Hábitos de vida de la serpiente cretácica Dinilysia patagonica Woodward . Ameghiana , 40 : 407 – 414 .
   Google Scholar
• Apesteguía , S. and Zaher , H. A. 2006 . A Cretaceous terrestrial snake with robust hindlimbs and a sacrum . Nature , 440 : 1037 – 1040 .
   PubMed Web of Science ®Google Scholar
• Apesteguía , S. , Makovicky , P. J. , Gaetano , L. , Smith , N. and Rougier , G. W. In press. Cerro Bandera: una nueva localidad con micro y mesovertebrados limayenses en Río Negro y la distribución de la fauna de ‘La Buitrera’. XXIII Jornadas Argentinas de Paleontología de Vertebrados . Ameghiniana ,
   Google Scholar
• Bolkay , St. J. 1925 . Mesophis nopcsai n. g., n. sp., ein neues, schlangenahnliches Reptil aus der unteren Kreide (Neocom) von Bilek-Selista (Ost-Hercegovina) . Glasnika Zemaljskog Mujeza u Bosni i Hercegovini , 27 : 125 – 126 .
   Google Scholar
• Brongniart , A. 1800 . Essai d’une classification naturelle des reptiles. 1ère partie. Establissement des Orders . Bulletin des Sciences, par la Societé Philomatique, Paris , 11 : 81 – 82 .
   Google Scholar
• Budney , L. A. , Caldwell , M. W. and Albino , A. 2006 . Unexpected tooth socket histology in the Cretaceous snake Dinilysia with a review of amniote dental attachment tissues . Journal of Vertebrate Paleontology , 26 : 138 – 145 .
   Web of Science ®Google Scholar
• Caldwell , M. W. 2007 . “ Snake phylogeny, origins, and evolution: the role, impact, and importance of fossils (1869–2006) ” . In Major Transitions in Vertebrate Evolution , Edited by: Anderson , J. S. and Sues , H.-D. 253 – 302 . Bloomington and Indianapolis , Indiana : Indiana University Press .
   Google Scholar
• Caldwell , M. W. and Albino , A. 2002 . Exceptionally preserved skeletons of the Cretaceous snake Dinilysia patagonica, Woodward, 1901 . Journal of Vertebrate Paleontology , 22 : 861 – 866 .
   Web of Science ®Google Scholar
• Caldwell , M. W. and Calvo , J. 2008 . Details of a new skull and articulated cervical column of Dinilysia patagonica Woodward, 1901 . Journal of Vertebrate Paleontology , 28 : 349 – 362 .
   Web of Science ®Google Scholar
• Carpenter , C. C. , Murphy , J. B. and Mitchell , L. A. 1978 . Combat bouts with spur use in the Madagascan boa (Sanzinia madagascariensis) . Herpetologica , 34 : 207 – 212 .
   Web of Science ®Google Scholar
• Conrad , J. L. 2008 . Phylogeny and systematics of Squamata (Reptilia) based on morphology . Bulletin of the American Museum of Natural History , 310 : 1 – 182 .
   Web of Science ®Google Scholar
• Cope , E. D. 1869 . On the reptilian orders Pythonomorpha and Streptosauria . Proceedings of the Boston Society of Natural History , 12 : 250 – 261 .
   Google Scholar
• Estes , R. , Frazzetta , T. H. and Williams , E. E. 1970 . Studies on the fossil snake Dinilysia patagonica Woodward: part 1. Cranial morphology . Bulletin of the Museum of Comparative Zoology , 140 : 25 – 74 .
   Google Scholar
• Estes , R. , de Quieroz , K. and Gauthier , J. 1988 . “ Phylogenetic relationships within Squamata ” . In Phylogenetic Relationships of the Lizard Families , Edited by: Estes , R. and Pregill , G. 119 – 281 . Stanford , California : Stanford University Press .
   Google Scholar
• Felsenstein , J. 1983 . Statistical inference of phylogenies . Journal of the Royal Statistical Society, Series A , 146 : 246 – 272 .
   Web of Science ®Google Scholar
• Felsenstein , J. 1985 . Confidence limits on phylogenies: an approach using the bootstrap . Evolution , 39 : 783 – 791 .
   PubMed Web of Science ®Google Scholar
• Fitch , W. M. 1971 . Toward defining the course of evolution: minimal change for a specific tree topology . Systematic Zoology , 20 : 406 – 416 .
   Google Scholar
• Frazzetta , T. H. 1966 . Studies on the morphology and function of the skull in the Boidae (Serpentes). Part II: morphology and function of the jaw apparatus in Python sebae and Python molurus . Journal of Morphology , 118 : 217 – 296 .
   PubMed Web of Science ®Google Scholar
• Gardner , J. and Cifelli , R. 1999 . A primitive snake from the Cretaceous of Utah . Special Papers in Palaeontology , 60 : 87 – 100 .
