Aristonectes quiriquinensis, sp. nov., a new highly derived elasmosaurid from the upper Maastrichtian of central Chile

LITERATURE CITED

• Andrews , C. W. 1910 . A Descriptive Catalogue of the Marine Reptiles of the Oxford Clay, Part I , 205 London : British Museum (Natural History) .
   Google Scholar
• Bardet , N. , Godefroit , P. and Sciau , J. 1999 . A new elasmosaurid plesiosaur from the Lower Jurassic of southern France . Palaeontology , 42 : 927 – 952 .
   Web of Science ®Google Scholar
• Benson , R. B. J. , Evans , M. and Druckenmiller , P. S. 2012 . High diversity, low disparity and small body size in plesiosaurs (Reptilia, Sauropterygia) from the Triassic–Jurassic boundary . PLoS ONE , 7 : e31838 doi: 10.1371/journal.pone.0031838
   PubMed Web of Science ®Google Scholar
• Biró-Bagóczky , L. 1982 . Revisión y redefinición de los ‘Estratos de Quiriquina’, Campaniano–Maastrichtiano, en su localidad tipo, en la Isla Quiriquina, 36°37′ Lat. Sur, Chile, Sudamérica, con un perfil complementario en Cocholgüe . Congreso Geológico Chileno No 3, November 6–14, 1982, Concepción, Chile, Actas 1:A29– A64
   Google Scholar
• Blainville , H. M. D. de . 1835 . Description de quelques espèces de reptiles de la Californie précédé de l’analyse d’un système général d’herpétologie et d’amphibiologie . Nouvelles Annales du Muséum d’Histoire Naturelle de Paris, Série 3 , 4 : 233 – 296 .
   Google Scholar
• Broili , F. 1930 . Plesiosaurierreste von der Insel Quiriquina . Neues Jahrbuch für Mineralogie, Geologie und Paläontologie Beilage-Band (B) , 63 : 497 – 514 .
   Google Scholar
• Brown , D. S. 1981 . The English Late Jurassic Plesiosauroidea (Reptilia) and a review of the phylogeny and classification of the Plesiosauria . Bulletin of the British Museum (Natural History), Geology , 4 : 225 – 234 .
   Google Scholar
• Cabrera , A. 1941 . Un Plesiosaurio nuevo del Cretáceo del Chubut . Revista del Museo de La Plata , 2 : 113 – 130 .
   Google Scholar
• Caldwell , M. 1997 . Limb osteology and ossification patterns in Cryptoclidus (Reptilia: Plesiosauroidea) with a review of sauropterygian limbs . Journal of Vertebrate Paleontology , 17 : 295 – 307 .
   Web of Science ®Google Scholar
• Caldwell , M. W. 2002 . From fins to limbs to fins: limb evolution in fossil marine reptiles . American Journal of Medical Genetics , 112 : 236 – 249 .
   PubMed Web of Science ®Google Scholar
• Carpenter , K. 1997 . “ Comparative cranial anatomy of two North American Cretaceous plesiosaurs ” . In Ancient Marine Reptiles , Edited by: Callaway , J. M. and Nicholls , E. L. 191 – 216 . San Diego : Academic Press .
   Google Scholar
• Carpenter , K. 1999 . Revision of North American elasmosaurs from the Cretaceous of the Western Interior . Paludicola , 2 : 148 – 173 .
   Google Scholar
• Carroll , R. L. 1969 . A middle Pennsylvanian captorhinomorph, and the interrelationships of primitive reptiles . Journal of Paleontology , 43 : 151 – 170 .
   Google Scholar
• Carroll , R. L. 1981 . Plesiosaur ancestors from the Upper Permian of Madagascar . Philosophical Transactions of the Royal Society of London, Series B , 293 : 315 – 383 .
   Web of Science ®Google Scholar
• Carroll , R. L. 1988 . Vertebrate Paleontology and Evolution , 698 New York : W. H. Freeman .
