Reassessment of the small ‘arctocyonid’ Prolatidens waudruae from the early Paleocene of Belgium, and its phylogenetic relationships with ungulate-like mammals

LITERATURE CITED

• Archibald , J. D. 1982 . “ A study of Mammalia and geology across the Cretaceous–Tertiary Boundary in Garfield County, Montana ” . 286 University of California Publications in Geological Sciences 122 .
   Google Scholar
• Archibald , J. D. 1996 . Fossil evidence for a Late Cretaceous Origin of “Hoofed” Mammals . Science , 272 : 1150 – 1153 .
   PubMed Web of Science ®Google Scholar
• Archibald , J. D. 1998 . “ Archaic ungulates (“Condylarthra”) ” . In Evolution of Tertiary Mammals of North America, Volume 1: Terrestrial Carnivores, Ungulates and Ungulate-like Mammals , Edited by: Janis , C. M. , Scott , K. M. and Jacobs , L. L. 292 – 331 . New York : Cambridge University Press .
   Google Scholar
• Archibald , J. D. and Averianov , A. O. 2003 . The Late Cretaceous placental mammal Kulbeckia . Journal of Vertebrate Paleontology , 23 : 404 – 419 .
   Web of Science ®Google Scholar
• Archibald , J. D. and Averianov , A. O. 2005 . Mammalian faunal succession in the Cretaceous of the Kyzylkum Desert . Journal of Mammalian Evolution , 12 : 9 – 22 .
   Google Scholar
• Archibald , J. D. and Averianov , A. O. 2012 . Phylogenetic analysis, taxonomic revision, and dental ontogeny of the Cretaceous Zhelestidae (Mammalia: Eutheria) . Zoological Journal of the Linnean Society , 164 : 361 – 426 .
   Web of Science ®Google Scholar
• Archibald , J. D. , Schoch , R. M. and Rigby , J. K. Jr . 1983 . A new subfamily, Conacodontinae, a new species, Conacodon kohlbergeri, of the Periptychidae (Condylarthra, Mammalia) . Postilla , 191 : 24
   Google Scholar
• Cifelli , R. L. 1983 . The origin and affinities of the South American Condylarthra and early Tertiary Litopterna (Mammalia) . American Museum Novitates , 2772 : 1 – 49 .
   Google Scholar
• De Bast , E. , Sigé , B. and Smith , T. 2012 . Diversity of the adapisoriculid mammals from the early Palaeocene of Hainin, Belgium . Acta Palaeontologica Polonica , 57 : 35 – 52 .
   Web of Science ®Google Scholar
• Folie , A. , Sigé , B. and Smith , T. 2005 . A new scincomorph lizard from the Palaeocene of Belgium and the origin of Scincoidea in Europe . Naturwissenschaften , 92 : 542 – 546 .
   PubMed Web of Science ®Google Scholar
• Gazin , C. L. 1968 . A study of the Eocene condylarthran mammal Hyopsodus . Smithsonian Miscellaneous Collections , 153 ( 4 ) : 1 – 90 .
   Google Scholar
• Gheerbrant , E. and Astibia , H. 1994 . Un nouveau mammifère du Maastrichtien de Laño (Pays Basque espagnol) . Comptes Rendus de l’Académie des Sciences de Paris, série II , 318 : 1125 – 1131 .
   Google Scholar
• Gheerbrant , E. and Astibia , H. 1999 . The Upper Cretaceous mammals from Laño (Spanish Basque Country) . Estudios del Museo de Ciencias Naturales de Alava , 14 ( Número especial 1 ) : 295 – 323 .
   Google Scholar
• Gingerich , P. D. 1989 . New earliest Wasatchian mammalian fauna from the Eocene of northwestern Wyoming: composition and diversity in a rarely sampled high-floodplain assemblage . University of Michigan Papers on Paleontology , 28 : 1 – 97 .
   Google Scholar
• Gingerich , P. D. , Abbas , S. G. and Arif , M. 1997 . Early Eocene Quettacyon parachai (Condylarthra) from the Ghazij Formation of Baluchistan (Pakistan): oldest Cenozoic land mammal from South Asia . Journal of Vertebrate Paleontology , 17 : 629 – 637 .
