A non-averostran neotheropod vertebra (Dinosauria: Theropoda) from the earliest Jurassic Whitmore Point Member (Moenave Formation) in southwestern Utah

LITERATURE CITED

• Antonietto, L. S., L. E. Park Boush, C. A. Suarez, A. R. C. Milner, and J. I. Kirkland. 2018. The ‘last hurrah of the reigning darwinulocopines’? Ostracoda (Arthropoda, Crustacea) from the Lower Jurassic Moenave Formation, Arizona and Utah, USA. Journal of Paleontology 92:648–660.
   Web of Science ®Google Scholar
• Bakker, R. T. 1986. The Dinosaur Heresies. William Morrow and Co., Inc., New York, 481 pp.
   Google Scholar
• Blackburn, T., P. E. Olsen, S. A. Bowring, N. M. McLean, D. V. Kent, J. Puffer, G. McHone, E. T. Rasbury, and M. Et-Touhami. 2013. Zircon U–Pb geochronology links the end-Triassic mass extinction with the Central Atlantic Magmatic Province. Science 340:941–945.
   PubMed Web of Science ®Google Scholar
• Britt, B. B., F. M. Dalla Vecchia, D. J. Chure, G. F. Engelmann, M. F. Whiting, and R. D. Scheetz. 2018. Caelestiventus hanseni gen. et sp. nov. extends the desert-dwelling pterosaur record back 65 million years. Nature Ecology & Evolution 2:1386–1392.
   PubMed Web of Science ®Google Scholar
• Brochu, C. A. 1996. Closure of neurocentral sutures during crocodilian ontogeny: implications for maturity assessment in fossil archosaurs. Journal of Vertebrate Paleontology 16:49–62.
   Web of Science ®Google Scholar
• Bonaparte, J. F. 1986. Les dinosaures (Carnosaures, Allosauridés, Sauropodes, Cétiosauridés) du Jurassique Moyen de Cerro Cóndor (Chubut, Argentine). Annales de Paléontologie 72:247–289.
   Google Scholar
• Camp, C. L. 1936. A new type of small bipedal dinosaur from the Navajo Sandstone of Arizona. University of California Publications, Bulletin of the Department of Geological Sciences 24:39–56.
   Google Scholar
• Carpenter, K. 1997. A giant coelophysoid (Ceratosauria) theropod from the Upper Triassic of New Mexico, USA. Neues Jahrbuch für Geologie und Paläontologie–Abhandlungen 205:189–208.
   Google Scholar
• Carrano, M. T., J. R. Hutchison, and S. D. Sampson. 2005. New information on Segisaurus halli, a small theropod dinosaur from the Early Jurassic of Arizona. Journal of Vertebrate Paleontology 25:835–849.
   Web of Science ®Google Scholar
• Clark, J. M., and D. E. Fastovsky. 1986. Vertebrate biostratigraphy of the Glen Canyon Group in northern Arizona; pp. 285–301 in K. Padian (ed.), The Beginning of the Age of Dinosaurs: Faunal Changes Across the Triassic–Jurassic Boundary. Cambridge University Press, Cambridge.
   Google Scholar
• Cohen, K. M., D. A. T. Harper, and P. L. Gibbard. 2020. ICS International Chronostratigraphic Chart 2020/03. International Commission on Stratigraphy, IUGS. www.stratigraphy.org (visited: 2020/07/28).
   Google Scholar
• Colbert, E. H. 1989. The Triassic dinosaur Coelophysis. Museum of Northern Arizona Bulletin 57:1–174.
   Google Scholar
• Cornet, B., and G. Waanders. 2006. Palynomorphs indicate Hettangian (Early Jurassic) age for the middle Whitmore Point Member of the Moenave Formation, Utah and Arizona. New Mexico. New Mexico Museum of Natural History and Science Bulletin 37:390–406.
   Google Scholar
• Ezcurra, M. D. 2017. A new early coelophysoid neotheropod from the Late Triassic of northwestern Argentina. Ameghiniana 54:506–538.
