Nasutoceratops titusi (Ornithischia, Ceratopsidae), a basal centrosaurine ceratopsid from the Kaiparowits Formation, southern Utah

LITERATURE CITED

• Anderson, B. G., R. E. Barrick, M. L. Droser, and K. L. Stadman. 1999. Hadrosaur skin impressions from the Upper Cretaceous Neslen Formation, Book Cliffs, Utah: morphology and paleoenvironmental context; pp. 295–301 in D. D. Gillette (ed.), Vertebrate Paleontology in Utah. Utah Geological Survey, Salt Lake City, Utah.
   Google Scholar
• Barrick, R. E., M. K. Stoskopf, J. D. Marcot, D. A. Russell, and W. J. Showers. 1998. The thermoregulatory functions of the Triceratops frill and horns: heat flow measured with oxygen isotopes. Journal of Vertebrate Paleontology 18:746–750.
   Web of Science ®Google Scholar
• Blakey, R. 2009. Mollweide plate tectonic maps. Available at http://cpgeosystems.com/paleomaps.html Accessed November 10, 2009.
   Google Scholar
• Brown, B. 1916. Corythosaurus casuarius: skeleton, musculature and epidermis. Bulletin of the American Museum of Natural History 35:709–716.
   Google Scholar
• Brown, B. 1917. A complete skeleton of the horned dinosaur Monoclonius, and a description of a second skeleton showing skin impressions. Bulletin of the American Museum of Natural History 37:281–306.
   Google Scholar
• Brown, C. M., A. P. Russell, and M. J. Ryan. 2009. Pattern and transition of surficial bone texture of the centrosaurine frill and their ontogenetic and taxonomic implications. Journal of Vertebrate Paleontology 29:132–141.
   Web of Science ®Google Scholar
• Campione, N. E., and R. Holmes. 2006. The anatomy and homologies of the ceratopsid syncervical. Journal of Vertebrate Paleontology 26:1014–1017.
   Web of Science ®Google Scholar
• Carr, T. D., T. E. Williamson, B. Britt, and K. Stadtman. 2011. Evidence for high taxonomic and morphologic tyrannosauroid diversity in the Late Cretaceous (late Campanian) of the American Southwest and a new short-skulled tyrannosaurid from the Kaiparowits Formation of Utah. Naturwissenschaften 98:241–246.
   PubMed Web of Science ®Google Scholar
• Clayton, K., M. Loewen, S. Sampson, A. Farke, and R. Irmis. 2009. Epiparietal homology within Centrosaurinae (Ornithischia: Ceratopsidae): a re-evaluation based on newly discovered basal taxa. Journal of Vertebrate Paleontology 29(3, Supplement):80A.
   Google Scholar
• Currie, P. J., W. Langston, and D. H. Tanke. 2008. A new species of Pachyrhinosaurus (Dinosauria: Ceratopsidae) from the Upper Cretaceous of Alberta, Canada; pp. 1–108 in P. J. Currie, W. Langston, and D. H. Tanke (eds.), A New Horned Dinosaur from an Upper Cretaceous Bone Bead in Alberta. NRC Research Press, Ottawa, Ontario, Canada.
   Google Scholar
• Davis, M. 2014. Census of dinosaur skin reveals lithology may not be the most important factor in increased preservation of hadrosaurid skin. Acta Palaeontologica Polonica 59:601–605. DOI: 10.4202/app.2012.0077.
   Web of Science ®Google Scholar
• Dodson, P. 1986. Avaceratops lammersi: a new ceratopsid from the Judith River Formation of Montana. Proceedings of the Academy of Natural Sciences, Philadelphia 138:305–317.
   Web of Science ®Google Scholar
• Dodson, P. 1990. On the status of the ceratopsids Monoclonius and Centrosaurus; pp. 231–243 in K. Carpenter and P. J. Currie (eds.), Dinosaur Systematics: Approaches and Perspectives. Cambridge University Press, New York, New York.