   Google Scholar
• Gauthier , J. A. , Kearney , M. , Maisano , J. A. , Rieppel , O. and Behlke , A. D. B. 2012 . Assembling the squamate tree of life: perspectives from the phenotype and the fossil record . Bulletin of the Peabody Museum of Natural History , 53 : 3 – 308 .
   Web of Science ®Google Scholar
• Greene , H. W. 1997 . Snakes: The Evolution of Mystery in Nature , 351 Berkeley , California : University of California Press .
   Google Scholar
• Greer , A. E. 1985 . The relationships of the lizard genera Anelytropsis and Dibamus . Journal of Herpetology , 19 : 116 – 156 .
   Web of Science ®Google Scholar
• Haas , G. 1979 . On a new snakelike reptile from the Lower Cenomanian of Ein Jabrud, near Jerusalem . Bulletin du Muséum National d’Histoire Naturelle, Paris, Series 4 , 1 : 51 – 64 .
   Google Scholar
• Hall , B. K. 2003 . Descent with modification: the unity underlying homology and homoplasy as seen through an analysis of development and evolution . Biological Review , 78 : 409 – 433 .
   PubMed Web of Science ®Google Scholar
• Hillis , D. M. and Bull , J. J. 1993 . An empirical test of bootstrapping as a method for assessing confidence in phylogenetic analyses . Systematic Biology , 42 : 182 – 192 .
   Web of Science ®Google Scholar
• Houssaye , A. , Xu , F. , Helfen , L. , De Buffréénil , V. , Baumbach , T. and Tafforeau , P. 2011 . Three-dimensional pelvis and limb anatomy of the Cenomanian hind-limbed snake Eupodophis descouensi (Squamata, Ophidia) revealed by synchrotron-radiation computed laminography . Journal of Vertebrate Paleontology , 31 : 2 – 7 .
   Web of Science ®Google Scholar
• Lee , M. S. Y. 1997 . The phylogeny of varanoid lizards and the affinities of snakes . Philosophical Transactions of the Royal Society of London, Series B , 352 : 53 – 91 .
   Web of Science ®Google Scholar
• Lee , M. S. Y. 1998 . Convergent evolution and character correlation in burrowing reptiles: towards a resolution of squamate relationships . Biological Journal of the Linnean Society , 65 : 369 – 453 .
   Web of Science ®Google Scholar
• Lee , M. S. Y. 2005 . Squamate phylogeny, taxon sampling, and data congruence . Organisms, Diversity and Evolution , 5 : 25 – 45 .
   Google Scholar
• Lee , M. S. Y. 2009 . Hidden support from unpromising data sets strongly unites snakes with anguimorph ‘lizards . Journal of Evolutionary Biology , 22 : 1308 – 1316 .
   PubMed Web of Science ®Google Scholar
• Lee , M. S. Y. and Caldwell , M. W. 1998 . Anatomy and relationships of Pachyrhachis, a primitive snake with hindlimbs . Philosophical Transactions of the Royal Society of London, Series B , 353 : 1521 – 1552 .
   Web of Science ®Google Scholar
• Lee , M. S. Y. and Caldwell , M. W. 2000 . Adriosaurus and the affinities of mosasaurs, dolichosaurs, and snakes . Journal of Paleontology , 74 : 915 – 937 .
   Web of Science ®Google Scholar
• Lee , M. S. Y. and Scanlon , J. D. 2002 . Snake phylogeny based on osteology, soft anatomy and ecology . Biological Reviews , 77 : 333 – 401 .
   PubMed Web of Science ®Google Scholar
• Lee , M. S. Y. , Caldwell , M. W. and Scanlon , J. D. 1999 . A second primitive marine snake: Pachyophis woodwardi from the Cretaceous of Bosnia-Herzegovina . Journal of Zoology , 248 : 509 – 520 .
   Web of Science ®Google Scholar
• Linnaeus , C. 1758 . Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis , ii – +824 . Stockholm : Editio decima, reformata. Laurentius Salvius Holmiae .
   Google Scholar
• Maschio , G. F. , Prudente , A. L. C. , Rodriguez , F. S. and Hoogmoed , M. S. 2010 . Food habits of Anilius scytale (Serpentes: Aniliidae) in the Brazilian Amazonia . Zoologia , 27 : 184 – 190 .
   Web of Science ®Google Scholar
• McDowell , S. B. 1975 . A catalogue of the snakes of New Guinea and the Solomons, with special reference to those in the Bernice P. Bishop Museum. Part II. Anilioidea and Pythoninae . Journal of Herpetology , 9 : 1 – 79 .
   Google Scholar
• Nopcsa , F. B. 1923 . Eidolosaurus und Pachyophis. Zwei neue Neocom-Reptilien . Palaeontographica , 65 : 99 – 154 .
   Google Scholar
• O’Shea , M. 2007 . Boas and Pythons of the World , 165 Princeton, Oxford , New Jersey, U.K : Princeton University Press .
