New late Eocene and Oligocene remains of the flightless, penguin-like plotopterids (Aves, Plotopteridae) from western Washington State, U.S.A.

LITERATURE CITED

• Barnes, L. G., and J. L. Goedert. 2001. Stratigraphy and paleoecology of Oligocene and Miocene desmostylian occurrences in western Washington State, U.S.A. Bulletin of Ashoro Museum of Paleontology 2:7–22.
   Google Scholar
• Baumel, J. J., and L. M. Witmer. 1993. Osteologia; pp. 45–132 in J. J. Baumel, A. S. King, J. E. Breazile, H. E. Evans, and J. C. Vanden Berge (eds.), Handbook of Avian Anatomy: Nomina Anatomica Avium, second edition. Publications of the Nuttall Ornithological Club 23.
   Google Scholar
• Christidis, L., and W. E. Boles. 2008. Systematics and Taxonomy of Australian Birds. CSIRO Publishing, Collingwood, Victoria, Australia.
   Google Scholar
• Dyke, G. J., X. Wang, and M. B. Habib. 2011. Fossil plotopterid seabirds from the Eo-Oligocene of the Olympic Peninsula (Washington State, USA): descriptions and functional morphology. PLoS ONE 6:e25672.
   PubMed Web of Science ®Google Scholar
• Felice, R. N., and P. M. O'Connor. 2014. Ecology and caudal skeletal morphology in birds: the convergent evolution of pygostyle shape in underwater foraging taxa. PLoS ONE 9:e89737.
   PubMed Web of Science ®Google Scholar
• Goedert, J. L. 1988. A new late Eocene species of Plotopteridae (Aves: Pelecaniformes) from northwestern Oregon. Proceedings of the California Academy of Sciences 45:97–102.
   Google Scholar
• Goedert, J. L., and J. Cornish. 2002. A preliminary report on the diversity and stratigraphic distribution of the Plotopteridae (Pelecaniformes) in Paleogene rocks of Washington State, USA; pp. 63–76 in Z. Zhou and F. Zhang (eds.), Proceedings of the 5th Symposium of the Society of Avian Paleontology and Evolution, Beijing, 1–4 June 2000. Science Press, Beijing, China.
   Google Scholar
• Ericson, P. G. P., C. L. Anderson, T. Britton, A. Elzanowski, U. S. Johansson, M. Källersjö, J. I. Ohlson, T. J. Parsons, D. Zuccon, and G. Mayr. 2006. Diversification of Neoaves: integration of molecular sequence data and fossils. Biology Letters 2:543–547.
   PubMed Web of Science ®Google Scholar
• Hackett, S. J., R. T. Kimball, S. Reddy, R. C. K. Bowie, E. L. Braun, M. J. Braun, J. L. Chojnowski, W. A. Cox, K.-L. Han, J. Harshman, C. J. Huddleston, B. D. Marks, K. J. Miglia, W. S. Moore, F. H. Sheldon, D. W. Steadman, C. C. Witt, and T. Yuri. 2008. A phylogenomic study of birds reveals their evolutionary history. Science 320:1763–1767.
   PubMed Web of Science ®Google Scholar
• Hasegawa, Y., S. Isotani, K. Nagai, K. Seki, T. Suzuki, H. Otsuka, M. Ota, and K. Ono. 1979. Preliminary notes on the Oligo-Miocene penguin-like birds from Japan (Parts I–VII). Bulletin of the Kitakyushu Museum of Natural History 1:41–60.
   Google Scholar
• Howard, H. 1969. A new avian fossil from Kern County, California. Condor 71:68–69.
   Web of Science ®Google Scholar
• Jarvis, E. D., S. Mirarab, A. J. Aberer, B. Li, P. Houde, C. Li, S. Y. W. Ho, B. C. Faircloth, B. Nabholz, J. T. Howard, A. Suh, C. C. Weber, R. R. da Fonseca, J. Li, F. Zhang, H. Li, L. Zhou, N. Narula, L. Liu, G. Ganapathy, B. Boussau, M. S. Bayzid, V. Zavidovych, S. Subramanian, T. Gabaldón, S. Capella-Gutiérrez, J. Huerta-Cepas, B. Rekepalli, K. Munch, M. Schierup, et al. (75 further co-authors) 2014. Whole-genome analyses resolve early branches in the tree of life of modern birds. Science 346:1320–1331.
   PubMed Web of Science ®Google Scholar
• Ksepka, D. T., R. E. Fordyce, T. Ando, and C. M. Jones. 2012. New fossil penguins (Aves, Sphenisciformes) from the Oligocene of New Zealand reveal the skeletal plan of stem penguins. Journal of Vertebrate Paleontology 32:235–254.
   Web of Science ®Google Scholar
• Linnaeus, C. 1758. Systema Naturae per Regna Tria Naturae, Secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis, Locis. Tomus I. Editio decima, reformata. Laurentii Salvii, Stockholm, 824 pp.
   Google Scholar
• Mayr, G. 2005. Tertiary plotopterids (Aves, Plotopteridae) and a novel hypothesis on the phylogenetic relationships of penguins (Spheniscidae). Journal of Zoological Systematics and Evolutionary Research 43:61–71.
   Web of Science ®Google Scholar
• Mayr, G., and J. Clarke. 2003. The deep divergences of neornithine birds: a phylogenetic analysis of morphological characters. Cladistics 19:527–553.
   PubMed Web of Science ®Google Scholar
• Mayr, G., J. L. Goedert, and O. Vogel. 2015. Oligocene plotopterid skulls from western North America and their bearing on the phylogenetic affinities of these penguin-like seabirds. Journal of Vertebrate Paleontology 35:e943764.