   Google Scholar
• Dodson, P. 2013. Ceratopsia increase: history and trends. Canadian Journal of Earth Sciences 50:294–305.
   Web of Science ®Google Scholar
• Dodson, P., and P. J. Currie. 1990. Neoceratopsia; pp. 593–618 in D. B. Weishampel, P. Dodson, and H. Osmólska (eds.), The Dinosauria. University of California Press, Berkeley, California.
   Google Scholar
• Dodson, P., C. A. Forster, and S. D. Sampson. 2004. Ceratopsidae; pp. 494–513 in D. B. Weishampel, P. Dodson, and H. Osmólska (eds.), The Dinosauria, second edition. University of California Press, Berkeley, California.
   Google Scholar
• Eaton, J. G., and R. L. Cifelli. 1988. Preliminary report on the Late Cretaceous mammalian faunas of the Kaiparowits Plateau, Utah. Contributions to Geology, University of Wyoming 26:245–251.
   Google Scholar
• Eaton, J. G., R. L. Cifelli, J. H. Hutchison, J. I. Kirkland, and J. M. Parrish. 1999. Cretaceous vertebrate faunas from the Kaiparowits Plateau, south central Utah; pp 345–353 in D. D. Gillette (ed.), Vertebrate Paleontology in Utah. Utah Geological Survey Miscellaneous Publication, Salt Lake City, Utah.
   Google Scholar
• Eberth, D. A., and A. P. Hamblin. 1993. Tectonic, stratigraphic, and sedimentologic significance of a regional discontinuity in the upper Judith River Group (Belly River Wedge) of southern Alberta, Saskatchewan, and northern Montana. Canadian Journal of Earth Sciences 30:174–200.
   Web of Science ®Google Scholar
• Evans, D. C. 2010. Cranial anatomy and systematics of Hypacrosaurus altispinus, and a comparative analysis of skull growth in lambeosaurine hadrosaurids (Dinosauria: Ornithischia). Zoological Journal of the Linnean Society 159:398–434.
   Web of Science ®Google Scholar
• Farke, A. A. 2004. Horn use in Triceratops (Dinosauria: Ceratopsidae): testing behavioral hypotheses using scale models. Palaeontologia Electronica 7(1):10p. http://palaeo-electronica.org/paleo/2004_3/horn/issue1_04.htm.
   Google Scholar
• Farke, A. A. 2006. Morphology and ontogeny of the cornual sinuses in chasmosaurine dinosaurs (Ornithischia: Ceratopsidae). Journal of Paleontology 80:780–785.
   Web of Science ®Google Scholar
• Farke, A. A., E. D. S. Wolff, and D. H. Tanke. 2009. Evidence of combat in Triceratops. PLoS ONE 4(1):e4252. DOI: 10.1371/journal.pone.0004252.
   PubMed Web of Science ®Google Scholar
• Farke, A. A., M. J. Ryan, P. M. Barrett, D. H. Tanke, D. R. Braman, M. A. Loewen, and M. R. Graham. 2011. A new centrosaurine from the Late Cretaceous of Alberta, Canada, and the evolution of parietal ornamentation in horned dinosaurs. Acta Palaeontologica Polonica 56:691–702.
   Web of Science ®Google Scholar
• Farlow, J. O., and P. Dodson. 1975. The behavioral significance of frill and horn morphology in ceratopsian dinosaurs. Evolution 29:353–361.
   PubMed Web of Science ®Google Scholar
• Forster, C. A. 1990. The cranial morphology of Triceratops, and a preliminary phylogeny of the Ceratopsia. Ph.D. dissertation, University of Pennsylvania, Philadelphia, Pennsylvania, 227 pp.
   Google Scholar
• Forster, C. A. 1996. Species resolution in Triceratops: cladistic and morphometric approaches. Journal of Vertebrate Paleontology 16:259–270.