   Google Scholar
• Oppel , M. 1811 . Die Ordnungen, Familien, und Gattungen der Reptilien als prodrom einer naturgeschichte derselben , 88 Munich : Akademie der Wissenschaften .
   Google Scholar
• Rage , J.-C. 1982 . La phylogenie des Lepidosauriens (Reptilia): une approche cladistique . Comptes Rendus de l’Academie des Sciences , 294 : 563 – 566 .
   Google Scholar
• Rage , J.-C. and Escuillié , F. 2000 . Un nouveau serpent bipède du Cénomanien (Crétacé). Implications phylètiques . Comptes Rendus de l’Academie des Sciences , 330 : 513 – 520 .
   Web of Science ®Google Scholar
• Rage , J.-C. and Escuillié , F. 2003 . The Cenomanian: stage of hindlimbed snakes . Carnets de Géologie (Notebooks on Geology) , 1 : 1 – 11 .
   Google Scholar
• Rieppel , O. and Head , J. J. 2004 . New specimens of the fossil snake genus Eupodophis Rage and Escuillié, from the Cenomanian (Late Cretaceous) of Lebanon . Memorie della Società Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano , 32 : 1 – 26 .
   Google Scholar
• Rieppel , O. , Zaher , H. , Tchernov , E. and Polcyn , M. J. 2003 . The anatomy and relationships of Haasiophis terrasanctus, a fossil snake with well developed hind limbs from the mid-Cretaceous of the Middle East . Journal of Paleontology , 77 : 536 – 558 .
   Web of Science ®Google Scholar
• Rogers , R. R. , Eberth , D. A. and Fiorillo , A. R. 2007 . Bonebeds: Genesis, Analysis, and Paleobiological Significance , 499 Chicago, Illinois, London , , U.K : The University of Chicago Press .
   Google Scholar
• Romer , A. S. 1956 . Osteology of the Reptiles , 772 Chicago , Illinois : University of Chicago Press .
   Google Scholar
• Sankey , J. T. and Baszio , S. 2008 . Vertebrate Microfossil Assemblages: Their Role in Paleoecology and Paleobiogeography , 278 Indiana : Indiana University Press, Bloomington .
   Google Scholar
• Smith-Woodward , A. 1901 . On some extinct reptiles from Patagonia of the genera Miolania,Dinilysia, and Genyodectes . Proceedings of the Zoological Society of London , 1901 : 169 – 184 .
   Google Scholar
• Swofford , D. L. 2002 . PAUP*: Phylogenetic Analysis Using Parsimony*, Version 4.0b10 , Sunderland , Massachusetts : Sinauer .
   Google Scholar
• Tchernov , E. , Rieppel , O. , Zaher , H. , Polcyn , M. and Jacobs , L. L. 2000 . A fossil snake with limbs . Science , 287 : 2010 – 2012 .
   PubMed Web of Science ®Google Scholar
• Townsend , T. , Larson , A. , Louis , E. J. and Macey , J. R. 2004 . Molecular phylogenetics of Squamata: the position of snakes, amphisbaenians, and dibamids, and the root of the squamate tree . Systematic Biology , 53 : 735 – 757 .
   PubMed Web of Science ®Google Scholar
• Vidal , N. and Hedges , S. B. 2004 . Molecular evidence for a terrestrial origin of snakes . Proceedings of the Royal Society of London, Series B , 271 : S226 – S229 .
   Web of Science ®Google Scholar
• Vidal , N. and Hedges , S. B. 2005 . The phylogeny of squamate reptiles (lizards, snakes, and amphisbaenians) inferred from nine nuclear protein-coding genes . Comptes Rendus Biologies , 328 : 1000 – 1008 .
   PubMed Web of Science ®Google Scholar
• Wilson , J. A. , Mohabey , D. M. , Peters , S. E. and Head , J. J. 2010 . Predation upon hatchling dinosaurs by a new snake from the Late Cretaceous of India . PLoS Biology , 8 : 1 – 10 .
   Web of Science ®Google Scholar
• Witmer , L. M. 1995 . “ The extant phylogenetic bracket and the importance of reconstructing soft tissues in fossils ” . In Functional Morphology in Vertebrate Paleontology , Edited by: Thompson , J. J. 19 – 33 . Cambridge , U.K : Cambridge University Press .
   Google Scholar
• Zaher , H. and Scanferla , C. A. 2011 . The skull of the Upper Cretaceous snake Dinilysia patagonica Smith-Woodward, 1901, and its phylogenetic position revisited . Zoological Journal of the Linnean Society , 164 : 194 – 238 .
   Web of Science ®Google Scholar
• Zaher , H. , Apesteguía , S. and Scanferla , C. A. 2009 . The anatomy of the Upper Cretaceous snake Najash rionegrina Apesteguía and Zaher, 2006, and the evolution of limblessness in snakes . Zoological Journal of the Linnean Society , 156 : 801 – 826 .
   Web of Science ®Google Scholar