   Web of Science ®Google Scholar
• Fronimos, J. A., and J. W. Wilson. 2017. Neurocentral suture complexity and stress distribution in the vertebral column of a sauropod dinosaur. Ameghiniana 54:36–49.
   Web of Science ®Google Scholar
• Gauthier, J. A. 1986. Saurischian monophyly and the origin of birds. Memoirs of the California Academy of Sciences 8:1–55.
   Google Scholar
• Gibson, S. G. 2013. Biodiversity and evolutionary history of †Lophionotus (Neopterygii: †Semionotiformes) from the western United States. Copeia 2013:582–603.
   Web of Science ®Google Scholar
• Griffin, C. T. 2018. Developmental patterns and variation among early theropods. Journal of Anatomy 232:604–640.
   PubMed Web of Science ®Google Scholar
• Griffin, C. T. 2019. Large neotheropods from the Upper Triassic of North America and the early evolution of large theropod body sizes. Journal of Paleontology 93:1010–1030.
   Web of Science ®Google Scholar
• Griffin, C. T., and S. J. Nesbitt. 2016. Anomalously high variation in postnatal development is ancestral for dinosaurs but lost in birds. Proceedings of the National Academy of Sciences 113:14757–14762.
   PubMed Web of Science ®Google Scholar
• Griffin, C. T., and S. J. Nesbitt. 2019. Does the maximum body size of theropods increase across the Triassic–Jurassic boundary? Integrating ontogeny, phylogeny, and body size. The Anatomical Record 303:1158–1169.
   Google Scholar
• Griffin, C. T., M. R. Stocker, C. Colleary, C. M. Stefanic, E. J. Lessner, M. Riegler, K. Formoso, K. Koeller, and S. J. Nesbitt. 2020. Assessing ontogenetic maturity in extinct saurian reptiles. Biological Reviews (early online). https://doi.org/https://doi.org/10.1111.brv.12666.
   Google Scholar
• Harris, J. D., and A. R. C. Milner. 2015. Tracks in Deep Time: the St. George Dinosaur Discovery Site at Johnson Farm. The University of Utah Press, Salt Lake City, Utah, 99 pp.
   Google Scholar
• Harshbarger, J. W., C. A. Repenning, and J. H. Irwin. 1957. Stratigraphy of the uppermost Triassic and the Jurassic rocks of the Navajo Country. United States Geological Survey Professional Paper 291:1–74.
   Google Scholar
• Heckert, A. B., K. E. Zeigler, S. G. Lucas, L. F. Rinehart, and J. D. Harris. 2000. Preliminary description of coelophysoids (Dinosauria: Theropoda) from the Upper Triassic (Revueltian: early–mid Norian) Snyder Quarry, north–central New Mexico. New Mexico Museum of Natural History and Science Bulletin 17:27–32.
   Google Scholar
• Horner, J. R., H. N. Woodward, and A. M. Bailleul. 2015. Mineralized tissues in dinosaurs interpreted as having formed through metaplasia: a preliminary evaluation. Comptes Rendus Palevol 15:176–196.
   Web of Science ®Google Scholar
• Hunt, A. P., S. G. Lucas, A. B. Heckert, R. M. Sullivan, and M. G. Lockley. 1998. Late Triassic dinosaurs from the western United States. Geobios 31:511–531.
   Web of Science ®Google Scholar
• Ikejiri, T. 2012. Histology-based morphology of the neurocentral synchondrosis in Alligator mississippiensis (Archosauria, Crocodylia). The Anatomical Record 295:18–31.
   Google Scholar
• Irmis, R. B. 2007. Axial skeleton ontogeny in the Parasuchia (Archosauria: Pseudosuchia) in its implications for ontogenetic determination in archosaurs. Journal of Vertebrate Paleontology 27:350–361.
   Web of Science ®Google Scholar
• Irmis, R. B. 2011. Evaluating hypotheses for the early diversification of dinosaurs. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 101:397–426.