   Google Scholar
• Casamiquela , R. 1969 . La presencia en Chile del género Aristonectes Cabrera (Plesiosauria), del Maestrichtiense del Chubut, Argentina. Edad y carácter de la transgresión ‘Rocaense’ . Jornadas Geológicas Argentinas No 4, April 6–16, 1969, Mendoza, Argentina, Actas 1:199–213
   Google Scholar
• Chatterjee , S. and Creisler , B. S. 1994 . Alwalkeria (Theropoda) and Morturneria (Plesiosauria), new names for preoccupied Walkeria Chatterjee, 1987 and Turneria Chatterjee and Small, 1989 . Journal of Vertebrate Paleontology , 14 : 142
   Web of Science ®Google Scholar
• Chatterjee , S. and Small , B. J. 1989 . “ New plesiosaurs from the Upper Cretaceous of Antarctica ” . Edited by: Crame , J. M. 197 – 215 . Origins and Evolution of the Antarctic Biota. Geological Society of London Special Publication 47
   Google Scholar
• Cecioni , G. 1955 . Distribuzione verticale di alcune Kossmaticeratidae della Patagonia cilena . Bolettino de la Societá Geológica Italiana , 74 : 141 – 148 .
   Google Scholar
• Cope , E. D. 1869 . Synopsis of the extinct Batrachia, Reptilia and Aves of North America . Transactions of the American Philosophical Society, New Series , 14 : 1 – 252 .
   Google Scholar
• Cruickshank , A. R. and Fordyce , R. E. 2002 . A new marine reptile (Sauropterygia) from New Zealand: further evidence for a Late Cretaceous austral radiation of cryptocleidid plesiosaurs . Palaeontology , 45 : 557 – 575 .
   Web of Science ®Google Scholar
• Druckenmiller , P. S. and Russell , A. P. 2006 . A new elasmosaurid plesiosaur (Reptilia: Sauropterygia) from the Lower Cretaceous Clearwater Formation, northeastern Alberta, Canada . Paludicola , 5 : 184 – 199 .
   Google Scholar
• Druckenmiller , P. S. and Russell , A. P. 2008 . A phylogeny of Plesiosauria (Sauropterygia) and its bearing on the systematic status of Leptocleidus Andrews, 1922 . Zootaxa , 1863 : 1 – 120 .
   Google Scholar
• Fuenzalida , H. 1956 . Los saurios de la Isla Quiriquina . Noticiario Mensual del Museo Nacional de Historia Natural (Santiago) , I ( 5 ) : 1
   Google Scholar
• Gasparini , Z. 1979 . Comentarios críticos sobre los vertebrados mesozoicos de Chile . : H15 – H32 . Congreso Geológico Chileno No. 2, Actas 3. August 6–11, 1979
   Google Scholar
• Gasparini , Z. and Biró-Bagoczky , L. 1986 . Osteopygis sp. (Reptilia, Testudines, Toxochelyidae). Tortuga Fósil de la Formación Quiriquina, Cretácico Superior, Sur de Chile . Revista Geológica de Chile, Nota Paleontológica , 27 : 85 – 90 .
   Google Scholar
• Gasparini , Z. , Salgado , L. and Casadío , S. 2003b. Maastrichtian plesiosaurs from northern Patagonia . Cretaceous Research , 24 157 – 170 .
   Web of Science ®Google Scholar
• Gasparini , Z. , Bardet , N. , Martin , J. E. and Fernandez , M. 2003a . The elasmosaurid plesiosaur Aristonectes Cabrera from the latest Cretaceous of South America and Antarctica . Journal of Vertebrate Paleontology , 23 : 104 – 115 .
   Web of Science ®Google Scholar
• Gay , C. 1847 . Historia Física y Política de Chile. Zoología, Tomo Primero , 495 Paris : Imprenta Maulde y Renou .
   Google Scholar
• Gay , C. 1848 . Historia Física y Política de Chile. Zoología, Tomo Segundo , 371 Paris : Imprenta Maulde y Renou .
   Google Scholar
• Gay , C. 1854 . Historia Física y Política de Chile. Zoología 3–4 , 186 Paris : Imprenta Maulde y Renou .
   Google Scholar
• Goloboff , P. , Farris , J. and Nixon , K. 2003 . T.N.T.: Tree Analysis Using New Technology. Program and documentation . Available at www.zmuc.dk/public/phylogeny. Accessed July 11, 2008
   Google Scholar
• Hector , J. 1874 . On the fossil reptiles of New Zealand . Transactions of the New Zealand Institute , 6 : 333 – 358 .