   Web of Science ®Google Scholar
• Forster, C. A., P. C. Sereno, T. W. Evans, and T. Rowe. 1993. A complete skull of Chasmosaurus mariscalensis (Dinosauria: Ceratopsidae) from the Aguja Formation (Late Campanian) of west Texas. Journal of Vertebrate Paleontology 13:161–170.
   Google Scholar
• Gates, T. A., and S. D. Sampson. 2007. A new species of Gryposaurus (Dinosauria: Hadrosauridae) from the Late Campanian Kaiparowits Formation, southern Utah, U.S.A. Zoological Journal of the Linnean Society 151:351–376.
   Web of Science ®Google Scholar
• Geist, V. 1971. Mountain Sheep. University of Chicago Press, Chicago, Illinois, 383 pp.
   Google Scholar
• Gelman, A., and D. B. Rubin. 1992. Inference from iterative simulation using multiple sequences. Statistical Sciences 7:457–472.
   Google Scholar
• Getty, M. A., M. A. Loewen, A. L. Titus, and S. D. Sampson. 2010. Taphonomy of horned dinosaurs (Ornithischia: Ceratopsidae) from the Late Campanian Kaiparowits Formation, Grand Staircase–Escalante National Monument, Utah; pp. 478–494 in M. J. Ryan, B. J. Chinnery-Allgeier, and D. A. Eberth (eds.), New Perspectives on Horned Dinosaurs, Royal Tyrrell Museum Ceratopsian Symposium, Indiana University Press, Bloomington and Indianapolis, Indiana.
   Google Scholar
• Goodwin, M. B., and A. L. Deino. 1989. The first radiometric ages from the Judith River Formation (Late Cretaceous), Hill County, Montana. Canadian Journal of Earth Sciences 26:1384–1391.
   Web of Science ®Google Scholar
• Gorscak, E., P. M. O'Connor, N. J. Stevens, and E. M. Roberts. 2014. The basal titanosaurian Rukwatitan bisepultus (Dinosauria: Sauropoda) from the middle Cretaceous Galula Formation, Rukwa Rift Basin, southwestern Tanzania. Journal of Vertebrate Paleontology 34:113–1154.
   Web of Science ®Google Scholar
• Hall, J. P., D. L. Wolberg, and S. West. 1988. Dinosaur skin impressions from the Fruitland Formation (Campanian–Maastrichtian) of the Fossil Forest, San Juan Basin, San Juan County, New Mexico. New Mexico Bureau of Mines and Mineral Resources Bulletin 122:23–27.
   Google Scholar
• Hatcher, J. B., O. C. Marsh, and R. S. Lull. 1907. The Ceratopsia. United States Geological Survey Monograph 49:1–300.
   Google Scholar
• Heckert, A. B., S. G. Lucas, and S. E. Krzyzanowski. 2003. Vertebrate fauna of the Late Cretaceous (Judithian) Fort Crittenden Formation, and the age of Cretaceous vertebrate faunas of southeastern Arizona (U.S.A.). Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen 227:343–364.
   Web of Science ®Google Scholar
• Holmes, R. B., C. A. Forster, M. Ryan, and K. M. Sheperd. 2001. A new species of Chasmosaurus from the Dinosaur Park Formation of southern Alberta. Canadian Journal of Earth Sciences 38:1423–1438.
   Web of Science ®Google Scholar
• Huelsenbeck, J. P., and F. Ronquist. 2001. MRBAYES: Bayesian inference of phylogeny. Bioinformatics 17:754–755.
   PubMed Web of Science ®Google Scholar
• Jinnah, Z. A., E. M. Roberts, A. L. Deino, J. S. Larsen, P. K. Link, and C. M. Fanning. 2009. New 40Ar-39Ar and detrital zircon U-Pb ages for the Upper Cretaceous Wahweap and Kaiparowits formations on the Kaiparowits Plateau, Utah: implications for regional correlation, provenance, and biostratigraphy. Cretaceous Research 30:287–299.