   Web of Science ®Google Scholar
• Nesbitt, E. A., R. A. Martin, N. P. Carroll, and J. Grieff. 2010. Reassessment of the Zemorrian foraminiferal stage and Juanian molluscan stage north of the Olympic Mountains, Washington State and Vancouver Island. Newsletters on Stratigraphy 43:275–291.
   Web of Science ®Google Scholar
• Olson, S. L. 1977. A Lower Eocene frigatebird from the Green River Formation of Wyoming (Pelecaniformes: Fregatidae). Smithsonian Contributions to Paleobiology 35:1–33.
   Google Scholar
• Olson, S. L. 1980. A new genus of penguin-like pelecaniform bird from the Oligocene of Washington (Pelecaniformes: Plotopteridae). Natural History Museum of Los Angeles County, Contributions in Science 330:51–57.
   Google Scholar
• Olson, S. L., and Y. Hasegawa. 1979. Fossil counterparts of giant penguins from the North Pacific. Science 206:688–689.
   PubMed Web of Science ®Google Scholar
• Olson, S. L., and Y. Hasegawa. 1985. A femur of Plotopterum from the early middle Miocene of Japan (Pelecaniformes: Plotopteridae). Bulletin of the National Science Museum, Tokyo, Series C 11:137–140.
   Google Scholar
• Olson, S. L., and Y. Hasegawa. 1996. A new genus and two new species of gigantic Plotopteridae from Japan (Aves: Plotopteridae). Journal of Vertebrate Paleontology 16:742–751.
   Web of Science ®Google Scholar
• Prothero, D. R., and K. G. Hankins. 2000. Magnetic stratigraphy and tectonic rotation of the Eocene-Oligocene Keasey Formation, northwest Oregon. Journal of Geophysical Research: Solid Earth (1978–2012) 105(B7):16473–16480.
   Web of Science ®Google Scholar
• Prothero, D. R., E. Draus, and C. Burns. 2009. Magnetostratigraphy and tectonic rotation of the Eocene-Oligocene Makah and Hoko River Formations, northwest Washington, USA. International Journal of Geophysics 2009:1–15.
   Google Scholar
• Prothero, D. R., A. Streig, and C. Burns. 2001. Magnetic stratigraphy and tectonic rotation of the upper Oligocene Pysht Formation, Clallam County, Washington. Society for Sedimentary Geology Book 91:224–233.
   Google Scholar
• Sakurai, K., M. Kimura, and T. Katoh. 2008. A new penguin-like bird (Pelecaniformes: Plotopteridae) from the Late Oligocene Tokoro Formation, northeastern Hokkaido, Japan. Oryctos 7:83–94.
   Google Scholar
• Slack, K. E., C. M. Jones, T. Ando, G. L. Harrison, R. E. Fordyce, U. Arnason, and D. Penny. 2006. Early penguin fossils, plus mitochondrial genomes, calibrate avian evolution. Molecular Biology and Evolution 23:1144–1155.
   PubMed Web of Science ®Google Scholar
• Smith, N. D. 2010. Phylogenetic analysis of Pelecaniformes (Aves) based on osteological data: implications for waterbird phylogeny and fossil calibration studies. PLoS ONE 5:e13354.
   PubMed Web of Science ®Google Scholar
• Snavely, P. D., Jr., N. S. MacLeod, and A. R. Niem. 1993. Geologic map of the Cape Flattery, Clallam Bay, Ozette Lake, and Lake Pleasant quadrangles, northwestern Olympic Peninsula, Washington. U.S. Geological Survey Miscellaneous Investigations Series Map I-1946.
   Google Scholar
• Snavely, P. D., Jr., A. R. Niem, N. S. MacLeod, J. E. Pearl, and W. W. Rau. 1980. Makah Formation – a deep-marginal-basin sequence of late Eocene and Oligocene age in the northwestern Olympic Peninsula, Washington. U.S. Geological Survey Professional Paper 1162-B:1–28.
   Google Scholar
• Wells, R. E. 1989. Geologic map of the Cape Disappointment–Naselle River area, Pacific and Wahkiakum counties, Washington. U.S. Geological Survey Miscellaneous Investigations Series Map I-1832.
   Google Scholar
• Wolfe, E. W., and E. H. McKee. 1968. Geology of the Grays River Qaudrangle, Wahkiakum and Pacific counties, Washington. State of Washington, Division of Mines and Geology, Geologic Map GM-4.
   Google Scholar
• Wolfe, E. W., and E. H. McKee. 1972. Sedimentary and igneous rocks of the Grays River quadrangle, Washington. U.S. Geological Survey Bulletin 1335:1–68.
   Google Scholar
• Yuri, T., R. T. Kimball, J. Harshman, R. C. K. Bowie, M. J. Braun, J. L. Chojnowski, K.-L. Han, S. J. Hackett, C.-J. Huddleston, W.-S. Moore, S. Reddy, F. H. Sheldon, D. W. Steadman, C. C. Witt, and E. L. Braun. 2013. Parsimony and model-based analyses of indels in avian nuclear genes reveal congruent and incongruent phylogenetic signals. Biology 2:419–444.
   PubMedGoogle Scholar

• Kaiser, G., Watanabe, J., and Johns, M.. 2015. A new member of the family Plotopteridae (Aves) from the late Oligocene of British Columbia, Canada. Palaeontologia Electronica 18:1–18.
   Google Scholar
• Handling editor: Trevor Worthy.
   Google Scholar