   Web of Science ®Google Scholar
• Irmis, R. B., S. J. Nesbitt, K. Padian, N. D. Smith, A. H. Turner, D. Woody, and A. Downs. 2007. A Late Triassic dinosauromorph assemblage from New Mexico and the rise of dinosaurs. Science 317:358–361.
   PubMed Web of Science ®Google Scholar
• Jenkins, X. A., J. R. Foster, and R. J. Gay. 2017. First unambiguous dinosaur specimen from the Upper Triassic Chinle Formation in Utah. Geology of the Intermountain West 4:231–242.
   Google Scholar
• Kirkland, J. I., and A. R. C. Milner. 2005. The Moenave Formation at the St. George Dinosaur Discovery Site at Johnson Farm; pp. 8–9 in Tracking Dinosaur Origins: the Triassic/Jurassic Terrestrial Transition Abstracts Volume.
   Google Scholar
• Kirkland, J. I., and A. R. C. Milner. 2006. The Moenave Formation at the St. George Dinosaur Discovery Site at Johnson Farm. New Mexico Museum of Natural History and Science Bulletin 37:289–309.
   Google Scholar
• Kirkland, J. I., A. R. C. Milner, P. E. Olsen, and J. E. Hargrave. 2014. The Whitmore Point Member of the Moenave Formation in its type area in northern Arizona and its age and correlation with the section in St. George, Utah: evidence for two major lacustrine sequences. Utah Geological Association Publication 43:321–356.
   Google Scholar
• Litwin, R. J. 1986. The palynostratigraphy and age of the Chinle and Moenave formations, southwestern U.S.A. Ph.D. dissertation, Pennsylvania State University, State College, Pennsylvania 266 pp.
   Google Scholar
• Lockley, M. G., and G. D. Gierliński. 2006. Diverse vertebrate ichnofaunas containing Anomoepus and other unusual trace fossils from the Lower Jurassic of the western United States: implications for paleoecology and palichnostratigraphy. New Mexico Museum of Natural History and Science Bulletin 37:176–191.
   Google Scholar
• Long, R. A., and P. A. Murry. 1995. Late Triassic (Carnian and Norian) tetrapods from the southwestern United States. New Mexico Museum of Natural History and Science Bulletin 4:1–254.
   Google Scholar
• Lucas, S. G., and A. B. Heckert. 2001. Theropod dinosaurs and the Early Jurassic age of the Moenave Formation, Arizona–Utah, USA. Neues Jahrbuch für Geologie und Paläontologie–Monatshefte 7:435–448.
   Google Scholar
• Lucas, S. G., and L. H. Tanner. 2006. The Springdale Member of the Kayenta Formation, Lower Jurassic of Utah–Arizona. New Mexico Museum of Natural History and Science Bulletin 37:71–76.
   Google Scholar
• Lucas, S. G., L. H. Tanner, and A. B. Heckert. 2005. Tetrapod biostratigraphy and biochronology across the Triassic–Jurassic boundary in northeastern Arizona. New Mexico Museum of Natural History and Science Bulletin 29:83–93.
   Google Scholar
• Lucas, S. G., H. Klein, M. G. Lockley, J. A. Spielmann, G. D. Gierliński, A. P. Hunt, and L. H. Tanner. 2006. Triassic–Jurassic stratigraphic distribution of the theropod footprint ichnogenus Eubrontes. New Mexico Museum of Natural History and Science Bulletin 37:86–93.
   Google Scholar
• Lucas, S. G., L. H. Tanner, L. L. Donohoo-Hurley, J. W. Geissman, A. B. Heckert, and R. E. Weems. 2011. Position of the Triassic–Jurassic boundary and timing of the end-Triassic extinctions on land: data from the Moenave Formation on the southern Colorado Plateau, USA. Palaeogeography, Palaeoclimatology, Palaeoecology 302:194–205.