   Web of Science ®Google Scholar
• Kear, B. P., and P. M. Barrett. 2011. Reassessment of the Lower Cretaceous (Barremian) pliosauroid Leptocleidus superstes Andrews, 1922 and other plesiosaur remains from the nonmarine Wealden succession of southern England. Zoological Journal of the Linnean Society 161:663–691.
   Web of Science ®Google Scholar
• Kirkland, J. I., and D. D. DeBlieux. 2010. New basal centrosaurine ceratopsian skulls from the Wahweap Formation (Middle Campanian), Grand Staircase–Escalante National Monument, southern Utah; pp. 117–140 in M. J. Ryan, B. J. Chinnery-Allgeier, and D. A. Eberth (eds.), New Perspectives on Horned Dinosaurs, Royal Tyrrell Museum Ceratopsian Symposium, Indiana University Press, Bloomington and Indianapolis, Indiana.
   Google Scholar
• Lambe, L. M. 1904. On the squamoso-parietal crest of two species of horned dinosaurs from the Cretaceous of Alberta. Ottawa Naturalist 18:81–84.
   Google Scholar
• Lambe, L. M. 1913. A new genus and species of Ceratopsia from the Belly River Formation of Alberta. Ottawa Naturalist 27:109–116.
   Google Scholar
• Lambe, L. M. 1914. On the fore-limb of a carnivorous dinosaur from the Belly River Formation of Alberta, and a new genus of Ceratopsia from the same horizon, with remarks on the integument of some Cretaceous herbivorous dinosaurs. Ottawa Naturalist 27:129–135.
   Google Scholar
• Lambe, L. M. 1915. On Eoceratops canadensis, gen. nov., with remarks on other genera of Cretaceous horned dinosaurs. Canada Geological Survey Museum Bulletin 12, Geological Series 24:1–49.
   Google Scholar
• Langston, W. L., Jr. 1975. The ceratopsian dinosaurs and associated lower vertebrates from the St. Mary River Formation (Maastrichtian) at Scabby Butte, southern Alberta. Canadian Journal of Earth Sciences 12:1576–1608.
   Google Scholar
• Larson, P., M. Larson, C. Ott, and R. Bakker. 2007. Skinning a Triceratops. Journal of Vertebrate Paleontology 27(3, Supplement):104A.
   Web of Science ®Google Scholar
• Lee, M. S. Y., and T. H. Worthy. 2012. Likelihood reinstates Archaeopteryx as a primitive bird. Biology Letters 8:299–303.
   PubMed Web of Science ®Google Scholar
• Lee, M. S. Y., A. Cau, D. Naish, and G. J. Dyke. 2014. Morphological clocks in paleontology and a mid-Cretaceous origin of crown Aves. Systematic Biology 63:442–449.
   PubMed Web of Science ®Google Scholar
• Lehman, T. M. 1989. Chasmosaurus mariscalensis, sp. nov., a new ceratopsian dinosaur from Texas. Journal of Vertebrate Paleontology 9:137–162.
   Google Scholar
• Lehman, T. M. 1990. The ceratopsian subfamily Chasmosaurinae: sexual dimorphism and systematics; pp. 211–229 in K. Carpenter and P. J. Currie (eds.), Dinosaur Systematics: Approaches and Perspectives. Cambridge University Press, New York, New York.
   Google Scholar
• Lehman, T. M. 1996. A horned dinosaur from the El Picacho Formation of west Texas, and a review of ceratopsian dinosaurs from the American Southwest. Journal of Paleontology 70:494–508.
   Web of Science ®Google Scholar
• Lehman, T. M. 1997. Late Campanian dinosaur biogeography in the western interior of North America; pp. 223–240 in D. L. Wolberg, E. Stump, and G. D. Rosenburg (eds.), Dinofest International Proceedings. Academy of Natural Sciences, Philadelphia, Pennsylvania.
   Google Scholar
• Lehman, T. M. 2001. Late Cretaceous dinosaur provinciality; pp. 310–328 in D. H. Tanke and K. Carpenter (eds.), Mesozoic Vertebrate Life. Indiana University Press, Bloomington, Indiana.
