The ontogenetic growth of Anhangueridae (Pterosauria, Pterodactyloidea) premaxillary crests as revealed by a crestless Anhanguera specimen

LITERATURE CITED

• Adams, D. C., and E. Otárola-Castillo. 2013. Geomorph: an R package for the collection and analysis of geometric morphometric shape data. Methods in Ecology and Evolution 4:393–399.
   Web of Science ®Google Scholar
• Andres, B., J. Clarck, and X. Xu. 2014. The earliest pterodactyloid and the origin of the group. Current Biology 24:1011–1016.
   PubMed Web of Science ®Google Scholar
• Assine, M. L. 1992. Análise Estratigráfica da Bacia do Araripe, Nordeste do Brasil. Revista Brasileira de Geociências 22:289–300.
   Google Scholar
• Assine, M. L., J. A. Periotto, M. A. Custódio, V. H. Neumann, F. G. Varejão, and P. C. Mescolotti. 2014. Sequências deposicionais do Andar Alagoas da Bacia do Araripe, Nordeste do Brasil. Boletim de Geociências da Petrobrás, Rio de Janeiro 22:3–28.
   Google Scholar
• Bantim, R. A. M., A. A. F. Saraiva, J. M. Sayão. 2014. A new toothed pterosaur (Pterodactyloidea: Anhangueridae) from the Early Cretaceous Romualdo Formation, NE Brazil. Zootaxa 3869:201–223.
   PubMed Web of Science ®Google Scholar
• Bennett, S. C. 1992. Sexual dimorphism of Pteranodon and other pterosaurs, with comments on cranial crests. Journal of Vertebrate Paleontology 12:422–434.
   Google Scholar
• Bennett, S. C. 1994. Taxonomy and systematics of the Late Cretaceous pterosaur Pteranodon (Pterosauria, Pterodactyloidea). Natural History Museum, University of Kansas Occasional Papers 169:1–70.
   Google Scholar
• Bennett, S. C. 2006. Juvenile specimens of the pterosaur Germanodactylus cristatus, with a review of the genus. Journal of Vertebrate Paleontology 26: 872–878.
   Web of Science ®Google Scholar
• Bennett, S. C. 2018. New smallest specimen of the pterosaur Pteranodon and ontogenetic niches in pterosaurs. Journal of Paleontology 92:254–271.
   Web of Science ®Google Scholar
• Campos, D. A., and A. W. A. Kellner. 1985. Panorama of the flying reptiles study in South America. Anais da Academia Brasileira de Ciências 57:453–466.
   Web of Science ®Google Scholar
• Carpenter, K., D. M., Unwin, K. Cloward, C. Miles, and C. Miles. 2003. A new scaphognathine pterosaur from the Upper Jurassic Morrison Formation of Wyoming, USA; pp217:45–54 in E. Buffetaut; J. M. Mazrn (eds) Evolution and Palaeobiology of Pterosaurs. Wyoming. Geological Society of London, Special Publications.
   Google Scholar
• Cerqueira, G. M., M. A. Santos, M. F. Marks, J. M. Sayão, and F. L. Pinheiro. 2021. A new azhdarchoid pterosaur from the Lower Cretaceous of Brazil and the paleobiogeography of the Tapejaridae. Acta Palaeontologica Polonica 66:555–570.
   Web of Science ®Google Scholar
• Duque, R. R. C., and A. M. F. Barreto. 2018. Novos sítios fossilíferos da Formação Romualdo, Cretáceo Inferior, Bacia do Araripe, Exu, Pernambuco, Nordeste do Brasil. Anuário do Instituto de Geociências 41:5–14.
   Google Scholar
• Fastnacht, M. 2001. First record of Coloborhynchus (Pterosauria) from the Santana Formation (Lower Cretaceous) of the Chapada do Araripe, Brazil. Paläontologische Zeitschrift 75:23.
   Google Scholar
• Frey, E., D. M. Martill, and M. C. Buchy. 2003. A new rested ornithocheirid from the lower Cretaceous of northeastern Brazil and the unusual death of an unusual pterosaur; pp. 217:55–63 in E. Buffettaut and J. M. Mazin (eds.) Evolution and Palaeobiology of Pterosaurs, Geological Society Special Publications.
   Google Scholar
• Goloboff, P. A., and S. A. Catalano. 2016. TNT version 1.5, including a full implementation of phylogenetic morphometrics. Cladistics 32:221–238.
   PubMed Web of Science ®Google Scholar
• Holgado, B., and R. V. Pêgas. 2020. A taxonomic and phylogenetic review of the anhanguerid pterosaur group Coloborhynchinae and the new clade Tropeognathinae. Acta Palaeontologica Polonica 65:743–761.