   Web of Science ®Google Scholar
• Madsen, J. H. 1976. Allosaurus fragilis: a revised osteology. Utah Geological and Mining Survey Bulletin 37:109–117.
   Google Scholar
• Madsen, S. K., J. I. Kirkland, D. D. DeBlieux, V. L. Santucci, P. Inkenbrandt, and J. S. Tweet. 2012. Paleontological resources inventory and monitoring: Arches National Park, Utah. Utah Geological Survey Contract Deliverable to the National Park Service:1–120.
   Google Scholar
• Marsh, A. D., and W. G. Parker. 2020. New dinosauromorph specimens from Petrified Forest National Park and a global biostratigraphic review of Triassic dinosauromorph body fossils. PaleoBios 37:01w536hs.
   Google Scholar
• Marsh, A. D., and T. B. Rowe. 2018. Anatomy and systematics of the sauropodomorph Sarahsaurus aurifontanalis from the Early Jurassic Kayenta Formation. PLoS ONE 13:e0204007.
   PubMed Web of Science ®Google Scholar
• Marsh, A. D., and T. B. Rowe. 2020. A comprehensive anatomical and phylogenetic evaluation of Dilophosaurus wetherilli (Dinosauria: Theropoda) with descriptions of new specimens from the Kayenta Formation of northern Arizona. Journal of Paleontology 94 (Memoir 78):1–103.
   Web of Science ®Google Scholar
• Marsh, A. D., T. B. Rowe, A. Simonetti, D. Stockli, and L. Stockli. 2014. The age of the Kayenta Formation of northeastern Arizona: overcoming the challenges of dating fossil bone. Journal of Vertebrate Paleontology, Program and Abstracts 2014:178.
   Google Scholar
• Marsh, O. C. 1881. Principal characters of American Jurassic dinosaurs, part V. American Journal of Science (Series 3) 21:417–423.
   Google Scholar
• Martínez, R. N., and C. Apaldetti. 2017. A late Norian–Rhaetian coelophysoid neotheropod (Dinosauria, Saurischia) from the Quebrada del Barro Formation, northwestern Argentina. Ameghiniana 54:488–505.
   Web of Science ®Google Scholar
• Martz, J. W., and B. J. Small. 2019. Non-dinosaurian dinosauromorphs from the Chinle Formation (Upper Triassic) of the Eagle Basin, northern Colorado: Dromomeron romeri (Lagerpetidae) and a new taxon, Kwanasaurus williamparkeri (Silesauridae). PeerJ 7:e7551.
   PubMed Web of Science ®Google Scholar
• Martz, J. W., J. I. Kirkland, A. R. C. Milner, W. G. Parker, and V. L. Santucci. 2017. Upper Triassic lithostratigraphy, depositional systems, and vertebrate paleontology across southern Utah. Geology of the Intermountain West 4:99–180.
   Google Scholar
• Milner, A. R. C., and J. I. Kirkland. 2006. Preliminary review of the Early Jurassic (Hettangian) freshwater Lake Dixie fish fauna in the Whitmore Point Member, Moenave Formation in southwest Utah. New Mexico Museum of Natural History and Science Bulletin 37:510–521.
   Google Scholar
• Milner, A. R. C., and J. I. Kirkland. 2007. The case for fishing dinosaurs at the St. George Dinosaur Discovery Site at Johnson Farm. Utah Geological Survey Notes 39:1–3.
   Google Scholar
• Milner, A. R. C., and M. G. Lockley. 2006. History, geology, and paleontology: St. George Dinosaur Discovery Site at Johnson Farm, Utah; pp. 35–48 in Reynolds, R.E. (ed.), Making Tracks Across the Southwest, Abstracts from the 2006 Desert Symposium. California State University, Desert Studies Consortium and LSA Associates, Inc.
   Google Scholar
• Milner, A. R. C., and S. Z. Spears. 2007. Mesozoic and Cenozoic paleoichnology of southwestern Utah. Utah Geological Association Publication 35:1–85.