   Google Scholar
• Lewis, P. O. 2001. A likelihood approach to estimating phylogeny from discrete morphological character data. Systematic Biology 50:913–925.
   PubMed Web of Science ®Google Scholar
• Loewen, M. A., S. D. Sampson, E. K. Lund, A. A. Farke, M. C. Aguillón-Martínez, C. A. de Leon, R. A. Rodríguez-de la Rosa, M. A. Getty, and D. A. Eberth. 2010. Horned dinosaurs (Ornithischia: Ceratopsidae) from the Upper Cretaceous (Campanian) Cerro del Pueblo Formation, Coahuila, Mexico; pp. 99–116 in M. J. Ryan, B. J. Chinnery-Allgeier, and D. A. Eberth (eds.), New Perspectives on Horned Dinosaurs, Royal Tyrrell Museum Ceratopsian Symposium. Indiana University Press, Bloomington and Indianapolis, Indiana.
   Google Scholar
• Lucas, S. G., R. M. Sullivan, and A. P. Hunt. 2006. Re-evaluation of Pentaceratops and Chasmosaurus (Ornithischia: Ceratopsidae) in the Upper Cretaceous of the Western Interior. New Mexico Museum of Natural History and Science Bulletin 35 :367–370.
   Google Scholar
• Lull, R. S. 1933. A revision of the Ceratopsia, or horned dinosaurs. Memoirs of the Peabody Museum of Natural History 3:1–175.
   Google Scholar
• Lull, R. S., and N. E. Wright. 1942. Hadrosaurian dinosaurs of North America. Geological Society of America Special Papers 40:1–242.
   Google Scholar
• Lundrigan, B. 1996. Morphology of horns and fighting behavior in the family Bovidae. Journal of Mammalogy 77:462–475.
   Web of Science ®Google Scholar
• Maddison, W. P., and D. R. Maddison. 2011. Mesquite: a modular system for evolutionary analysis, version 2.75. Available at http://mesquiteproject.org. Accessed September 1, 2014.
   Google Scholar
• Makovicky, P. J., and M. A. Norell. 2006. Yamaceratops dorngobiensis, a new primitive ceratopsian (Dinosauria: Ornithischia) from the Cretaceous of Mongolia. American Museum Novitates 3530:1–41.
   Web of Science ®Google Scholar
• Marsh, O. C. 1888. A new family of horned Dinosauria from the Cretaceous. American Journal of Science 36:477–478.
   Google Scholar
• Marsh, O. C. 1889. Notice of gigantic horned Dinosauria from the Cretaceous. American Journal of Science 38:173–175.
   Google Scholar
• Marsh, O. C. 1890. Additional characters of the Ceratopsidae with notice of new Cretaceous dinosaurs. American Journal of Science 39:418–426.
   Google Scholar
• Marsh, O. C. 1891. Notice of new vertebrate fossils. American Journal of Science 3:265–269.
   Google Scholar
• Mayr, G., D. S. Peters, G. Plodowski, and O. Vogel. 2002. Bristle-like integumentary structures at the tail of the horned dinosaur Psittacosaurus. Naturwissenschaften 89:361–365.
   PubMed Web of Science ®Google Scholar
• McDonald, A. T., and J. R. Horner. 2010. New material of Styracosaurus ovatus from the Two Medicine Formation of Montana; pp. 156–168 in M. J. Ryan, B. J. Chinnery-Allgeier, and D. A. Eberth (eds.), New Perspectives on Horned Dinosaurs, Royal Tyrrell Museum Ceratopsian Symposium. Indiana University Press, Bloomington and Indianapolis, Indiana.
   Google Scholar
• Müller, J., and R. Reisz. 2006. The phylogeny of early eureptiles: comparing parsimony and Bayesian approaches in the investigation of a basal fossil clade. Systematic Biology 55:503–511.