   Web of Science ®Google Scholar
• Holgado, B., R. V. Pêgas, J. I. Canudo, J. Fortuny, T. Rodrigues, J. Company, and A. W. A. Kellner. 2019. On a new crested pterodactyloid from the Early Cretaceous of the Iberian Penisula and the radiation of the clade Anhangueria. Scientific Reports 9:1–10.
   PubMed Web of Science ®Google Scholar
• Hone, D. W., Naish, D. and Cuthill, I. C. 2012. Does mutual sexual selection explain the evolution of head crests in pterosaurs and dinosaurs? Lethaia 45(2):139–156.
   Web of Science ®Google Scholar
• Kaup, J. J. 1834. Versuch einer Eintheilung der Saugethiere in 6 Stämme und der Amphibien in 6 Ordnungen. Isis 3:311–315.
   Google Scholar
• Kellner, A. W. A. 1984. Ocorrência de uma mandíbula de Pterosauria (Brasileodactylus araripensis, nov. gen.; nov. sp.) na Formação Santana, Cretáceo da Chapada do Araripe, Ceará – Brasil. Anais do Congresso Brasileiro de Geologia, 33:578–590.
   Google Scholar
• Kellner, A. W. A. 1990. Os répteis voadores do Cretáceo brasileiro. Anuário do Instituto de Geociências 12:86–106.
   Google Scholar
• Kellner, A. W. A. 1996. Description of new material of Tapejaridae and Anhangueridae (Pterosauria, Pterodactyloidea) and discussion of pterosaur phylogeny. Ph.D. dissertation, Columbia University, New York, 347 pp.
   Google Scholar
• Kellner, A. W. A. 2003. Pterosaur phylogeny and comments on the evolutionary history of the group; pp: 217:105–137 in E. Buffetaut; J. M. Mazin (eds.). Evolution and Paleobiology of Pterosaurs. Geological Society, London, Special Publications.
   Google Scholar
• Kellner, A. W. A., and Y. Tomida. 2000. Description of a new species of Anhangueridae (Pterodactyloidea) with comments on the pterosaur fauna from the Santana Formation (Aptian-Albian), northeastern Brazil. National Science Museum Monographs 17:1–135
   Google Scholar
• Klingenberg, C. P. 2011. MorphoJ: an integrated sohtware package for geometric morphometrics. Molecular Ecology Resources 11:353–357
   PubMed Web of Science ®Google Scholar
• Longrich, N. R., D. M. Martill, and B. Andres. 2018. Late Maastrichtian pterosaurs from North Africa and mass extinction of Pterosauria at the Cretaceous-Paleogene boundary. Plos Biology 16: e2001663
   PubMed Web of Science ®Google Scholar
• Lü, J.; D. M. Unwin, D. C. Deeming, X. Jin, Y. Liu, and Q. Ji. 2011. An egg-adult association, gender, and reproduction in pterosaurs. Science 331:3211–324.
   Web of Science ®Google Scholar
• Manzig, P. C., A. W. A. Kellner, L. C. Weinschütz, C. E. Fragoso, C. S. Vega, G. B. Guimarães, L. C. Godoy, A. Liccardo, J. H. C. Ricetti, and C. C. Moura. 2014. Discovery of a rare pterosaur bone bed in a Cretaceous Desert with insights on ontogeny and behavior of flying reptiles. Plos One 9(8): e100005.
   PubMed Web of Science ®Google Scholar
• May, P., P. Reser, and P. Leiggi. 1994. Macrovertebrates preparation; pp. 113–128 in P. Leiggi; P. May. (eds) Vertebrates Paleontological Techniques, volume I. Cambridge University Press.
   Google Scholar
• Neumann, V. H., and L. Cabrera. 1999. Uma nueva propuesta estratigráfica para la tectonocuencia post-rifte de la Cuenca de Araripe, nordeste de Brasil. In: Simpósio sobre o Cretáceo do Brasil, 5. 1963, Rio Claro, Boletim. Rio Claro: UNESP. 279–285.
   Google Scholar
• Pêgas, R. V., B. Holgado, and M. E. C. Leal. 2019. On Targaryendraco wiedenrothi gen. nov. (Pterodactyloidea, Pteranodontoidea, Lanceodontia) and recognition of a new cosmopolitan lineage of Cretaceous toothed pterodactyloids. Historical Biology 33:1266–1280.
   Web of Science ®Google Scholar
• Pinheiro, F. L., and T. Rodrigues. 2017. Anhanguera taxonomy revisited: is our understanding of Santana Group pterosaur diversity biased by poor biological and stratigraphic control? PeerJ 5.3285:1–29.