   Google Scholar
• Milner, A. R. C., M. G. Lockley, and J. I. Kirkland. 2006a. A large collection of well-preserved theropod dinosaur swim tracks from the Lower Jurassic Moenave Formation, St. George, Utah. New Mexico Museum of Natural History and Science Bulletin 37:315–328.
   Google Scholar
• Milner, A. R. C., M. G. Lockley, and S. B. Johnson. 2006b. The story of the St. George Dinosaur Discovery Site at Johnson Farm: an important new Lower Jurassic dinosaur tracksite from the Moenave Formation of southwestern Utah. New Mexico Museum of Natural History and Science Bulletin 37:329–345.
   Google Scholar
• Milner, A. R. C., J. D. Harris, M. G. Lockley, J. I. Kirkland, and N. A. Matthews. 2009. Bird-like anatomy, posture, and behavior revealed by an Early Jurassic theropod dinosaur resting trace. PLoS ONE 4(3):e4591.
   PubMed Web of Science ®Google Scholar
• Milner, A. R. C., T. A. Birthisel, J. I. Kirkland, B. H. Breithaupt, N. A. Matthews, M. G. Lockley, V. L. Santucci, S. Z. Gibson, D. D. DeBlieux, M. Hurlbut, J. D. Harris, and P. E. Olsen. 2012. Tracking Early Jurassic dinosaurs across southwestern Utah and the Triassic–Jurassic transition. Nevada State Museum Paleontological Papers 1:1–107.
   Google Scholar
• Morales, M. 1994. First dinosaur body fossils from the Lower Jurassic Dinosaur Canyon Member, Moenave Formation of northeastern Arizona. Journal of Vertebrate Paleontology, SVP Program and Abstracts Book 1994:39A.
   Google Scholar
• Nesbitt, S. J. 2011. The early evolution of archosaurs: relationships and the origin of major clades. Bulletin of the American Museum of Natural History 352:1–292.
   Web of Science ®Google Scholar
• Nesbitt, S. J., and M. D. Ezcurra. 2015. The early fossil record of dinosaurs in North America: a new neotheropod from the base of the Upper Triassic Dockum Group of Texas. Acta Palaeontologica Polonica 60:513–526.
   Web of Science ®Google Scholar
• Nesbitt, S. J., R. B. Irmis, and W. G. Parker. 2007. A critical re-evaluation of the Late Triassic dinosaur taxa of North America. Journal of Systematic Palaeontology 5:209–243.
   Web of Science ®Google Scholar
• Nesbitt, S. J., N. D. Smith, R. B. Irmis, A. H. Turner, A. Downs, and M. A. Norell. 2009. A complete skeleton of a Late Triassic saurischian and the early evolution of dinosaurs. Science 326:1530–1533.
   PubMed Web of Science ®Google Scholar
• Norman, D. B. 2020. Scelidosaurus harrisonii from the Early Jurassic of Dorset, England: postcranial skeleton. Zoological Journal of the Linnean Society 189:47–157.
   Web of Science ®Google Scholar
• O’Connor, P. M. 2007. The postcranial axial skeleton of Majungasaurus crenatissimus (Theropoda: Abelisauridae) from the Late Cretaceous of Madagascar. Journal of Vertebrate Paleontology 27 (Memoir 8):127–162.
   Web of Science ®Google Scholar
• Olsen, P. E. 1983. Relationship between biostratigraphic subdivisions and igneous activity in the Newark Supergroup. Geological Society of America, Abstracts with Programs 14:71.
   Google Scholar
• Olsen, P. E., and P. M. Galton. 1984. A review of the reptile and amphibian assemblages from the Stormberg of southern Africa, with special emphasis on the footprints and the age of the Stormberg. Palaeontologia Africana 25:87–110.
   Google Scholar
• Olsen, P. E., and K. Padian. 1986. Earliest records of Batrachopus from the southwestern United States, and a revision of some early Mesozoic crocodylomorph ichnogenera; pp. 258–273 in K. Padian (ed.), The Beginning of the Age of Dinosaurs: Faunal Changes Across the Triassic–Jurassic Boundary. Cambridge University Press, Cambridge.