   Web of Science ®Google Scholar
• Gingerich , P. D. , Arw , M. , Khan , I. H. and Abbas , S. G. 1998 . “ First Early Eocene land mammals from the upper Ghazij Formation of the Sor Range, Baluchistan ” . In Siwaliks of South Asia. Proceedings of the Third GEOSAS Workshop, Islamabad, Pakistan, 1–5 March 1997 , Edited by: Ghaznavi , M. I. , Raza , S. M. and Hasan , M. T. 1 – 17 . Islamabad : Geological Survey of Pakistan .
   Google Scholar
• Gingerich , P. D. , Arif , M. , Khan , I. H. , Clyde , W. C. and Bloch , J. I. 1999 . Machocyon Abbasz, a new Early Eocene Quettacyonid (Mammalia, Condylarthra) from the middle Ghazij Formation of Mach and Daghari coal fields . Baluchistan (Pakistan). Contributions from the Museum of Paleontology of the University of Michigan , 30 : 233 – 250 .
   Google Scholar
• Godfriaux , I. and Robaszinski , F. 1974 . Le Montien continental et le Dano–Montien marin des sondages de Hainin (Hainaut, Belgique) . Annales de la Société Géologique de Belgique , 97 : 185 – 200 .
   Google Scholar
• Godinot , M. 1980 . La variabilité morphologique dentaire du Landenodon (Condylarthra) de Dormaal (Eocene, Belgique) . Palaeovertebrata, Mémoire Jubilaire R. Lavocat , : 167 – 196 .
   Google Scholar
• Goloboff , P. , Farris , J. and Nixon , K. 2008 . Vol. 24 , 1–13 T.N.T.: A free program for phylogenetic analysis. Cladistics
   Google Scholar
• Hooker , J. J. and Russell , D. E. 2012 . Early Paleogene Lousinidae (Macroscelidea, Mammalia), their relationships and north European diversity . Zoological Journal of the Linnean Society , 164 : 856 – 936 .
   Web of Science ®Google Scholar
• Jepsen , G. L. 1930 . Stratigraphy and paleontology of the Paleocene of the Northeastern Park County, Wyoming . Proceedings of the American Philosophical Society , 69 : 463 – 528 .
   Google Scholar
• Kielan-Jaworowska , Z. 1968 . Preliminary data on the Upper Cretaceous eutherian mammals from Bayn Dzak, Gobi Desert . Palaeontologia Polonica , 19 : 171 – 191 .
   Google Scholar
• Kondrashov , P. E. and Lucas , S. G. 2004 . “ Oxyclaenus from the early Paleocene of New Mexico and the status of the Oxyclaeninae (Mammalia, Arctocyonidae) ” . In Paleogene Mammals , Edited by: Lucas , S. G. , Zeigler , K. E. and Kondrashov , P. E. 21 – 31 . New Mexico Museum of Natural History and Science Bulletin 26 .
   Google Scholar
• Ladevèze , S. , Missiaen , P. and Smith , T. 2010 . First skull of Orthaspidotherium edwardsi (Mammalia, “Condylarthra”) from the late Paleocene of Berru (France) and phylogenetic affinities of the enigmatic European family Pleuraspidotheriidae . Journal of Vertebrate Paleontology , 30 : 1559 – 1578 .
   Web of Science ®Google Scholar
• Lillegraven , J. A. 1969 . Latest Cretaceous mammals of upper part of Edmonton Formation of Alberta, Canada, and review of Marsupial-Placental dichotomy in mammalian evolution . The University of Kansas Paleontological Contributions , 50 ( Vertebrata 12 ) : 1 – 122 .
   Google Scholar
• Linnaeus , C. 1758 . “ Systema Naturae Per Regna Tria Naturae, Secundum Classes, Ordines, Genera, Species, Cum Characteribus, Differentiis, Synonymis, Locis. Vol. 1: Regnum Animale. Editio decima, reformata ” . Stockholm : Laurentii Salvii .
   Google Scholar
• Lofgren , D. L. 1995 . The Bug Creek problem and the Cretaceous– Tertiary transition at McGuire Creek, Montana . University of California Publications in Geological Sciences , 140 : 1 – 185 .
