Cryodrakon boreas, gen. et sp. nov., a Late Cretaceous Canadian azhdarchid pterosaur

LITERATURE CITED

• Andres, B., and T. S. Myers. 2012. Lone Star pterosaurs. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 103:383–398. doi: 10.1017/S1755691013000303
   Web of Science ®Google Scholar
• Averianov, A. O. 2007. New records of azhdarchids (Pterosauria, Azhdarchidae) from the Late Cretaceous of Russia, Kazakhstan, and Central Asia. Paleontological Journal 41:189–197. doi: 10.1134/S0031030107020098
   Web of Science ®Google Scholar
• Averianov, A. O. 2010. The osteology of Azhdarcho lancicollis (Nessov, 1984) (Pterosauria, Azhdarchidae) from the late Cretaceous of Uzbekistan. Proceedings of the Zoological Institute, Russian Academy of Sciences 314:264–317.
   Google Scholar
• Averianov, A. O. 2014. Review of taxonomy, geographic distribution, and paleoenvironments of Azhdarchidae (Pterosauria). ZooKeys 432:1–107. doi: 10.3897/zookeys.432.7913
   Web of Science ®Google Scholar
• Averianov, A. O., M. S. Arkhangelsky, and E. M. Pervushov. 2008. A new late Cretaceous azhdarchid (Pterosauria, Azhdarchidae) from the Volga Region. Paleontological Journal 42:634–642. doi: 10.1134/S0031030108060099
   Web of Science ®Google Scholar
• Bennett, S. C. 1996. Year-classes of pterosaurs from the Solnhofen Limestone of Germany: taxonomic and systematic implications. Journal of Vertebrate Paleontology 16:432–444. doi: 10.1080/02724634.1996.10011332
   Web of Science ®Google Scholar
• Bennett, S. C. 2001. Osteology and functional morphology of the Late Cretaceous pterosaur Pteranodon Part I: general description of osteology. Palaeontographica Abteilung A 260:1–112.
   Google Scholar
• Buffetaut, E., D. Grigorescu, and Z. Csiki. 2003. Giant azhdarchid pterosaurs from the terminal Cretaceous of Transylvania (western Romania); pp. 91–104 in E. Buffetaut and J.-M. Mazin (eds.), Evolution and Palaeobiology of Pterosaurs. Geological Society, London, Special Publications 217.
   Google Scholar
• Cai, Z., and F. Wei. 1994. Zhejiangopterus linhaiensis (Pterosauria) from the Upper Cretaceous of Linhai, Zhejiang, China. Vertebrata PalAsiatica 32:181–194.
   Google Scholar
• Carroll, N. 2015. Reassignment of Montanazhdarcho minor as a non-azhdarchid member of the Azhdarchoidea. Journal of Vertebrate Paleontology 35 (Programs and Abstracts):104.
   Google Scholar
• Cohen, J. E., T. C. Hunt, J. A. Fredrickson, J. L. Berry, and R. L. Cifelli. 2018. Azhdarchid pterosaur from the Upper Cretaceous (Turonian) of Utah, USA. Cretaceous Research 86:60–65. doi: 10.1016/j.cretres.2017.12.013
   Web of Science ®Google Scholar
• Currie, P. J., and A. R. Jacobsen. 1995. An azhdarchid pterosaur eaten by a velociraptorine theropod. Canadian Journal of Earth Sciences 32:922–925. doi: 10.1139/e95-077
   Web of Science ®Google Scholar
• Currie, P. J., and E. B. Koppelhus (eds.). 2005. Dinosaur Provincial Park: A Spectacular Ancient Ecosystem Revealed. Indiana University Press, Bloomington, Indiana, 672 pp. + CD supplement.
   Google Scholar
• Currie, P. J., and D. A. Russell. 1982. A giant pterosaur (Reptilia: Archosauria) from the Judith River (Oldman) Formation of Alberta. Canadian Journal of Earth Sciences 19:894–897. doi: 10.1139/e82-074
   Web of Science ®Google Scholar
• Eberth, D. A., J. Ramenazi, E. M. Roberts, R. R. Rogers, D. R. Braman, and D. C. Evans. 2017. Biostratigraphic importance of the new CA-TIMS U-Pb geochronology for the Belly River Group at Dinosaur Provincial Park (Campanian, Alberta, Canada). Geological Society of America Abstracts with Programs 49:5.
   Google Scholar
• Elgin, R. A., and D. W. E. Hone. 2013. Pneumatization of an immature azhdarchoid pterosaur. Cretaceous Research 45:16–24. doi: 10.1016/j.cretres.2013.06.006
   Web of Science ®Google Scholar
• Frey, E., and D. M. Martill. 1996. A reappraisal of Arambourgiania (Pterosauria, Pterodactyloidea): one of the world’s largest flying animals. Neues Jahrbuch für Geologie und Paläontologie Abhandlungen 199:221–247. doi: 10.1127/njgpa/199/1996/221
   Google Scholar
• Godfrey, S. J., and P. J. Currie. 2005. Pterosaurs; pp. 292–311 in P. J. Currie and E. B. Koppelhus (eds.), Dinosaur Provincial Park: A Spectacular Ancient Ecosystem Revealed. Indiana University Press, Bloomington, Indiana.