   Google Scholar
• Olsen, P. E., A. R. McCune, and K. S. Thomson. 1982. Correlation of the early Mesozoic Newark Supergroup by vertebrates, principally fishes. American Journal of Science 282:1–44.
   Web of Science ®Google Scholar
• Olsen, P. E., D. V. Kent, H.-D. Sues, C. Koeberl, H. Huber, A. Montanari, E. C. Rainforth, S. J. Fowell, M. J. Szajna, and B. W. Hartline. 2002. Ascent of dinosaurs linked to an iridium anomaly at the Triassic–Jurassic boundary. Science 296:1305–1307.
   PubMed Web of Science ®Google Scholar
• Owen, R. 1842. Report on British fossil reptiles, part II. Report of the British Association of the Advancement of Science 11:60–104.
   Google Scholar
• Padian, K. 1986. On the type material of Coelophysis Cope (Saurischia: Theropoda) and a new specimen from the Petrified Forest of Arizona (Late Triassic: Chinle Formation); pp. 45–60 in K. Padian (ed.), The Beginning of the Age of Dinosaurs: Faunal Changes Across the Triassic–Jurassic Boundary. Cambridge University Press, Cambridge.
   Google Scholar
• Padian, K., and C. L. May. 1993. The earliest dinosaurs. New Mexico Museum of Natural History and Science Bulletin 3:379–381.
   Google Scholar
• Parker, W. G., and R. B. Irmis. 2005. Advances in Late Triassic vertebrate paleontology based on new material from Petrified Forest National Park, Arizona. New Mexico Museum of Natural History and Science Bulletin 29:45–58.
   Google Scholar
• Raath, M. A. 1977. The anatomy of the Triassic theropod Syntarsus rhodesiensis (Saurischia: Podokesauridae) and a consideration of its biology. Ph.D. dissertation, Rhodes University, Grahamstown, South Africa, 233 pp.
   Google Scholar
• Riggs, N. R., and R. C. Blakey. 1993. Early and Middle Jurassic paleogeography and volcanology of Arizona and adjacent areas. Pacific Section SEPM Book 71:347–373.
   Google Scholar
• Rowe, T. 1989. A new species of the theropod dinosaur Syntarsus from the Early Jurassic Kayenta Formation of Arizona. Journal of Vertebrate Paleontology 9:125–136.
   Google Scholar
• Rowe, T. B., H.-D. Sues, and R. R. Reisz. 2010. Dispersal and diversity in the earliest North American sauropodomorph dinosaurs, with a description of a new taxon. Proceedings of the Royal Society B: Biological Sciences 278:1044–1053.
   PubMed Web of Science ®Google Scholar
• Santa Luca, A. P. 1980. The postcranial skeleton of Heterodontosaurus tucki (Reptilia, Ornithischia) from the Stormberg of South Africa. Annals of the South African Museum 79:159–211.
   Google Scholar
• Seeley, H. G. 1887. On the classification of the fossil animals commonly named Dinosauria. Proceedings of the Royal Society of London 43:165–171.
   Google Scholar
• Sereno, P. C. 1998. A rationale for phylogenetic definitions, with application to the higher-level taxonomy of Dinosauria. Neues Jahrbuch für Geologie und Paläontologie–Abhandlungen 210:41–83.
   Google Scholar
• Schudack, M. E. 2006. Basal Jurassic nonmarine ostracods from the Moenave Formation of St. George, Utah. New Mexico Museum of Natural History and Science Bulletin 37:427–431.
   Google Scholar
• Steiner M. 2014. Age of Lower Jurassic Springdale Sandstone of southwestern Utah from its magnetic polarity sequence. Volumina Jurassica 12:23–30.