   PubMed Web of Science ®Google Scholar
• O'Malley, J. F. 1924. Evolution of the nasal cavities and sinuses in relation to function. Journal of Laryngology and Otology 39:57–64.
   Google Scholar
• Osborn, H. F. 1911. A dinosaur mummy. American Museum Journal 2:7–11.
   Google Scholar
• Ostrom, J. H. 1964. A functional analysis of jaw mechanics in the dinosaur Triceratops. Postilla 88:1–35.
   Google Scholar
• Ostrom, J. H. 1966. Functional morphology and evolution of the ceratopsian dinosaurs. Evolution 20:290–308.
   PubMed Web of Science ®Google Scholar
• Padian, K., and J. R. Horner. 2010. The evolution of ‘bizarre structures’ in dinosaurs: biomechanics, sexual selection, social selection or species recognition? Journal of Zoology 283:3–17.
   Web of Science ®Google Scholar
• Penkalski, P., and P. Dodson. 1999. The morphology and systematics of Avaceratops, a primitive horned dinosaur from the Judith River Formation (Late Campanian) of Montana, with the description of a second skull. Journal of Vertebrate Paleontology 19:692–711.
   Web of Science ®Google Scholar
• Prieto-Márquez, A. 2010. Global phylogeny of Hadrosauridae (Dinosauria: Ornithopoda) using parsimony and Bayesian methods. Zoological Journal of the Linnean Society 159:435–502.
   Web of Science ®Google Scholar
• Roberts, E. M. 2007. Facies architecture and depositional environments of the Upper Cretaceous Kaiparowits Formation, southern Utah. Sedimentary Geology 197:207–233.
   Web of Science ®Google Scholar
• Roberts, E. M., S. D. Sampson, A. L. Deino, and S. I. Bowring. 2013. The Kaiparowits Formation: a remarkable record of Late Cretaceous terrestrial environments, ecosystems and evolution in Western North America; pp. 85–106 in A. L. Titus and M. A. Loewen (eds.), At the Top of the Grand Staircase: The Late Cretaceous of Southern Utah. Indiana University Press, Bloomington, Indiana.
   Google Scholar
• Rogers, R. R. 1990. Taphonomy of three horned dinosaur bone beds in the Upper Cretaceous Two Medicine Formation of northwestern Montana: evidence for drought-related mortality. Palaios 5:394–413.
   Google Scholar
• Rogers, R. R., C. C. Swisher, and J. R. Horner. 1993. 40Ar/39Ar age and correlation of the nonmarine Two Medicine Formation (Upper Cretaceous), northwestern Montana. Canadian Journal of Earth Sciences 30:1066–1075.
   Web of Science ®Google Scholar
• Ronquist, F., and J. P. Huelsenbeck. 2003. MrBayes 3: Bayesian phylogenetic inference under mixed models. Bioinformatics 19:1572–1574.
   PubMed Web of Science ®Google Scholar
• Ryan, M. J. 2007. A new basal centrosaurine ceratopsid from the Oldman Formation, southeastern Alberta. Journal of Paleontology 81:376–396.
   Web of Science ®Google Scholar
• Ryan, M. J., and A. P. Russell. 2005. A new centrosaurine ceratopsid from the Oldman Formation of Alberta and its implications for centrosaurine taxonomy and systematics. Canadian Journal of Earth Science 42:1369–1387.
   Web of Science ®Google Scholar
• Ryan, M. J., D. C. Evans, and K. M. Shepard. 2012. A new ceratopsid from the Foremost Formation (middle Campanian) of Alberta. Canadian Journal of Earth Sciences 49:1251–1262.
   Web of Science ®Google Scholar
• Ryan, M. J., A. P. Russell, D. A. Eberth, and P. J. Currie. 2001. Taphonomy of a Centrosaurus (Ornithischia: Ceratopsidae) bone bed from the Dinosaur Park Formation (Upper Campanian), Alberta, Canada, with comments on cranial ontogeny. Palaios 16:482–506.