   Google Scholar
• Habib, M. 2010. 10,000 miles: maximum range and soaring efficiency of azhdarchid pterosaurs. Journal of Paleontology 30(Supplement 2):99A–100A.
   Google Scholar
• Horner, J. R., and P. J. Currie. 1994. Embryonic and neonatal morphology and ontogeny of a new species of Hypacrosaurus (Ornithischia, Lambeosauridae) from Montana and Alberta; pp. 312–336 in K. Carpenter, K. F. Hirsch, and J. R. Horner (eds.), Dinosaur Eggs and Babies. Cambridge University Press, Cambridge, U.K.
   Google Scholar
• Ibrahim, N., D. M., Unwin, D. M., Martill, L., Baidder, and S. Zouhri 2010. A new pterosaur (Pterodactyloidea: Azhdarchidae) from the Upper Cretaceous of Morocco. PLoS ONE 5:e10875. doi: 10.1371/journal.pone.0010875
   PubMed Web of Science ®Google Scholar
• Kaup, J. J. 1834. Versuch einer Eintheilung der Säugethiere in 6 Stämme und der Amphibien in 6 Ordnungen. Isis von Oken. 3:311–315.
   Google Scholar
• Kellner, A. W. A. 2003. Pterosaur phylogeny and comments on the evolutionary history of the group; pp. 105–137 in E. Buffetaut and J.-M. Mazin (eds.), Evolution and Palaeobiology of Pterosaurs. Geological Society, London, Special Publications 217. London.
   Google Scholar
• Kellner, A. W. A., and W. Langston Jr. 1996. Cranial remains of Quetzalcoatlus (Pterosauria, Azhdarchidae) from Late Cretaceous sediments of Big Bend National Park, Texas. Journal of Vertebrate Paleontology 16:222–231. doi: 10.1080/02724634.1996.10011310
   Web of Science ®Google Scholar
• Lawson, D. A. 1975. Pterosaur from the Latest Cretaceous of West Texas: discovery of the largest flying creature. Science 185:947–948. doi: 10.1126/science.187.4180.947
   Web of Science ®Google Scholar
• Longrich, N. R., D. M. Martill, and B. Andres. 2018. Late Maastrichtian pterosaurs from North Africa and mass extinction of Pterosauria at the Cretaceous-Paleogene boundary. PLoS Biology 16:e2001663. doi: 10.1371/journal.pbio.2001663
   PubMed Web of Science ®Google Scholar
• Martill, D. M., and M. Moser. 2018. Topotype specimens probably attributable to the giant azhdarchid pterosaur Arambourgiania philadelphiae (Arambourg 1959); pp. 159–169 in D. W. E. Hone, M. P. Witton, and D. M. Martill (eds.), Evolution and Palaeobiology of Pterosaurs. Geological Society, London, Special Publications 455. London.
   Google Scholar
• Martill, D. M., E. Frey, R. M. Sadaqah, and H. N. 1998. Discovery of the holotype of the giant pterosaur Titanopteryx philadelphiae Arambourg 1959, and the status of Arambourgiania and Quetzalcoatlus. Neues Jahrbuch für Geologie und Paläontologie Abhandlungen 207:57–76. doi: 10.1127/njgpa/207/1998/57
   Google Scholar
• McGowen, M. R., K. Padian, M. A. De Sosa, and R. J. Harmon. 2002. Description of Montanazhdarcho minor, an azhdarchid pterosaur from the Two Medicine Formation (Campanian) of Montana. PaleoBios 22:1–9.
   Google Scholar
• Naish, D., and M. P. Witton. 2017. Neck biomechanics indicate that giant Transylvanian azhdarchid pterosaurs were short-necked arch predators. PeerJ 5:e2908. doi: 10.7717/peerj.2908
   PubMed Web of Science ®Google Scholar
• Nessov, L. A. 1984. Upper Cretaceous pterosaurs and birds from central Asia. Paleontological Journal 1984:38–49.
   Google Scholar
• Ősi, A., E. Buffetaut, and E. Prondvai. 2011. New pterosaurian remains from the Late Cretaceous (Santonian) of Hungary (Iharkút, Csehbánya Formation). Cretaceous Research 32:456–463. doi: 10.1016/j.cretres.2011.01.011
   Web of Science ®Google Scholar
• Ősi, A., D. B. Weishampel, and C. M. Jianu. 2005. First evidence of azhdarchid pterosaurs from the Late Cretaceous of Hungary. Acta Palaeontologica Polonica 50:777–787.