   Google Scholar
• Stefanic, C. M., and S. J. Nesbitt. 2019. The evolution and role of the hyposphene-hypantrum articulation in Archosauria: phylogeny size and/or mechanics? Royal Society Open Science 6: e190258.
   PubMed Web of Science ®Google Scholar
• Suarez, C. A., T. K. Knobbe, J. L. Crowley, J. I. Kirkland, and A. R. C. Milner. 2017. A chronostratigraphic assessment of the Moenave Formation, USA using C-isotope chemostratigraphy and detrital zircon geochronology: implications for the terrestrial end Triassic extinction. Earth and Planetary Science Letters 475:83–93.
   Web of Science ®Google Scholar
• Sues, H.-D., S. J. Nesbitt, D. S. Berman, and A. C. Henrici. 2011. A late-surviving basal theropod dinosaur from the latest Triassic of North America. Proceedings of the Royal Society B 278:3459–3464.
   PubMed Web of Science ®Google Scholar
• Turner, A. H., and S. J. Nesbitt. 2013. Body size evolution during the Triassic archosauriform radiation. Geological Society Special Publications 379:573–597.
   Google Scholar
• Tykoski, R. S. 1998. The osteology of Syntarsus kayentakatae and its implications for ceratosaurid phylogeny. M.Sc. thesis, The University of Texas at Austin, Austin, Texas, 217 pp.
   Google Scholar
• Vitkus, A. R., K. Chin, J. I. Kirkland, A. R. C. Milner, E. L. Simpson, and E. T. Ellison. 2020. Unusual fossiliferous concretions from lacustrine deposits in the Lower Jurassic Moenave Formation in St. George, Utah, USA: implications for ancient fish mass mortalities. Palaios 35:77–93.
   Web of Science ®Google Scholar
• Welles, S. P. 1954. New Jurassic dinosaur from the Kayenta Formation of Arizona. Geological Society of America Bulletin 65:591–598.
   Web of Science ®Google Scholar
• Welles, S. P. 1970. Dilophosaurus (Reptilia, Saurischia), a new name for a dinosaur. Journal of Paleontology 44:989.
   Google Scholar
• Welles, S. P. 1984. Dilophosaurus wetherilli (Dinosauria, Theropoda): osteology and comparisons. Palaeontographica Abteilung A 185:84–180.
   Google Scholar
• Wilson, J. A. 1999. A nomenclature for vertebral laminae in sauropod dinosaurs and other saurischian dinosaurs. Journal of Vertebrate Paleontology 19:639–653.
   Web of Science ®Google Scholar
• Wilson, J. A., M. D. D’Emic, T. Ikejiri, E. M. Moacdieh, and J. A. Whitlock. 2011. A nomenclature for vertebral fossae in sauropods and other saurischian dinosaurs. PLoS ONE 6:e17114.
   PubMed Web of Science ®Google Scholar
• Wilson, J. A., D. C. Woodruff, J. D. Gardner, H. M. Flora, J. R. Horner, and C. L. Organ. 2016. Vertebral adaptations to large body size in theropod dinosaurs. PLoS ONE 11:e0158962.
   PubMed Web of Science ®Google Scholar
• Wilson, R. F. 1967. Whitmore Point: a new member of the Moenave Formation in Utah and Arizona. Plateau 40:29–40.
   Google Scholar
• Woodruff, D. C. 2014. The anatomy of the bifurcated neural spine and its occurrence within Tetrapoda. Journal of Morphology 275:1053–1065.
   PubMed Web of Science ®Google Scholar
• You, H.-L., Y. Azuma, T. Wang, Y.-M. Wang, and Z.-M. Dong. 2014. The first well-preserved coelophysoid theropod dinosaur from Asia. Zootaxa 3873:233–249.
   PubMed Web of Science ®Google Scholar
• Zahner, M., and W. Brinkmann. 2019. A Triassic averostran-line theropod from Switzerland and the early evolution of dinosaurs. Nature Ecology & Evolution 3:1146–1152.
   PubMed Web of Science ®Google Scholar