   Google Scholar
• Luo , Z. 1991 . Variability of dental morphology and the relationships of the earliest arctocyonid species . Journal of Vertebrate Paleontology , 11 : 452 – 471 .
   Google Scholar
• Maddison , W. P. and Maddison , D. R. 2010 . “ Mesquite: A Modular System for Evolutionary Analysis, version 2.73 ” . Available at http://mesquiteproject.org. Accessed January 11, 2012.
   Google Scholar
• Marsh , O. C. 1889 . Discovery of Cretaceous Mammalia . American Journal of Science , 38 : 81 – 92 .
   Google Scholar
• Matthew , W. D. 1897 . A revision of the Puerco fauna . Bulletin of the American Museum of Natural History , 9 : 259 – 323 .
   Google Scholar
• Matthew , W. D. 1901 . Additional observations on the Creodonta . Bulletin of the American Museum of Natural History , 14 : 1 – 38 .
   Google Scholar
• Matthew , W. D. 1915 . A review of the Lower Eocene Wasatch and Wind River faunas. Part I.—Order Ferae (Carnivora). Suborder Creodonta . Bulletin of the American Museum of Natural History , 34 : 1 – 103 .
   Google Scholar
• Matthew , W. D. 1918 . “ A review of the Lower Eocene Wasatch and Wind River faunas ” . Part V.—Insectivora (continued), Glires, Edentata. Bulletin of the American Museum of Natural History 38:565– 657.
   Google Scholar
• Matthew , W. D. 1937 . Paleocene faunas of the San Juan Basin, New Mexico . Transactions of the American Philosophical Society , 30 : 1 – 510 .
   Google Scholar
• Matthew , W. D. and Granger , W. 1915 . A revision of the Lower Eocene Wasatch and Wind River faunas . Bulletin of the American Museum of Natural History , 34 : 1 – 103 .
   Google Scholar
• McKenna , M. C. and Bell , S. K. 1997 . “ Classification of Mammals above the Species Level ” . 631 New York : Columbia University Press .
   Google Scholar
• Nessov , L. A. 1985 . [New mammals from the Cretaceous of Kyzylkum] . Vestnik Leningradskogo Universiteta , 17 : 8 – 18 . [Russian]
   Google Scholar
• Nessov , L. A. 1987 . [Results of search and study of Cretaceous and early Paleogene mammals on the territory of SSSR] . Ezhegodnik Vsesoyuznogo Paleontologicheskogo Obshchestva , 30 : 199 – 218 . [Russian]
   Google Scholar
• Nessov , L. A. 1993 . [New Mesozoic mammals of middle Asia and Kazakhstan, and comments about evolution of theriofaunas of Cretaceous coastal plains of ancient Asia] . Trudy Zoologicheskogo Instituta RAN , 249 : 105 – 133 . [Russian]
   Google Scholar
• Nessov , L. A. , Archibald , J. D. and Kielan-Jaworowska , Z. 1998 . “ Ungulate-like mammals from the Late Cretaceous of Uzbekistan and a phylogenetic analysis of Ungulatomorpha ” . In Dawn of the Age of Mammals in Asia , Edited by: Beard , C. K. and Dawson , M. R. 44 – 88 . Bulletin of the Carnegie Museum of Natural History 34 .
   Google Scholar
• Peláez-Campomanes , P. , López-Martínez , N. , Álvarez-Sierra , M. A. and Daams , R. 2000 . The earliest mammal of the European Paleocene: the multituberculate Hainina . Journal of Paleontology , 74 : 701 – 711 .
   Web of Science ®Google Scholar
• Pol , C. , Buscalioni , A. D. , Carballeira , J. , Francés , V. , López Martinez , N. , Marandat , B. , Joaquin Moratalla , J. , Luis Sanz , J. , Sigé , B. and Villate , J. 1992 . Reptiles and mammals from the Late Cretaceous new locality Quintanilla del Coco (Burgos Province, Spain) . Neues Jahrbuch für Geologie und Paläontologie Abhandlungen , 184 : 279 – 314 .
   Google Scholar
• Prothero , D. R. , Manning , E. M. and Fischer , M. 1988 . “ The phylogeny of the ungulates ” . In The Phylogeny and Classification of the Tetrapods, Volume II: Mammals , Edited by: Benton , M. J. 173 – 181 . New York : Springer-Verlag .