   Google Scholar
• Hiller , N. and Mannering , A. 2004 . Elasmosaur (Reptilia: Plesiosauria) skull remains from the Upper Cretaceous of North Canterbury. New Zealand . Records of the Canterbury Museum , 18 : 1 – 7 .
   Google Scholar
• Hiller , N. and Mannering , A. 2005 . An unusual new elasmosaurid plesiosaur (Sauropterygia) from the Upper Haumurian (Maastrichtian) of the South Island, New Zealand . Memoirs of the Queensland Museum , 51 : 27 – 37 .
   Google Scholar
• Hiller , N. , Mannering , A. , Jones , C. and Cruickshank , A. 2005 . The nature of Mauisaurus haasti Hector, 1874 (Reptilia: Plesiosauria) . Journal of Vertebrate Paleontology , 25 : 588 – 601 .
   Web of Science ®Google Scholar
• Jiménez-Huidobro , P. , Otero , R. A. and Rubilar-Rogers , D. 2010 . First evidence of a tylosaurine mosasaur from Chile . Journal of Vertebrate Paleontology, Program and Abstracts , 2010 : 112A
   Google Scholar
• Kear , B. P. 2005 . A new elasmosaurid plesiosaur from the Lower Cretaceous of Queensland, Australia . Journal of Vertebrate Paleontology , 25 : 792 – 805 .
   Web of Science ®Google Scholar
• Kear , B. P. 2007 . Taxonomic clarification of the Australian elasmosaurid genus Eromangasaurus, with reference to other austral elasmosaur taxa . Journal of Vertebrate Paleontology , 27 : 241 – 246 .
   Web of Science ®Google Scholar
• Ketchum , H. F. and Benson , R. B. J. 2010 . Global interrelationships of Plesiosauria (Reptilia, Sauropterygia) and the pivotal role of taxon sampling in determining the outcome of phylogenetic analyses . Biological Reviews , 85 : 361 – 392 .
   PubMed Web of Science ®Google Scholar
• Ketchum , H. F. and Benson , R. B. J. 2011a . A new pliosaurid (Sauropterygia, Plesiosauria) from the Oxford Clay Formation (Middle Jurassic, Callovian) of England: evidence for a gracile, longirostrine grade of early Middle Jurassic pliosaurids . Special Papers in Palaeontology , 89 : 109 – 129 .
   Google Scholar
• Ketchum , H. F. and Benson , R. B. J. 2011b . The cranial anatomy and taxonomy of Peloneustes philarchus (Sauropterygia, Pliosauridae) from the Peterborough Member (Callovian, Middle Jurassic) of the United Kingdom . Palaeontology , 54 : 639 – 665 .
   Web of Science ®Google Scholar
• Lambrecht , K. 1929 . Neogaeornis wetzeli n. g. n. sp., der erste Kreidevogel der südlichen Hemisphäre . Paläontogische Zeitschrift , 11 : 121 – 129 .
   Google Scholar
• O’Gorman , J. P. , Gasparini , Z. and Reguero , M. 2010 . Aristonectes parvidens Cabrera (Sauropterygia, Plesiosauria) from Cape Lamb, Vega Island (Upper Cretaceous), Antarctica . XXXI Scientific Committee on Antarctic Research (SCAR) Open Science Conference, August 3–6, 2010, Buenos Aires, abstract 557
   Google Scholar
• O’Gorman , J. P. , Gasparini , Z. and Salgado , L. 2013 . Postcranial morphology of Aristonectes Cabrera, 1941 (Plesiosauria, Elasmosauridae) from the Upper Cretaceous of Patagonia and Antarctica . Antarctic Science , 25 : 71–82
   Web of Science ®Google Scholar
• O’Gorman , J. P. , Santillana , S. , Reguero , M. and Moly , J. J. 2012 . “ Primer registro de gastrolitos asociados a un espécimen de Aristonectes sp. (Plesiosauria, Elasmosauridae), Isla Seymour (Is. Marambio), Antártida ” . In Abriendo Ventanas al Pasado, Libro de Resúmenes III Simposio—Paleontología en Chile Edited by: Leppe , M. , Aravena , J. C. and Villa-Martínez , R. 121 – 123 . Punta Arenas , Chile October 11–13, 2012
   Google Scholar
• O’Keefe , F. R. 2001 . A cladistic analysis and taxonomic revision of the Plesiosauria (Reptilia: Sauropterygia) . Acta Zoologica Fennica , 213 : 1 – 63 .