   Web of Science ®Google Scholar
• Sampson, S. D. 1995. Two new horned dinosaurs from the Upper Cretaceous Two Medicine Formation of Montana; with a phylogenetic analysis of the Centrosaurinae (Ornithischia: Ceratopsidae). Journal of Vertebrate Paleontology 15:743–760.
   Web of Science ®Google Scholar
• Sampson, S. D. 1997. Dinosaur combat and courtship; pp. 383–393 in J. O. Farlow and M. K. Brett-Surman (eds.), The Complete Dinosaur. Indiana University Press, Bloomington, Indiana.
   Google Scholar
• Sampson, S. D., and M. A. Loewen. 2010. Unraveling a radiation: a review of the diversity, stratigraphic distribution, biogeography, and evolution of horned dinosaurs (Ornithischia: Ceratopsidae); pp. 405–427 in M. J. Ryan, B. J. Chinnery-Allgeier, and D. A. Eberth (eds.), New Perspectives on Horned Dinosaurs, Royal Tyrrell Museum Ceratopsian Symposium. Indiana University Press, Bloomington and Indianapolis, Indiana.
   Google Scholar
• Sampson, S. D., and M. J. Ryan. 1997. Variation; pp. 773–780 in P. J. Currie and K. Padian (eds.), Encyclopedia of Dinosaurs. Academic Press, San Diego, California.
   Google Scholar
• Sampson, S. D., M. J. Ryan, and D. H. Tanke. 1997. Craniofacial ontogeny in centrosaurine dinosaurs (Ornithischia: Ceratopsidae): taxonomic and behavioral implications. Zoological Journal of the Linnean Society 121:293–337.
   Web of Science ®Google Scholar
• Sampson, S. D., M. A. Loewen, A. A. Farke, E. M. Roberts, and C. A. Forster. 2010. New horned dinosaurs from Utah provide evidence for intracontinental dinosaur endemism. PLoS ONE 5(9):e12292. DOI: 10.1371/journal.pone.0012292.
   PubMed Web of Science ®Google Scholar
• Sampson, S. D., E. K. Lund, M. A. Loewen, A. A. Farke, and K. E. Clayton. 2013. A remarkable short-snouted horned dinosaur from the Late Cretaceous (late Campanian) of southern Laramidia. Proceedings of the Royal Society B: Biological Sciences 280(1766):20131186.
   PubMed Web of Science ®Google Scholar
• Sampson, S. D., M. A. Loewen, E. M. Roberts, J. A. Smith, L. E. Zanno, and T. A. Gates. 2004. Provincialism in Late Cretaceous terrestrial faunas: new evidence from the Campanian Kaiparowits Formation of Utah. Journal of Vertebrate Paleontology 24(3, Supplement):108A.
   Web of Science ®Google Scholar
• Schaffer, W. M., and C. A. Reed. 1972. The co-evolution of social behavior and cranial morphology in sheep and goats (Bovidae, Caprini). Fieldiana 61:1–88.
   Google Scholar
• Sereno, P. C. 1986. Phylogeny of the bird-hipped dinosaurs (order Ornithischia). National Geographic Research 2:234–256.
   Google Scholar
• Spassov, N. B. 1979. Sexual selection and the evolution of horn-like structures of ceratopsian dinosaurs. Paleontology, Stratigraphy, Lithology 11:37–48.
   Google Scholar
• Sternberg, C. M. 1925. Integument of Chasmosaurus belli. Canadian Field-Naturalist 34:108–110.
   Google Scholar
• Sternberg, C. M. 1949. The Edmonton fauna and description of a new Triceratops from the Upper Edmonton Member: phylogeny of the Ceratopsidae. Bulletin of the National Museum of Canada 113:33–46.
   Google Scholar
• Sternberg, C. M. 1950. Pachyrhinosaurus canadensis, representing a new family of Ceratopsia. Bulletin of the National Museum of Canada 118:109–120.