   Web of Science ®Google Scholar
• Padian, K., A. J. de Ricqles, and J. R. Horner. 1995. Bone histology determines identification of a new fossil taxon of pterosaur (Reptilia: Archosauria). Comptes Rendus de l'Académie des Sciences Paris, serie 2 Sciences de la Terre et des Planetes Fascicule A 320:77–84.
   Google Scholar
• Plieninger, F. 1901. Beiträge zur Kenntnis der Flugsaurier. Palaeontographica 48:65–90.
   Google Scholar
• Russell, D. A. 1972. A pterosaur from the Oldman Formation (Cretaceous) of Alberta. Canadian Journal of Earth Sciences 9:1338–1340. doi: 10.1139/e72-119
   Web of Science ®Google Scholar
• Suberbiola, X. P., N. Bardet, S. Jouve, M. Iarochène, B. Bouya, and M. Amaghzaz. 2003. A new azhdarchid pterosaur from the Late Cretaceous phosphates of Morocco; pp. 79–90 in E. Buffetaut and J.-M. Mazin (eds.), Evolution and Palaeobiology of Pterosaurs. Geological Society, London, Special Publications 217. London.
   Google Scholar
• Sullivan, R. M., and D. W. Fowler. 2011. Navajodactylus boerei, n. gen., n. sp. (Pterosauria, ?Azhdarchidae) from the Upper Cretaceous Kirtland Formation (Upper Campanian) of New Mexico. New Mexico Museum of Natural History and Science Bulletin 53:393–404.
   Google Scholar
• Unwin, D. M. 2005. The Pterosaurs from Deep Time. Pi Press, New York, 352 pp.
   Google Scholar
• Vidovic, S. U., and D. M. Martill. 2017. The taxonomy and phylogeny of Diopecephalus kochi (Wagner, 1837) and ‘Germanodactylus rhamphastinus’ (Wagner, 1851); pp. 125–147 in D. W. E. Hone, M. P. Witton, and D. M. Martill (eds.), Evolution and Palaeobiology of Pterosaurs. Geological Society, London, Special Publications 455.
   Google Scholar
• Vremir, M., A. W. A. Kellner, D. Naish, and G. J. Dyke. 2013. A new azhdarchid pterosaur from the Late Cretaceous of the Transylvanian Basin, Romania: implications for azhdarchid diversity and distribution. PLoS ONE 8:e54268. doi: 10.1371/journal.pone.0054268
   PubMed Web of Science ®Google Scholar
• Vremir, M., M. Witton, D. Naish, G. J. Dyke, S. L. Brusatte, M. Norell, and R. Totoianu. 2015. A medium-sized robust-necked azhdarchid pterosaur (Pterodactyloidea: Azhdarchidae) from the Maastrichtian of Pui (Hauteg Basin, Transylvania, Romania). American Museum Novitates 3827:1–16. doi: 10.1206/3827.1
   Google Scholar
• Vullo, R., G. Garcia, P. Godefroit, A. Cincotta, and X. Valentin. 2018. Mistralazhdarcho maggii, gen. et sp. nov., a new azhdarchid pterosaur from the Upper Cretaceous of southeastern France. Journal of Vertebrate Paleontology. doi: 10.1080/02724634.2018.1502670.
   Web of Science ®Google Scholar
• Watabe, M., T. Tsuihiji, S. Suzuki, and K. Tsogtbaatar. 2009. The first discovery of pterosaurs from the Upper Cretaceous of Mongolia. Acta Palaeontologica Polonica 54:231–242. doi: 10.4202/app.2006.0068
   Web of Science ®Google Scholar
• Wellnhofer, P. 1970. Die Pterodactyloidea (Pterosauria) der Oberjura-Plattenkalke Süddeutschlands. Bayerische Akademie der Wissenschaften, Mathematisch Wissenschaftlichen Klasse, Abhandlugen 141:1–133.
   Google Scholar
• Wellnhofer, P. 1991. The Illustrated Encyclopedia of Pterosaurs. Salamander Books, London, 192 pp.
   Google Scholar
• Witton, M. P. 2013. Pterosaurs: Natural History, Evolution, Anatomy. Princeton University Press, Princeton, New Jersey, 304 pp.
   Google Scholar
• Witton, M. P., and M. B. Habib. 2010. On the size and flight diversity of giant pterosaurs, the use of birds as pterosaur analogues and comments on pterosaur flightlessness. PLoS ONE 5:e13982 doi: 10.1371/journal.pone.0013982
   PubMed Web of Science ®Google Scholar
• Witton, M. P., and D. Naish. 2008. A reappraisal of azhdarchid pterosaur functional morphology and paleoecology. PLoS ONE 3:e2271. doi: 10.1371/journal.pone.0002271
   PubMed Web of Science ®Google Scholar