   Google Scholar
• O’Keefe , F. R. 2002 . The evolution of plesiosaur and pliosaur morphotypes in the Plesiosauria (Reptilia: Sauropterygia) . Paleobiology , 28 : 101 – 112 .
   Web of Science ®Google Scholar
• O’Keefe , F. R. and Hiller , N. 2006 . Morphologic and ontogenetic patterns in elasmosaur neck length, with comments on the utility of neck length variables . Paludicola , 5 : 206 – 229 .
   Google Scholar
• O’Keefe , F. R. and Street , H. P. 2009 . Osteology of the cryptocleidoid plesiosaur Tatenectes laramiensis, with comments on the taxonomic status of the Cimoliasauridae . Journal of Vertebrate Paleontology , 29 : 48 – 57 .
   Web of Science ®Google Scholar
• O’Keefe , F. R. and Wahl , W. 2003 . Current taxonomic status of the plesiosaur Pantosaurus striatus from the Upper Jurassic of Sundance Formation, Wyoming . Paludicola , 4 : 37 – 46 .
   Google Scholar
• O’Keefe , F. R. , Street , H. P. , Wilhelm , B. C. , Richards , C. D. and Zhu , H. 2011 . A new skeleton of the cryptoclidid plesiosaur Tatenectes laramiensis reveals a novel body shape among plesiosaurs . Journal of Vertebrate Paleontology , 31 : 330 – 33 .
   Web of Science ®Google Scholar
• Olson , S. L. 1992 . Neogaeornis wetzeli Lambrecht, a Cretaceous loon from Chile (Aves: Gaviidae) . Journal of Vertebrate Paleontology , 12 : 122 – 124 .
   Google Scholar
• Osborn , H. F. 1903 . The reptilian subclasses Diapsida and Synapsida and the early history of the Diaptosauria . Memoirs of the American Museum of Natural History , 1 : 451 – 507 .
   Google Scholar
• Otero , R. A. and O’Gorman , J. P. 2013 . Identification of the first postcranial skeleton of Aristonectes Cabrera (Plesiosauroidea, Elasmosauridae) from the upper Maastrichtian of the south-eastern Pacific, based on a bivariate-graphic method . 41 : 86–89 Cretaceous Research
   Google Scholar
• Otero , R. A. , Soto-Acuña , S. and Rubilar-Rogers , D. 2012 . A postcranial skeleton of an elasmosaurid plesiosaur from the Maastrichtian of central Chile, with comments on the affinities of Late Cretaceous plesiosauroids from the Weddellian Biogeographic Province . Cretaceous Research , 37 : 89 – 99 .
   Web of Science ®Google Scholar
• Owen , R. 1860 . On the orders of fossil and recent Reptilia and their distribution in time . Report of the British Association for the Advancement of Science , 29 : 153 – 166 .
   Google Scholar
• Philippi , R. A. 1887 . Los Fósiles Terciarios i Cuartarios de Chile. F . A , : 236 Brockhaus, Leipzig
   Google Scholar
• Romer , A. S. 1956 . 772 Chicago , Illinois : University of Chicago Press . Osteology of the Reptiles
   Google Scholar
• Salazar , C. , Stinnesbeck , W. and Quinzio-Sinn , L. A. 2010 . Ammonites from the Maastrichtian (Upper Cretaceous) Quiriquina Formation in central Chile . Neues Jahrbuch für Geologie und Paläontologie Abhandlungen , 257 : 181 – 236 .
   Web of Science ®Google Scholar
• Sato , T. 2003 . Terminonatator pointeixensis, a new elasmosaur (Reptilia; Sauropterygia) from the Upper Cretaceous of Saskatchewan . Journal of Vertebrate Paleontology , 23 : 89 – 103 .
   Web of Science ®Google Scholar
• Sato , T. , Hasegawa , Y. and Manabe , M. 2006 . A new elasmosaurid plesiosaur from the Upper Cretaceous of Fukushima, Japan . Palaeontology , 49 : 467 – 484 .