   Google Scholar
• Plieninger, F. 1901. Beiträge zur Kenntnis der Flugsaurier. Palaeontographica 48:65–90.
   Google Scholar
• Rodrigues, T., and A. W. A. Kellner. 2013. Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England. ZooKeys 308:1–112
   PubMed Web of Science ®Google Scholar
• Rohlf, F. J. 2010. TpsDig, version 216. Departamento of Ecology and Evolution, State University of New York at Stony Brook. Available at http://life.bio.sunysb.edu/morph/.
   Google Scholar
• Santos, O. S. 1985. Técnicas de Preparação de Fósseis; pp. 26:101–127 in D. A., Campos; R. C. G., Armestro, (eds.), Série Geológica, Seção Paleontologia e Estratigrafia.
   Google Scholar
• Unwin, D. M. 2001. An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England. Mitteilungen aus dem Museum für Naturkunde in Berlin, Geowissenschafiliche Reihe 4:189–221.
   Google Scholar
• Unwin, D. M. 2005. The Pterosaurs: from deep time. New Yorkm, Pi Press, 246 pp.
   Google Scholar
• Veldmeijer, A. J. 2003. Preliminary description of a skull and wing of a Brazilian Cretaceous (Santana Formation; Aptian-Albian) pterosaur (Pterodactyloidea) in the collection of the AMNH. PalArch’s Journal of Vertebrate Palaeontology 0:1–14.
   Google Scholar
• Veldmeijer, A. J., H. J. Meijer, and M. Signore. 2006. Coloborhynchus from the Lower Cretaceous Santana Formation, Brazil (Pterosauria, Pterodactyloidea, Anhangueridae); an update. PalArch’s Journal of Vertebrate Palaeontology, 3:15–29.
   Google Scholar
• Veldmeijer, A. J., H. J. Meijer, and M. Signore. 2009. Description of Pterosaurian (Pterodactyloidea: Anhangueridae, Brasileodactylus) remains from the Lower Cretaceous of Brazil. Deinsea 13:9–40.
   Google Scholar
• Viana, M. S. S., and V. H. L. Neumann. 2002. Membro Crato da Formação Santana, Chapada do Araripe, CE: Riquíssimo registro de fauna e flora do Cretáceo; pp. 113–120 in Schobbenhaus, C., Campos, D. A., Queiroz, E. T., Winge, M., and Berbert-Born, M. (orgs.). Sítios Geológicos e Paleontológicos do Brasil, Brasília, DNPM/CPRM/SIGEP.
   Google Scholar
• Vila Nova, B. C., J. M. Sayão, V. H. M. L. Neumann, and Kellner, A.W.A. 2014. Redescription of Cearadactylus atrox (Pterosauria, Pterodactyloidea) from the Early Cretaceous Romualdo Formation (Santana Group) of the Araripe Basin, Brazil. Journal of Vertebrate Paleontology 34:126–134.
   Web of Science ®Google Scholar
• Wang, X., A. W. A. Kellner, S. Juang, Q. Wang, Y. Ma, Y. Paidoula, X. Cheng, T. Rodrigues, X. Meng, J. Zhang, N. Li, and Z. Zhou. 2014. Sexually dimorphic tridimensionally preserves pterosaurs and their eggs from China. Current Biology 24:1323–1330.
   PubMed Web of Science ®Google Scholar
• Wang, X., A. W. A. Kellner, X. Cheng, S. Jiang, Q. Wang, J. M. Sayão, T. Rodrigues, F. Costa, N. Li, X. Meng and Z. Zhou. 2015. Eggshell and histology provide insight on the life history of a pterosaur with two functional ovaries. Anais da Academis Brasileira de Ciências 87: 1599–1609.
   PubMed Web of Science ®Google Scholar
• Wellnhofer, P. 1987. New crested pterosaurs from the Lower Cretaceous of Brazil. Mitteilungen der Bayerischen Staatssammlung für Paläontologie und Historische Geologie 27: 175–186.
   Google Scholar
• Wellnhofer, P. 1991. Weitere Pterosaurierfunde aus der Santana-Formation (Apt) der Chapada do Araripe, Brasilien. Palaeontographica A 215:43–101.
   Google Scholar
• Zhou, X., Pêgas, R. V., Ma, W., Han, G., Jin, X., Leal, M. E., Bonde, N., Kobayashi, Y., Lautenschlager, S., Wei, X., Shen, C. and Ji, S.. 2021. A new darwinopteran reveals arborealism and an opposed thumb. Current Biology 31(11): 2429–2436.
   PubMed Web of Science ®Google Scholar