   Google Scholar
• Sweeney, F., and W. M. Boyden. 1993. A first report of the southernmost occurrence of the ceratopsian dinosaur Styracosaurus albertensis, the first found in the United States. Journal of Vertebrate Paleontology 13(3, Supplement):59A.
   Google Scholar
• Tait, J., and B. Brown. 1928. How the Ceratopsia carried and used their head. Transactions of the Royal Society of Canada, Series 3, 22:13–23.
   Google Scholar
• Williamson, T. E. 1997. A new Late Cretaceous (early Campanian) vertebrate fauna from the Allison Member, Menefee Formation, San Juan Basin, New Mexico; pp. 51–59 in S. G. Lucus, J. W. Estep, T. E. Williamson, and G. S. Morgan (eds.), New Mexico's Fossil Record 1. New Mexico Museum of Natural History Science Bulletin 11. New Mexico Museum of Natural History and Science, Albuquerque, New Mexico.
   Google Scholar
• Witmer, L. M. 1990. The craniofacial air sac system of Mesozoic birds (Aves). Zoological Journal of the Linnean Society 100:327–378.
   Web of Science ®Google Scholar
• Witmer, L. M. 1995. The extant phylogenetic bracket and the importance of reconstructing soft tissues in fossils; pp. 19–33 in J. J. Thomason (ed.), Functional Morphology in Vertebrate Paleontology. Cambridge University Press, New York, New York.
   Google Scholar
• Witmer, L. M. 1997a. Craniofacial air sinus systems; pp. 151–159 in P. J. Currie and K. Padian (eds.), The Encyclopedia of Dinosaurs. Academic Press, New York, New York.
   Google Scholar
• Witmer, L. M. 1997b. The evolution of the antorbital cavity in archosaurs: a study in soft tissue reconstruction in the fossil record with an analysis of the function of pneumaticity. Society of Vertebrate Paleontology Memoirs 3, Journal of Vertebrate Paleontology 17(1, Supplement):1–73.
   Web of Science ®Google Scholar
• Witmer, L. M. 1999. The phylogenetic history of paranasal air sinuses; pp. 21–34 in T. Koppe, H. Nagai, and K. W. Alt (eds.), The Paranasal Sinuses of Higher Primates: Development, Function, and Evolution. Quintessence, Chicago, Illinois.
   Google Scholar
• Wolfe, D. G., and J. I. Kirkland. 1998. Zuniceratops christopheri n. gen. & n. sp., a ceratopsian dinosaur from the Moreno Hill Formation (Cretaceous, Turonian) of west-central New Mexico; pp. 303–317 in S. G. Lucas, J. I. Kirkland, and J. W. Jacobs (eds.), Lower and Middle Cretaceous Terrestrial Ecosystems. New Mexico Museum of Natural History and Science Bulletin 14. New Mexico Museum of Natural History and Science, Albuquerque, New Mexico.
   Google Scholar
• Xu, X., K. Wang, X. Zhou, and D. Li. 2010. First ceratopsid dinosaur from China and its biogeographical implications. Chinese Science Bulletin 55:1631–1635.
   Google Scholar
• You, H., and P. Dodson. 2004. Basal ceratopsia; pp. 478–493 in D. B. Weishampel, P. Dodson, and H. Osmólska (eds.), The Dinosauria, second edition. University of California Press, Berkeley, California.
   Google Scholar
• Zanno, L. E., and S. D. Sampson. 2006. A new oviraptorosaur (Theropoda: Maniraptora) from the late Campanian of Utah and the status of the North American Oviraptorosauria. Journal of Vertebrate Paleontology 25:897–904.
   Web of Science ®Google Scholar
• Zanno, L. E., D. J. Varricchio, P. M. O'Connor, A. L. Titus, and M. J. Knell. 2011. A new troodontid theropod, Talos sampsoni gen. et sp. nov., from the Upper Cretaceous Western Interior Basin of North America. PLoS ONE 6(9):e24487. DOI: 10.1371/journal.pone.0024487.
   PubMed Web of Science ®Google Scholar