   Web of Science ®Google Scholar
• Steinmann , G. , Deecke , W. and Möricke , W. 1895 . Das Alter und die Fauna der Quiriquina-Schichten in Chile . Neues Jahrbuch für Mineralogie Geologie und Paleontologie , 14 : 1 – 118 .
   Google Scholar
• Stinnesbeck , W. 1986 . Zu den faunistischen und palökologischen Verhältnissen in der Quiriquina Formation (Maastrichtium) Zentral-Chiles . Palaeontographica, Abteilung A , 194 : 99 – 237 .
   Google Scholar
• Stinnesbeck , W. 1996 . “ Ammonite extinctions and environmental changes across the Cretaceous-Tertiary Boundary in Central Chile; pp ” . In The Cretaceous-Tertiary Boundary Mass Extinction: Biotic and Environmental Events , Edited by: MacLeod , N. and Keller , G. 289 – 302 . New York : Norton Press .
   Google Scholar
• Stinnesbeck , W. , Ifrim , C. and Salazar , C. 2012 . The last Cretaceous ammonites in Latin America . Acta Paleontologica Polonica , 57 : 717 – 728 .
   Web of Science ®Google Scholar
• Suárez , M. E. and Fritis , O. 2002 . Nuevo registro de Aristonectes sp. (Plesiosauroidea incertae sedis) del Cretácico Tardío de la Formación Quiriquina, Cocholgüe, Chile . Boletín de la Sociedad de Biología de Concepción , 73 : 87 – 93 .
   Google Scholar
• Suárez , M. E. and Otero , R. A. 2009 . “ Nuevos hallazgos de vertebrados marinos en el Campaniano-Maastrichtiano de Loanco, VII Región ” . In I Simposio—Paleontología en Chile , Edited by: Rubilar , A. , Rubilar-Rogers , D. and Gutstein , C. 78 – 82 . Libro de Actas . October 2–3, 2008, Santiago, Chile
   Google Scholar
• Suárez , M. E. , Quinzio , L. A. , Fritis , O. and Bonilla , R. 2003 . Aportes al conocimiento de los vertebrados marinos de la Formación Quiriquina; pp. 1–7 in Congreso Geológico Chileno No. 10, Resúmenes (CD-ROM), October 6–10, 2003, Concepción, Chile .
   Google Scholar
• Vincent , P. , Bardet , N. , Pereda-Suberbiola , X. , Bouya , B. , Amaghzaz , M. and Meslouh , S. 2010 . Zarafasaura oceanis, a new elasmosaurid (Reptilia: Sauropterygia) from the Maastrichtian phosphates of Morocco and the palaeobiogeography of latest Cretaceous plesiosaurs . Gondwana Research , 19 : 1062 – 1073 .
   Web of Science ®Google Scholar
• Welles , S. P. 1943 . Elasmosaurid plesiosaurs with description of new material from California and Colorado . Memoirs of the University of California , 13 : 125 – 254 .
   Google Scholar
• Welles , S. P. 1952 . A review of the North American Cretaceous elasmosaurs . University of California Publications in Geological Sciences , 29 : 47 – 144 .
   Google Scholar
• Welles , S. P. 1962 . A new species of elasmosaur from the Aptian of Colombia and a review of the Cretaceous plesiosaurs . University of California, Publications in Geological Sciences , 44 : 1 – 96 .
   Google Scholar
• Wetzel , W. 1930 . Die Quiriquina-Schichten als Sediment und paläontologischen Archiv . Palaeontographica , 73 : 49 – 106 .
   Google Scholar
• Wiffen , J. and Moisley , W. 1986 . Late Cretaceous reptiles (families Elasmosauridae, Pliosauridae) from the Mangahouanga Stream, North Island, New Zealand . New Zealand Journal of Geology and Geophysics , 29 : 205 – 252 .
   Web of Science ®Google Scholar
• Zinsmeister , W. J. 1979 . “ Biogeographic significance of the Late Mesozoic and early Tertiary molluscan faunas of Seymour Island (Antarctic Peninsula) to the final break-up of Gondwanaland ” . In Historical Biogeography, Plate Tectonics and the Changing Environment , Edited by: Gray , J. and Boucot , A. J. 349 – 355 . Corvallis , Oregon : Oregon State University Press .
   Google Scholar