   Google Scholar
• Russell , D. E. 1964 . “ Les Mammifères Paléocènes d’Europe ” . In Université de Paris, Paris , 324 Éditions du Muséum .
   Google Scholar
• Russell , D. E. 1982 . Tetrapods of the northwest European Tertiary Basin . Geologisches Jahrbuch , 60 : 5 – 74 .
   Google Scholar
• Schmidt-Kittler , N. 1987 . European levels and correlation tables . Münchner Geowissenschaftliche Abhandlungen A , 10 : 15 – 31 .
   Google Scholar
• Secord , R. 2008 . The Tiffanian Land-Mammal Age (Middle and Late Paleocene) in the Northern Bighorn Basin, Wyoming . University of Michigan Papers on Paleontology , 35 : 192
   Google Scholar
• Sigé , B. and Marandat , B. 1997 . “ Apport à la faune du Paléocène inférieur d’Europe: un plésiadapiforme du Montien de Hainin (Belgique) ” . In Actes du Congrès BiochroM’97, Montpellier, France, 14–17 April 1997 , Edited by: Aguilar , J.-P. , Legendre , S. and Michaux , J. 679 – 686 . Mémoires et Travaux de l’E.P.H.E., Institut de Montpellier, 21 .
   Google Scholar
• Simpson , G. G. 1927 . Mammalian fauna of the Hell Creek Formation of Montana . American Museum Novitates , 267 : 1 – 8 .
   Google Scholar
• Simpson , G. G. 1935 . New Paleocene mammals from the Fort Union of Montana . Proceedings of the United States National Museum , 83 : 221 – 244 .
   Google Scholar
• Simpson , G. G. 1937 . The Fort Union of the Crazy Mountain Field, Montana and its mammalian faunas . United States National Museum Bulletin , 169 : 1 – 287 .
   Google Scholar
• Simpson , G. G. 1945 . The principles of classification and a classification of mammals . Bulletin of the American Museum of Natural History , 85 : vi – xvi . 1 – 350 .
   Google Scholar
• Sloan , R. E. and van Valen , L. 1965 . Cretaceous mammals from Montana . Science , 148 : 220 – 227 .
   PubMed Web of Science ®Google Scholar
• Sudre , J. and Russell , D. E. 1982 . Les mammifères montiens de Hainin (Paléocène moyen de Belgique) Part II: les Condylarthres . Palaeovertebrata , 12 : 173 – 184 .
   Google Scholar
• Steurbaut , E. 1998 . High-resolution holostratigraphy of Middle Paleocene to Early Eocene strata in Belgium and adjacent areas . Palaeontographica Abteilung A , 247 : 91 – 156 .
   Google Scholar
• Swofford , D. L. 2003 . “ PAUP*. Phylogenetic Analysis Using Parsimony (*And Other Methods), version 4.0b10 ” . Sunderland , Massachusetts : Sinauer Associates .
   Google Scholar
• Tabuce , R. , Clavel , J. and Telles Antugues , M. 2011 . A structural intermediate between triisodontids and mesonychians (Mammalia, Acreodi) from the earliest Eocene of Portugal . Naturwissenschaften , 98 : 145 – 155 .
   PubMed Web of Science ®Google Scholar
• Thewissen , J. G. M. 1990 . Evolution of Paleocene and Eocene Phenacodontidae (Mammalia, Condylarthra) . University of Michigan Papers on Paleontology , 29 : 120
   Google Scholar
• Van Valen , L. 1978 . The beginning of the Age of Mammals . Evolutionary Theory , 4 : 45 – 80 .
   Google Scholar
• Weigelt , J. 1960 . Die Arctocyoniden von Walbeck . Freiberger Forschungshefte , 77 : 1 – 241 .
   Google Scholar
• Williamson , T. E. and Carr , T. D. 2007 . Revision of the problematic Early Paleocene genus Oxyclaenus (Mammalia: Oxyclaenidae) and a new species of Carcinodon . Journal of Vertebrate Paleontology , 27 : 973 – 986 .
   Web of Science ®Google Scholar